544 lines
64 KiB
XML
544 lines
64 KiB
XML
<document id="4160BEA4DE491C39F193992878229E13" ID-DOI="10.11646/zootaxa.4039.1.3" ID-GBIF-Dataset="0514ea20-09ff-45ba-8a9e-758160a3015a" ID-ISSN="1175-5326" ID-Zenodo-Dep="236302" ID-ZooBank="C9B651D5-15D3-4849-9853-18A055E4600C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461324274228" checkinUser="plazi" docAuthor="Doughty, Paul, Kealley, Luke, Shoo, Luke P. & Melville, Jane" docDate="2015" docId="530887F58D49FFED56F4CD668A72FA62" docLanguage="en" docName="zt04039p117.pdf" docOrigin="Zootaxa 4039 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tympanocryptis pseudopsephos Doughty, Kealley, Shoo & Melville, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="110" masterDocId="AF31FF8D8D5FFFF45663CC168808FFFF" masterDocTitle="Revision of the Western Australian pebble-mimic dragon species-group (Tympanocryptis cephalus: Reptilia: Agamidae)" masterLastPageNumber="117" masterPageNumber="85" pageNumber="107" updateTime="1698618285306" updateUser="plazi">
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<mods:mods id="997E935DD3C9108875E2300029F8E66A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D8C06B2A93F6C30A2E22FD1EB5E9E523">
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<mods:title id="99DB7382A592F478D3662EFE4CED89F5">Revision of the Western Australian pebble-mimic dragon species-group (Tympanocryptis cephalus: Reptilia: Agamidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="10CE2D5CC872929135CDA3EB30802F94" type="personal">
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<mods:role id="AC0CFD31C2680F8BCC77E60F70070A76">
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<mods:roleTerm id="5267F3D1CB527591202AAC4DED999035">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8B5D9D7E170D47764EE9E15B1A7C89D8">Doughty, Paul</mods:namePart>
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</mods:name>
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<mods:name id="2AB82ED459457420A60220DBCCF6BB9F" type="personal">
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<mods:roleTerm id="02B5ACAA31DABE9D5E66F98D60DFE4CC">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F15988043837324DDECD8CE19857FA15">Kealley, Luke</mods:namePart>
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</mods:name>
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<mods:name id="D0F6513BEED89CA7A975CAE70D6D39FB" type="personal">
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<mods:roleTerm id="4B154A1EAA3349181DDDD3C6CBA59AD3">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="399548AB4F956393507BCC48AE0677C2">Shoo, Luke P.</mods:namePart>
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</mods:name>
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<mods:name id="5B73A00E3F8F04038E095AD02D6660CE" type="personal">
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<mods:role id="339E26ABFFAE461DD1C016639D603AF4">
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<mods:roleTerm id="E07D46BB3653E3BCFA028C8F7F3452E1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="51D086EE76F1C4A748E1943E454ECA5A">Melville, Jane</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FA5A0C03A15B2DDA030F4B2624CC1D72">text</mods:typeOfResource>
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<mods:title id="F3B20ACFE2564E1D24F9CA2DC6CC4D51">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D53D6BE475F25A4FC112C5515DCFDE48">
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<mods:date id="5B281AB50103283F85EA43593F474073">2015</mods:date>
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<mods:detail id="C8E8D9A1CD96B9B06AE3E9B7A9D0CFAA" type="volume">
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<mods:number id="2DB144772C05FD35E7556400EB918AA6">4039</mods:number>
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</mods:detail>
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<mods:detail id="074196723C0BF7B5DE9A378D5FC22D83" type="issue">
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<mods:number id="91F4161A5803F0B139A48BC32FD47975">1</mods:number>
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<mods:start id="A8FB4CD825736A4BB2A690E063B98674">85</mods:start>
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<mods:classification id="BFB270C5655C4A04501258F8C444A84B">journal article</mods:classification>
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<mods:identifier id="18984C14C7FF231FE2B957F432D1484C" type="DOI">10.11646/zootaxa.4039.1.3</mods:identifier>
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<mods:identifier id="2C66DF51AD428049514A6AF6D2043B85" type="GBIF-Dataset">0514ea20-09ff-45ba-8a9e-758160a3015a</mods:identifier>
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<mods:identifier id="9315F0C512ED96AE202D386821800326" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BD2EC5B8E715F282DD4B0B1CD895FC1E" type="Zenodo-Dep">236302</mods:identifier>
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<treatment id="530887F58D49FFED56F4CD668A72FA62" ID-DOI="http://doi.org/10.5281/zenodo.6120057" ID-GBIF-Taxon="119643907" ID-Zenodo-Dep="6120057" LSID="urn:lsid:plazi:treatment:530887F58D49FFED56F4CD668A72FA62" httpUri="http://treatment.plazi.org/id/530887F58D49FFED56F4CD668A72FA62" lastPageId="25" lastPageNumber="110" pageId="22" pageNumber="107">
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<subSubSection id="93BB65688D49FFE256F4CD668979FE30" pageId="22" pageNumber="107" type="nomenclature">
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<paragraph id="DB1E36E38D49FFE256F4CD668A70FE75" blockId="22.[151,632,368,463]" box="[151,632,368,394]" pageId="22" pageNumber="107">
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<heading id="8056818F8D49FFE256F4CD668A70FE75" bold="true" box="[151,632,368,394]" fontSize="11" level="1" pageId="22" pageNumber="107" reason="1">
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||
<emphasis id="E9D5EAF18D49FFE256F4CD668A70FE75" bold="true" box="[151,632,368,394]" pageId="22" pageNumber="107">
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||
<taxonomicName id="1CA14D608D49FFE256F4CD668A1CFE75" box="[151,532,368,394]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="107" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
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<emphasis id="E9D5EAF18D49FFE256F4CD668A1CFE75" bold="true" box="[151,532,368,394]" italics="true" pageId="22" pageNumber="107">Tympanocryptis pseudopsephos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F2E6578A8D49FFE25478CD668A70FE75" box="[539,632,368,394]" pageId="22" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="DB1E36E38D49FFE256F4CD848979FE30" blockId="22.[151,632,368,463]" pageId="22" pageNumber="107">
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Goldfields pebble-mimic dragons
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<figureCitation id="439A2A668D49FFE256F4CDA088EBFE30" box="[151,227,438,463]" captionStart="FIGURE 5" captionStartId="9.[151,250,1479,1501]" captionTargetBox="[176,1412,186,1446]" captionTargetId="figure@9.[163,1424,173,1458]" captionTargetPageId="9" captionText="FIGURE 5. Photographs in life of Tympanocryptis cephalus species-group members (left to right, top to bottom): a) T. cephalus (photo—G. Harold), b) T. gigas (M. Peterson), c) T. diabolicus sp. nov. (M. Peterson), d) T. fortescuensis sp. nov. (G. Harold), e) T. pseudopsephos sp. nov. — plain form (G. Harold), f) T. pseudopsephos sp. nov. — with charcoal wash on dorsum (G. Harold)." httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="22" pageNumber="107">Figs. 5</figureCitation>
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E, 5F, 14, 15
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="93BB65688D49FFE256F4CDE88D48FD34" pageId="22" pageNumber="107" type="materials_examined">
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<paragraph id="DB1E36E38D49FFE256F4CDE88C88FDC4" blockId="22.[151,1437,510,715]" pageId="22" pageNumber="107">
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<emphasis id="E9D5EAF18D49FFE256F4CDE88903FDE8" bold="true" box="[151,267,510,535]" pageId="22" pageNumber="107">
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<typeStatus id="041A88418D49FFE256F4CDE8890EFDE8" box="[151,262,510,535]" pageId="22" pageNumber="107" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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<collectionCode id="BDB0AE268D49FFE25775CDE98952FDE8" LSID="urn:lsid:biocol.org:col:34981" box="[278,346,511,535]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="107">WAM</collectionCode>
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||
R154790, an adult male collected
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<quantity id="1C599B068D49FFE2549FCDE98B4FFDE8" box="[764,839,510,535]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="22" pageNumber="107" unit="km" value="31.0">31 km</quantity>
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south-west of Doolgunna Homestead (
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||
<geoCoordinate id="BE9550248D49FFE25376CDE88D9FFDE8" box="[1301,1431,510,535]" direction="south" orientation="latitude" pageId="22" pageNumber="107" precision="15" value="-25.875834">25°52'33"S</geoCoordinate>
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||
,
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||
<geoCoordinate id="BE9550248D49FFE256F4CE35892FFDC4" box="[151,295,547,571]" direction="east" orientation="longitude" pageId="22" pageNumber="107" precision="15" value="119.02806">119°01'41"E</geoCoordinate>
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), Western
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<collectingCountry id="A3B676738D49FFE257FDCE348A00FDC4" box="[414,520,546,571]" name="Australia" pageId="22" pageNumber="107">Australia</collectingCountry>
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||
, on
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<date id="AF1F10238D49FFE25455CE348ACDFDC4" box="[566,709,546,571]" pageId="22" pageNumber="107" value="2004-04-05">5 April 2004</date>
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by B. Maryan and B. Budrey (
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||
<figureCitation id="439A2A668D49FFE2527CCE348C7BFDC4" box="[1055,1139,546,571]" captionStart="FIGURE 14" captionStartId="22.[151,250,1989,2011]" captionTargetBox="[151,1436,767,1966]" captionTargetId="figure@22.[151,1436,767,1966]" captionTargetPageId="22" captionText="FIGURE 14. Holotype of Tympanocryptis pseudopsephos sp. nov. (scale bar = 1 cm)." httpUri="https://zenodo.org/record/236315/files/figure.png" pageId="22" pageNumber="107">Fig. 14</figureCitation>
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||
).
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||
</paragraph>
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||
<paragraph id="DB1E36E38D49FFE256A4CE508D48FD34" blockId="22.[151,1437,510,715]" pageId="22" pageNumber="107">
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||
<emphasis id="E9D5EAF18D49FFE256A4CE50894EFDA0" bold="true" box="[199,326,582,607]" pageId="22" pageNumber="107">
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||
<typeStatus id="041A88418D49FFE256A4CE508949FDA0" box="[199,321,582,607]" pageId="22" pageNumber="107" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
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||
<collectionCode id="BDB0AE268D49FFE25732CE51899EFDA0" LSID="urn:lsid:biocol.org:col:34981" box="[337,406,583,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="107">WAM</collectionCode>
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||
R114550 (female),
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||
<quantity id="1C599B068D49FFE254E0CE508AC7FDA0" box="[643,719,582,607]" metricMagnitude="4" metricUnit="m" metricValue="6.7" pageId="22" pageNumber="107" unit="km" value="67.0">67 km</quantity>
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||
south of Capricorn Roadhouse,
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||
<quantity id="1C599B068D49FFE25233CE518C93FDA0" box="[1104,1179,582,607]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="22" pageNumber="107" unit="km" value="18.0">18 km</quantity>
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||
east of Bulloo Down homestead (
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||
<geoCoordinate id="BE9550248D49FFE25742CE7D8973FD7C" box="[289,379,619,643]" direction="south" orientation="latitude" pageId="22" pageNumber="107" precision="925" value="-24.0">24°00'S</geoCoordinate>
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||
,
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||
<geoCoordinate id="BE9550248D49FFE257E9CE7C89FCFD7C" box="[394,500,618,643]" direction="east" orientation="longitude" pageId="22" pageNumber="107" precision="925" value="119.75">119°45'E</geoCoordinate>
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||
);
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||
<collectionCode id="BDB0AE268D49FFE25469CE7D8A47FD7C" LSID="urn:lsid:biocol.org:col:34981" box="[522,591,619,643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="107">WAM</collectionCode>
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R136459 (female), Norseman Area (
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||
<geoCoordinate id="BE9550248D49FFE2559ECE7D8C50FD7C" box="[1021,1112,619,643]" direction="south" orientation="latitude" pageId="22" pageNumber="107" precision="925" value="-32.2">32°12'S</geoCoordinate>
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||
,
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||
<geoCoordinate id="BE9550248D49FFE25205CE7D8CD9FD7C" box="[1126,1233,619,643]" direction="east" orientation="longitude" pageId="22" pageNumber="107" precision="925" value="121.78333">121°47'E</geoCoordinate>
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||
);
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||
<collectionCode id="BDB0AE268D49FFE2528BCE7D8D25FD7C" LSID="urn:lsid:biocol.org:col:34981" box="[1256,1325,619,643]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="107">WAM</collectionCode>
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R164347 (male),
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||
<quantity id="1C599B068D49FFE25693CE988947FD58" box="[240,335,654,679]" metricMagnitude="4" metricUnit="m" metricValue="4.61" pageId="22" pageNumber="107" unit="km" value="46.1">46.1 km</quantity>
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east of Leonora (
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<geoCoordinate id="BE9550248D49FFE2547ECE988A96FD58" box="[541,670,654,679]" direction="south" orientation="latitude" pageId="22" pageNumber="107" precision="15" value="-28.86">28°51'36"S</geoCoordinate>
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||
,
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||
<geoCoordinate id="BE9550248D49FFE254CECE998B37FD58" box="[685,831,655,679]" direction="east" orientation="longitude" pageId="22" pageNumber="107" precision="15" value="121.80334">121°48'12"E</geoCoordinate>
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||
);
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<collectionCode id="BDB0AE268D49FFE25535CE998B93FD58" LSID="urn:lsid:biocol.org:col:34981" box="[854,923,655,679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="107">WAM</collectionCode>
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||
R164350 (female),
|
||
<quantity id="1C599B068D49FFE252E0CE998CE8FD58" box="[1155,1248,654,679]" metricMagnitude="4" metricUnit="m" metricValue="1.18" pageId="22" pageNumber="107" unit="km" value="11.8">11.8 km</quantity>
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||
east of Leonora (
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||
<geoCoordinate id="BE9550248D49FFE256FCCEA4892AFD34" box="[159,290,690,715]" direction="south" orientation="latitude" pageId="22" pageNumber="107" precision="15" value="-28.90639">28°54'23"S</geoCoordinate>
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||
,
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||
<geoCoordinate id="BE9550248D49FFE2574ECEA489C8FD34" box="[301,448,690,715]" direction="east" orientation="longitude" pageId="22" pageNumber="107" precision="15" value="121.44556">121°26'44"E</geoCoordinate>
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||
);
|
||
<collectionCode id="BDB0AE268D49FFE257B7CEA58A11FD34" LSID="urn:lsid:biocol.org:col:34981" box="[468,537,691,715]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="22" pageNumber="107">WAM</collectionCode>
|
||
R167478 (male),
|
||
<quantity id="1C599B068D49FFE25487CEA48B48FD34" box="[740,832,690,715]" metricMagnitude="4" metricUnit="m" metricValue="4.95" pageId="22" pageNumber="107" unit="km" value="49.5">49.5 km</quantity>
|
||
north of Leinster (
|
||
<geoCoordinate id="BE9550248D49FFE25270CEA48C9DFD34" box="[1043,1173,690,715]" direction="south" orientation="latitude" pageId="22" pageNumber="107" precision="15" value="-27.376667">27°22'36"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="BE9550248D49FFE252C2CEA48D3CFD34" box="[1185,1332,690,715]" direction="east" orientation="longitude" pageId="22" pageNumber="107" precision="15" value="120.58611">120°35'10"E</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8FDE666B8D49FFE256F4CBD38C0AF825" box="[151,1026,1988,2011]" httpUri="https://zenodo.org/record/236315/files/figure.png" pageId="22" pageNumber="107" targetBox="[151,1436,767,1966]" targetPageId="22">
|
||
<paragraph id="DB1E36E38D49FFE256F4CBD38C0AF825" blockId="22.[151,1026,1988,2011]" box="[151,1026,1988,2011]" pageId="22" pageNumber="107">
|
||
<emphasis id="E9D5EAF18D49FFE256F4CBD38917F825" bold="true" box="[151,287,1989,2011]" pageId="22" pageNumber="107">FIGURE 14.</emphasis>
|
||
Holotype of
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<taxonomicName id="1CA14D608D49FFE257C6CBD08AEEF825" box="[421,742,1988,2011]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="107" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D49FFE257C6CBD08AEEF825" box="[421,742,1988,2011]" italics="true" pageId="22" pageNumber="107">Tympanocryptis pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D49FFE2548FCBD38B35F825" box="[748,829,1989,2010]" pageId="22" pageNumber="107" rank="species">
|
||
<emphasis id="E9D5EAF18D49FFE2548FCBD38B35F825" bold="true" box="[748,829,1989,2010]" pageId="22" pageNumber="107">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(scale bar = 1 cm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8FDE666B8D48FFE356F4CF5E8C0EFCA1" box="[151,1030,840,862]" httpUri="https://zenodo.org/record/236316/files/figure.png" pageId="23" pageNumber="108" targetBox="[151,1436,193,818]" targetPageId="23">
|
||
<paragraph id="DB1E36E38D48FFE356F4CF5E8C0EFCA1" blockId="23.[151,1030,840,862]" box="[151,1030,840,862]" pageId="23" pageNumber="108">
|
||
<emphasis id="E9D5EAF18D48FFE356F4CF5E8917FCA2" bold="true" box="[151,287,840,862]" pageId="23" pageNumber="108">FIGURE 15.</emphasis>
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Paratypes of
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<taxonomicName id="1CA14D608D48FFE357C9CF5F8AE2FCA1" box="[426,746,840,862]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="108" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D48FFE357C9CF5F8AE2FCA1" box="[426,746,840,862]" italics="true" pageId="23" pageNumber="108">Tympanocryptis pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E9D5EAF18D48FFE35492CF5E8B49FCA2" bold="true" box="[753,833,840,861]" pageId="23" pageNumber="108">
|
||
<taxonomicNameLabel id="F2E6578A8D48FFE35492CF5E8B49FCA2" box="[753,833,840,861]" pageId="23" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
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||
(scale bar = 1 cm).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="93BB65688D48FFE356A4CF9C8B6CFB84" pageId="23" pageNumber="108" type="diagnosis">
|
||
<paragraph id="DB1E36E38D48FFE356A4CF9C8B6CFB84" blockId="23.[151,1437,906,2011]" pageId="23" pageNumber="108">
|
||
<emphasis id="E9D5EAF18D48FFE356A4CF9C8949FC5C" bold="true" box="[199,321,906,931]" pageId="23" pageNumber="108">Diagnosis.</emphasis>
|
||
Distinguished from other
|
||
<taxonomicName id="1CA14D608D48FFE35409CF9A8B14FC5C" box="[618,796,908,931]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="108" phylum="Chordata" rank="genus">
|
||
<emphasis id="E9D5EAF18D48FFE35409CF9A8B14FC5C" box="[618,796,908,931]" italics="true" pageId="23" pageNumber="108">Tympanocryptis</emphasis>
|
||
</taxonomicName>
|
||
by the following combination of character states: rotund body shape, presence of two pre-cloacal pores, lack of longitudinal stripes on the dorsum, presence of enlarged scales with raised spines arranged in short transverse rows of 4–7 scales, concave snout formed by protruding mouth, scales on snout rugose with feeble keels, rostral width ~1–2 times height, keels on scales of upper arm not aligned, well-defined row of enlarged scales at anterior and dorsal edge of thigh forming conspicuous ridge, scales on dorsal surface of thigh not aligned, ventrals smooth, and reddish-brown ground color, lacking lines but often possessing a dark charcoal-like wash over the head and dorsum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D48FFEC56A4C8908CFFF85D" lastPageId="24" lastPageNumber="109" pageId="23" pageNumber="108" type="description">
|
||
<paragraph id="DB1E36E38D48FFE356A4C8908CD5F944" blockId="23.[151,1437,906,2011]" pageId="23" pageNumber="108">
|
||
<emphasis id="E9D5EAF18D48FFE356A4C8908951FB60" bold="true" box="[199,345,1158,1183]" pageId="23" pageNumber="108">Description.</emphasis>
|
||
A small-bodied (to
|
||
<quantity id="1C599B068D48FFE3545DC8908AACFB61" box="[574,676,1158,1183]" metricMagnitude="-2" metricUnit="m" metricValue="5.65" pageId="23" pageNumber="108" unit="mm" value="56.5">56.5 mm</quantity>
|
||
SVL), rotund dragon; small head with blunt snout with protruding mouth; short neck; moderately short gracile limbs, ArmL%SVL—0.185 (0.016), LegL%SVL—0.235 (0.017); moderately short gracile digits; short tail. Head small, HeadL%SVL—0.307 (0.017), HeadW%SVL—0.258 (0.013), HeadD%SVL—0.158 (0.008); neck ~3/4 of widest part of head; snout short, SnoutL%HeadL—0.259 (0.025); snout straight or concave when viewed laterally, if concave then snout tip projects forward; line of mouth slightly projecting forming a shelf-like appearance, narrowing to blunt tip; canthus defined but rounded, forming continuous line with projecting brow ridge; nostril located below canthus in enlarged scale, opening projecting dorsally; eye moderate, EyeL%HeadL—0.258 (0.022); eyes with laterally-projecting scaly eyelids forming a fringe, often projecting past brow when viewed dorsally; tympana covered with fine scales, encircled by scattered enlarged scales with raised apex; scales on snout rugose, occasionally with feeble unaligned keels; scales on crown slightly larger with feeble unaligned keels; scales on back of head small with few variably-sized low scattered spines; rostral scale ~ 1–2 times wider than tall; 10–15 supralabial scales, 4–5 rows of scales above supralabial row, keeled, uppermost row slightly larger with larger keels and continuing to temporal region, forming edge of eye socket; loosely defined cluster of enlarged spines at posterior and lateral corners of head; mental 1.5 times long as wide; 11–16 infralabials with low keels; 4–5 rows of scales below infralabials with low keels parallel to angle of jaw, creating a slightly terraced appearance; gulars kite-shaped and smooth; prominent gular fold.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D48FFE356A4CAD08A35F824" blockId="23.[151,1437,906,2011]" pageId="23" pageNumber="108">Body dorsoventrally compressed, ovoid in dorsal view with widest part ~2 times wider than neck and pelvis; TrunkL%SVL—0.473 (0.048); dorsum with heterogeneous scales in size and shape; largest scales with sharp spines angled 10–30° posteriorly and ~4–7 times larger than smallest scales; large dorsal scales with spines tending to occur in transversely-aligned clusters of ~4–6; smaller dorsal scales rugose or with weak, low keels, arranged in loose whorls around clusters of large, spiny scales; dorsolateral edge of pelvis always with a cluster of enlarged spines at posterior edge where skin is fused to bone; ventral scales homogeneous, approximately half the size of large dorsal scales, kite-shaped and arranged in diagonal rows; ventral scales smooth, or at most with raised midline (no keel or protruding spine).</paragraph>
|
||
<paragraph id="DB1E36E38D47FFEC56A4CC818AD6FDC3" blockId="24.[151,1436,151,752]" pageId="24" pageNumber="109">Limbs largely covered in elongate kite or teardrop-shaped scales with prominent low keels, spine usually protruding beyond distal edge of scale, interspersed with occasional small unkeeled scales; arms and legs moderately long, ArmL%SVL—0.185 (0.016), LegL%SVL—0.235 (0.017); scales on dorsal surface of upper arm large with keels tending not to align; keels of dorsal scales on lower arm tending to align, often forming lines that extend to hand and fingers; keels of ventral scales mostly aligned forming lines along the length of the arm to palmar surfaces; scales on underside of digit with two rows of spiny lamellae; claw long and recurved, lower portion terminating with circular opening, upper portion continuing past ultimate lamellae to form sharp claw; finger length: 4> 3> 2 = 5> 1; scales on legs kite or teardrop-shaped; tops of upper and lower leg with large heterogeneous non-aligned scales, elsewhere with low keels that align and extend to feet and toes; scales at insertion of limb to body small, rectangular and lacking keels; anterior edge of thigh with enlarged scales forming conspicuous ridge; abrupt transition from dorsal to posterior edge of leg (large to small scales); scales on underside of toes as for fingers; toe length: 4 >> 3> 2 = 5> 1.</paragraph>
|
||
<paragraph id="DB1E36E38D47FFEC56A4CE5188ECFD32" blockId="24.[151,1436,151,752]" pageId="24" pageNumber="109">Pre-cloacal pores 2, set among 3–5 scales, positioned anterior to distal edges of cloaca, midway between anterior and posterior edge of leg; tail short and thin with blunt tip, TailL%SVL—1.268 (0.102); scattered enlarged scales aligning along most of the length of the tail to terminus; lateral surfaces of tail base with short protruding spines.</paragraph>
|
||
<paragraph id="DB1E36E38D47FFEC56A4CECE8B5BFD0F" blockId="24.[151,1436,151,752]" box="[199,851,727,752]" pageId="24" pageNumber="109">
|
||
Measurements for the
|
||
<typeStatus id="041A88418D47FFEC57A1CECE89FAFD0F" box="[450,498,728,752]" pageId="24" pageNumber="109">type</typeStatus>
|
||
series are presented in
|
||
<tableCitation id="962303588D47FFEC549BCEC18B46FD0F" box="[760,846,727,752]" captionStart="TABLE 5" captionStartId="24.[151,239,796,819]" captionTargetBox="[159,1338,846,1612]" captionTargetPageId="24" captionText="TABLE 5. Summary of meristic (mm) and mensural data for the type series of Tympanocryptis psuedopsephos sp. nov." httpUri="http://table.plazi.org/id/8FDE666B8D47FFEC56F4CF0A8D9BFCCC" pageId="24" pageNumber="109" tableUuid="8FDE666B8D47FFEC56F4CF0A8D9BFCCC">Table 5</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="8FDE666B8D47FFEC56F4CF0A8D9BFCCC" ID-Table-UUID="8FDE666B8D47FFEC56F4CF0A8D9BFCCC" box="[151,1427,796,819]" httpUri="http://table.plazi.org/id/8FDE666B8D47FFEC56F4CF0A8D9BFCCC" pageId="24" pageNumber="109" targetBox="[159,1338,846,1612]" targetIsTable="true" targetPageId="24">
|
||
<paragraph id="DB1E36E38D47FFEC56F4CF0A8D9BFCCC" blockId="24.[151,1427,796,819]" box="[151,1427,796,819]" pageId="24" pageNumber="109">
|
||
<emphasis id="E9D5EAF18D47FFEC56F4CF0A8902FCCC" bold="true" box="[151,266,796,819]" pageId="24" pageNumber="109">TABLE 5.</emphasis>
|
||
Summary of meristic (mm) and mensural data for the type series of
|
||
<taxonomicName id="1CA14D608D47FFEC5587CF0A8D3FFCCC" box="[996,1335,796,819]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="109" phylum="Chordata" rank="species" species="psuedopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D47FFEC5587CF0A8D3FFCCC" box="[996,1335,796,819]" italics="true" pageId="24" pageNumber="109">Tympanocryptis psuedopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D47FFEC535ECF0A8D9BFCCC" box="[1341,1427,796,819]" pageId="24" pageNumber="109" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DB1E36E38D47FFEC56FCCF598D06F9B3" pageId="24" pageNumber="109">
|
||
<table id="A9A1C4438D47000B56FCCF588D32F9B3" box="[159,1338,846,1612]" gridcols="6" gridrows="19" pageId="24" pageNumber="109">
|
||
<tr id="659134A18D47000B56FCCF588D32FC9B" box="[159,1338,846,868]" gridrow="0" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCF5889A1FC9B" box="[159,425,846,868]" gridcol="0" gridrow="0" pageId="24" pageNumber="109">R# 154790</th>
|
||
<th id="26405DDD8D47000B546DCF588A52FC9B" box="[526,602,846,868]" gridcol="1" gridrow="0" pageId="24" pageNumber="109">164347</th>
|
||
<th id="26405DDD8D47000B54A5CF588B19FC9B" box="[710,785,846,868]" gridcol="2" gridrow="0" pageId="24" pageNumber="109">114550</th>
|
||
<th id="26405DDD8D47000B551DCF588BC2FC9B" box="[894,970,846,868]" gridcol="3" gridrow="0" pageId="24" pageNumber="109">167478</th>
|
||
<th id="26405DDD8D47000B5255CF588C8AFC9B" box="[1078,1154,846,868]" gridcol="4" gridrow="0" pageId="24" pageNumber="109">136459</th>
|
||
<th id="26405DDD8D47000B528DCF588D32FC9B" box="[1262,1338,846,868]" gridcol="5" gridrow="0" pageId="24" pageNumber="109">164350</th>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCCF6E8D32FC71" box="[159,1338,888,910]" gridrow="1" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCF6E89A1FC71" box="[159,425,888,910]" gridcol="0" gridrow="1" pageId="24" pageNumber="109">Sex Male</th>
|
||
<td id="26405DDD8D47000B546DCF6E8A52FC71" box="[526,602,888,910]" gridcol="1" gridrow="1" pageId="24" pageNumber="109">Male</td>
|
||
<td id="26405DDD8D47000B54A5CF6E8B19FC71" box="[710,785,888,910]" gridcol="2" gridrow="1" pageId="24" pageNumber="109">Female</td>
|
||
<td id="26405DDD8D47000B551DCF6E8BC2FC71" box="[894,970,888,910]" gridcol="3" gridrow="1" pageId="24" pageNumber="109">Male</td>
|
||
<td id="26405DDD8D47000B5255CF6E8C8AFC71" box="[1078,1154,888,910]" gridcol="4" gridrow="1" pageId="24" pageNumber="109">Female</td>
|
||
<td id="26405DDD8D47000B528DCF6E8D32FC71" box="[1262,1338,888,910]" gridcol="5" gridrow="1" pageId="24" pageNumber="109">Female</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCCFB78D32FC48" box="[159,1338,929,951]" gridrow="2" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCFB789A1FC48" box="[159,425,929,951]" gridcol="0" gridrow="2" pageId="24" pageNumber="109">SVL 49.5</th>
|
||
<td id="26405DDD8D47000B546DCFB78A52FC48" box="[526,602,929,951]" gridcol="1" gridrow="2" pageId="24" pageNumber="109">44.5</td>
|
||
<td id="26405DDD8D47000B54A5CFB78B19FC48" box="[710,785,929,951]" gridcol="2" gridrow="2" pageId="24" pageNumber="109">52.5</td>
|
||
<td id="26405DDD8D47000B551DCFB78BC2FC48" box="[894,970,929,951]" gridcol="3" gridrow="2" pageId="24" pageNumber="109">46.5</td>
|
||
<td id="26405DDD8D47000B5255CFB78C8AFC48" box="[1078,1154,929,951]" gridcol="4" gridrow="2" pageId="24" pageNumber="109">49.5</td>
|
||
<td id="26405DDD8D47000B528DCFB78D32FC48" box="[1262,1338,929,951]" gridcol="5" gridrow="2" pageId="24" pageNumber="109">44.5</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCCFDC8D32FC1F" box="[159,1338,970,992]" gridrow="3" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCFDC89A1FC1F" box="[159,425,970,992]" gridcol="0" gridrow="3" pageId="24" pageNumber="109">TrunkL 18.6</th>
|
||
<td id="26405DDD8D47000B546DCFDC8A52FC1F" box="[526,602,970,992]" gridcol="1" gridrow="3" pageId="24" pageNumber="109">19.3</td>
|
||
<td id="26405DDD8D47000B54A5CFDC8B19FC1F" box="[710,785,970,992]" gridcol="2" gridrow="3" pageId="24" pageNumber="109">21.7</td>
|
||
<td id="26405DDD8D47000B551DCFDC8BC2FC1F" box="[894,970,970,992]" gridcol="3" gridrow="3" pageId="24" pageNumber="109">25.1</td>
|
||
<td id="26405DDD8D47000B5255CFDC8C8AFC1F" box="[1078,1154,970,992]" gridcol="4" gridrow="3" pageId="24" pageNumber="109">21.3</td>
|
||
<td id="26405DDD8D47000B528DCFDC8D32FC1F" box="[1262,1338,970,992]" gridcol="5" gridrow="3" pageId="24" pageNumber="109">20.3</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCCFE28D32FBF5" box="[159,1338,1012,1034]" gridrow="4" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCFE289A1FBF5" box="[159,425,1012,1034]" gridcol="0" gridrow="4" pageId="24" pageNumber="109">TailL 69.5</th>
|
||
<td id="26405DDD8D47000B546DCFE28A52FBF5" box="[526,602,1012,1034]" gridcol="1" gridrow="4" pageId="24" pageNumber="109">55.5</td>
|
||
<td id="26405DDD8D47000B54A5CFE28B19FBF5" box="[710,785,1012,1034]" gridcol="2" gridrow="4" pageId="24" pageNumber="109">73.0</td>
|
||
<td id="26405DDD8D47000B551DCFE28BC2FBF5" box="[894,970,1012,1034]" gridcol="3" gridrow="4" pageId="24" pageNumber="109">68.0</td>
|
||
<td id="26405DDD8D47000B5255CFE28C8AFBF5" box="[1078,1154,1012,1034]" gridcol="4" gridrow="4" pageId="24" pageNumber="109">58.5</td>
|
||
<td id="26405DDD8D47000B528DCFE28D32FBF5" box="[1262,1338,1012,1034]" gridcol="5" gridrow="4" pageId="24" pageNumber="109">55.0</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC80B8D32FBCC" box="[159,1338,1053,1075]" gridrow="5" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC80B89A1FBCC" box="[159,425,1053,1075]" gridcol="0" gridrow="5" pageId="24" pageNumber="109">HeadL 16.4</th>
|
||
<td id="26405DDD8D47000B546DC80B8A52FBCC" box="[526,602,1053,1075]" gridcol="1" gridrow="5" pageId="24" pageNumber="109">13.7</td>
|
||
<td id="26405DDD8D47000B54A5C80B8B19FBCC" box="[710,785,1053,1075]" gridcol="2" gridrow="5" pageId="24" pageNumber="109">16.8</td>
|
||
<td id="26405DDD8D47000B551DC80B8BC2FBCC" box="[894,970,1053,1075]" gridcol="3" gridrow="5" pageId="24" pageNumber="109">14.3</td>
|
||
<td id="26405DDD8D47000B5255C80B8C8AFBCC" box="[1078,1154,1053,1075]" gridcol="4" gridrow="5" pageId="24" pageNumber="109">14.4</td>
|
||
<td id="26405DDD8D47000B528DC80B8D32FBCC" box="[1262,1338,1053,1075]" gridcol="5" gridrow="5" pageId="24" pageNumber="109">14.3</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC8508D32FBA3" box="[159,1338,1094,1116]" gridrow="6" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC85089A1FBA3" box="[159,425,1094,1116]" gridcol="0" gridrow="6" pageId="24" pageNumber="109">HeadW 13.1</th>
|
||
<td id="26405DDD8D47000B546DC8508A52FBA3" box="[526,602,1094,1116]" gridcol="1" gridrow="6" pageId="24" pageNumber="109">11.7</td>
|
||
<td id="26405DDD8D47000B54A5C8508B19FBA3" box="[710,785,1094,1116]" gridcol="2" gridrow="6" pageId="24" pageNumber="109">13.7</td>
|
||
<td id="26405DDD8D47000B551DC8508BC2FBA3" box="[894,970,1094,1116]" gridcol="3" gridrow="6" pageId="24" pageNumber="109">12.7</td>
|
||
<td id="26405DDD8D47000B5255C8508C8AFBA3" box="[1078,1154,1094,1116]" gridcol="4" gridrow="6" pageId="24" pageNumber="109">12.6</td>
|
||
<td id="26405DDD8D47000B528DC8508D32FBA3" box="[1262,1338,1094,1116]" gridcol="5" gridrow="6" pageId="24" pageNumber="109">11.2</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC8668D32FB79" box="[159,1338,1136,1158]" gridrow="7" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC86689A1FB79" box="[159,425,1136,1158]" gridcol="0" gridrow="7" pageId="24" pageNumber="109">HeadD 8.3</th>
|
||
<td id="26405DDD8D47000B546DC8668A52FB79" box="[526,602,1136,1158]" gridcol="1" gridrow="7" pageId="24" pageNumber="109">6.9</td>
|
||
<td id="26405DDD8D47000B54A5C8668B19FB79" box="[710,785,1136,1158]" gridcol="2" gridrow="7" pageId="24" pageNumber="109">7.9</td>
|
||
<td id="26405DDD8D47000B551DC8668BC2FB79" box="[894,970,1136,1158]" gridcol="3" gridrow="7" pageId="24" pageNumber="109">7.5</td>
|
||
<td id="26405DDD8D47000B5255C8668C8AFB79" box="[1078,1154,1136,1158]" gridcol="4" gridrow="7" pageId="24" pageNumber="109">8.1</td>
|
||
<td id="26405DDD8D47000B528DC8668D32FB79" box="[1262,1338,1136,1158]" gridcol="5" gridrow="7" pageId="24" pageNumber="109">6.9</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC88F8D32FB50" box="[159,1338,1177,1199]" gridrow="8" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC88F89A1FB50" box="[159,425,1177,1199]" gridcol="0" gridrow="8" pageId="24" pageNumber="109">SnoutL 3.9</th>
|
||
<td id="26405DDD8D47000B546DC88F8A52FB50" box="[526,602,1177,1199]" gridcol="1" gridrow="8" pageId="24" pageNumber="109">3.4</td>
|
||
<td id="26405DDD8D47000B54A5C88F8B19FB50" box="[710,785,1177,1199]" gridcol="2" gridrow="8" pageId="24" pageNumber="109">4.3</td>
|
||
<td id="26405DDD8D47000B551DC88F8BC2FB50" box="[894,970,1177,1199]" gridcol="3" gridrow="8" pageId="24" pageNumber="109">3.6</td>
|
||
<td id="26405DDD8D47000B5255C88F8C8AFB50" box="[1078,1154,1177,1199]" gridcol="4" gridrow="8" pageId="24" pageNumber="109">3.7</td>
|
||
<td id="26405DDD8D47000B528DC88F8D32FB50" box="[1262,1338,1177,1199]" gridcol="5" gridrow="8" pageId="24" pageNumber="109">3.6</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC8D48D32FB27" box="[159,1338,1218,1240]" gridrow="9" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC8D489A1FB27" box="[159,425,1218,1240]" gridcol="0" gridrow="9" pageId="24" pageNumber="109">SnoutW 5.9</th>
|
||
<td id="26405DDD8D47000B546DC8D48A52FB27" box="[526,602,1218,1240]" gridcol="1" gridrow="9" pageId="24" pageNumber="109">5.0</td>
|
||
<td id="26405DDD8D47000B54A5C8D48B19FB27" box="[710,785,1218,1240]" gridcol="2" gridrow="9" pageId="24" pageNumber="109">6.1</td>
|
||
<td id="26405DDD8D47000B551DC8D48BC2FB27" box="[894,970,1218,1240]" gridcol="3" gridrow="9" pageId="24" pageNumber="109">5.9</td>
|
||
<td id="26405DDD8D47000B5255C8D48C8AFB27" box="[1078,1154,1218,1240]" gridcol="4" gridrow="9" pageId="24" pageNumber="109">6.1</td>
|
||
<td id="26405DDD8D47000B528DC8D48D32FB27" box="[1262,1338,1218,1240]" gridcol="5" gridrow="9" pageId="24" pageNumber="109">5.1</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC8FA8D32FAFD" box="[159,1338,1260,1282]" gridrow="10" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC8FA89A1FAFD" box="[159,425,1260,1282]" gridcol="0" gridrow="10" pageId="24" pageNumber="109">EyeL 4.1</th>
|
||
<td id="26405DDD8D47000B546DC8FA8A52FAFD" box="[526,602,1260,1282]" gridcol="1" gridrow="10" pageId="24" pageNumber="109">3.7</td>
|
||
<td id="26405DDD8D47000B54A5C8FA8B19FAFD" box="[710,785,1260,1282]" gridcol="2" gridrow="10" pageId="24" pageNumber="109">3.8</td>
|
||
<td id="26405DDD8D47000B551DC8FA8BC2FAFD" box="[894,970,1260,1282]" gridcol="3" gridrow="10" pageId="24" pageNumber="109">3.8</td>
|
||
<td id="26405DDD8D47000B5255C8FA8C8AFAFD" box="[1078,1154,1260,1282]" gridcol="4" gridrow="10" pageId="24" pageNumber="109">3.8</td>
|
||
<td id="26405DDD8D47000B528DC8FA8D32FAFD" box="[1262,1338,1260,1282]" gridcol="5" gridrow="10" pageId="24" pageNumber="109">3.5</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC9038D32FAD4" box="[159,1338,1301,1323]" gridrow="11" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC90389A1FAD4" box="[159,425,1301,1323]" gridcol="0" gridrow="11" pageId="24" pageNumber="109">InterOrb 1.8</th>
|
||
<td id="26405DDD8D47000B546DC9038A52FAD4" box="[526,602,1301,1323]" gridcol="1" gridrow="11" pageId="24" pageNumber="109">1.7</td>
|
||
<td id="26405DDD8D47000B54A5C9038B19FAD4" box="[710,785,1301,1323]" gridcol="2" gridrow="11" pageId="24" pageNumber="109">1.7</td>
|
||
<td id="26405DDD8D47000B551DC9038BC2FAD4" box="[894,970,1301,1323]" gridcol="3" gridrow="11" pageId="24" pageNumber="109">1.8</td>
|
||
<td id="26405DDD8D47000B5255C9038C8AFAD4" box="[1078,1154,1301,1323]" gridcol="4" gridrow="11" pageId="24" pageNumber="109">2.1</td>
|
||
<td id="26405DDD8D47000B528DC9038D32FAD4" box="[1262,1338,1301,1323]" gridcol="5" gridrow="11" pageId="24" pageNumber="109">2.3</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC9288D32FAAB" box="[159,1338,1342,1364]" gridrow="12" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC92889A1FAAB" box="[159,425,1342,1364]" gridcol="0" gridrow="12" pageId="24" pageNumber="109">ArmL 9.2</th>
|
||
<td id="26405DDD8D47000B546DC9288A52FAAB" box="[526,602,1342,1364]" gridcol="1" gridrow="12" pageId="24" pageNumber="109">8.1</td>
|
||
<td id="26405DDD8D47000B54A5C9288B19FAAB" box="[710,785,1342,1364]" gridcol="2" gridrow="12" pageId="24" pageNumber="109">9.6</td>
|
||
<td id="26405DDD8D47000B551DC9288BC2FAAB" box="[894,970,1342,1364]" gridcol="3" gridrow="12" pageId="24" pageNumber="109">8.3</td>
|
||
<td id="26405DDD8D47000B5255C9288C8AFAAB" box="[1078,1154,1342,1364]" gridcol="4" gridrow="12" pageId="24" pageNumber="109">9.3</td>
|
||
<td id="26405DDD8D47000B528DC9288D32FAAB" box="[1262,1338,1342,1364]" gridcol="5" gridrow="12" pageId="24" pageNumber="109">9.4</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC97E8D32FA81" box="[159,1338,1384,1406]" gridrow="13" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC97E89A1FA81" box="[159,425,1384,1406]" gridcol="0" gridrow="13" pageId="24" pageNumber="109">HeadL 7.3</th>
|
||
<td id="26405DDD8D47000B546DC97E8A52FA81" box="[526,602,1384,1406]" gridcol="1" gridrow="13" pageId="24" pageNumber="109">6.4</td>
|
||
<td id="26405DDD8D47000B54A5C97E8B19FA81" box="[710,785,1384,1406]" gridcol="2" gridrow="13" pageId="24" pageNumber="109">8.5</td>
|
||
<td id="26405DDD8D47000B551DC97E8BC2FA81" box="[894,970,1384,1406]" gridcol="3" gridrow="13" pageId="24" pageNumber="109">7.3</td>
|
||
<td id="26405DDD8D47000B5255C97E8C8AFA81" box="[1078,1154,1384,1406]" gridcol="4" gridrow="13" pageId="24" pageNumber="109">7.8</td>
|
||
<td id="26405DDD8D47000B528DC97E8D32FA81" box="[1262,1338,1384,1406]" gridcol="5" gridrow="13" pageId="24" pageNumber="109">8.9</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC9878D32FA58" box="[159,1338,1425,1447]" gridrow="14" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC98789A1FA58" box="[159,425,1425,1447]" gridcol="0" gridrow="14" pageId="24" pageNumber="109">LegL 12.4</th>
|
||
<td id="26405DDD8D47000B546DC9878A52FA58" box="[526,602,1425,1447]" gridcol="1" gridrow="14" pageId="24" pageNumber="109">10.8</td>
|
||
<td id="26405DDD8D47000B54A5C9878B19FA58" box="[710,785,1425,1447]" gridcol="2" gridrow="14" pageId="24" pageNumber="109">12.7</td>
|
||
<td id="26405DDD8D47000B551DC9878BC2FA58" box="[894,970,1425,1447]" gridcol="3" gridrow="14" pageId="24" pageNumber="109">11.2</td>
|
||
<td id="26405DDD8D47000B5255C9878C8AFA58" box="[1078,1154,1425,1447]" gridcol="4" gridrow="14" pageId="24" pageNumber="109">11.5</td>
|
||
<td id="26405DDD8D47000B528DC9878D32FA58" box="[1262,1338,1425,1447]" gridcol="5" gridrow="14" pageId="24" pageNumber="109">11.2</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC9AC8D32FA2F" box="[159,1338,1466,1488]" gridrow="15" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC9AC89A1FA2F" box="[159,425,1466,1488]" gridcol="0" gridrow="15" pageId="24" pageNumber="109">FootL 15.5</th>
|
||
<td id="26405DDD8D47000B546DC9AC8A52FA2F" box="[526,602,1466,1488]" gridcol="1" gridrow="15" pageId="24" pageNumber="109">12.0</td>
|
||
<td id="26405DDD8D47000B54A5C9AC8B19FA2F" box="[710,785,1466,1488]" gridcol="2" gridrow="15" pageId="24" pageNumber="109">14.6</td>
|
||
<td id="26405DDD8D47000B551DC9AC8BC2FA2F" box="[894,970,1466,1488]" gridcol="3" gridrow="15" pageId="24" pageNumber="109">13.8</td>
|
||
<td id="26405DDD8D47000B5255C9AC8C8AFA2F" box="[1078,1154,1466,1488]" gridcol="4" gridrow="15" pageId="24" pageNumber="109">14.0</td>
|
||
<td id="26405DDD8D47000B528DC9AC8D32FA2F" box="[1262,1338,1466,1488]" gridcol="5" gridrow="15" pageId="24" pageNumber="109">13.5</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCC9F28D32FA05" box="[159,1338,1508,1530]" gridrow="16" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCC9F289A1FA05" box="[159,425,1508,1530]" gridcol="0" gridrow="16" pageId="24" pageNumber="109">HeadW/SVL 0.3</th>
|
||
<td id="26405DDD8D47000B546DC9F28A52FA05" box="[526,602,1508,1530]" gridcol="1" gridrow="16" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B54A5C9F28B19FA05" box="[710,785,1508,1530]" gridcol="2" gridrow="16" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B551DC9F28BC2FA05" box="[894,970,1508,1530]" gridcol="3" gridrow="16" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B5255C9F28C8AFA05" box="[1078,1154,1508,1530]" gridcol="4" gridrow="16" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B528DC9F28D32FA05" box="[1262,1338,1508,1530]" gridcol="5" gridrow="16" pageId="24" pageNumber="109">0.3</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCCA1B8D32F9DC" box="[159,1338,1549,1571]" gridrow="17" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCA1B89A1F9DC" box="[159,425,1549,1571]" gridcol="0" gridrow="17" pageId="24" pageNumber="109">FootL/SVL 0.3</th>
|
||
<td id="26405DDD8D47000B546DCA1B8A52F9DC" box="[526,602,1549,1571]" gridcol="1" gridrow="17" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B54A5CA1B8B19F9DC" box="[710,785,1549,1571]" gridcol="2" gridrow="17" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B551DCA1B8BC2F9DC" box="[894,970,1549,1571]" gridcol="3" gridrow="17" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B5255CA1B8C8AF9DC" box="[1078,1154,1549,1571]" gridcol="4" gridrow="17" pageId="24" pageNumber="109">0.3</td>
|
||
<td id="26405DDD8D47000B528DCA1B8D32F9DC" box="[1262,1338,1549,1571]" gridcol="5" gridrow="17" pageId="24" pageNumber="109">0.3</td>
|
||
</tr>
|
||
<tr id="659134A18D47000B56FCCA208D32F9B3" box="[159,1338,1590,1612]" gridrow="18" pageId="24" pageNumber="109">
|
||
<th id="26405DDD8D47000B56FCCA2089A1F9B3" box="[159,425,1590,1612]" gridcol="0" gridrow="18" pageId="24" pageNumber="109">TailL/SVL 1.4</th>
|
||
<td id="26405DDD8D47000B546DCA208A52F9B3" box="[526,602,1590,1612]" gridcol="1" gridrow="18" pageId="24" pageNumber="109">1.3</td>
|
||
<td id="26405DDD8D47000B54A5CA208B19F9B3" box="[710,785,1590,1612]" gridcol="2" gridrow="18" pageId="24" pageNumber="109">1.4</td>
|
||
<td id="26405DDD8D47000B551DCA208BC2F9B3" box="[894,970,1590,1612]" gridcol="3" gridrow="18" pageId="24" pageNumber="109">1.5</td>
|
||
<td id="26405DDD8D47000B5255CA208C8AF9B3" box="[1078,1154,1590,1612]" gridcol="4" gridrow="18" pageId="24" pageNumber="109">1.2</td>
|
||
<td id="26405DDD8D47000B528DCA208D32F9B3" box="[1262,1338,1590,1612]" gridcol="5" gridrow="18" pageId="24" pageNumber="109">1.2</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D47FFEC56A4CA9B8BD4F881" blockId="24.[151,1436,1677,2026]" pageId="24" pageNumber="109">
|
||
<emphasis id="E9D5EAF18D47FFEC56A4CA9B89DFF95A" box="[199,471,1677,1702]" italics="true" pageId="24" pageNumber="109">Coloration and pattern.</emphasis>
|
||
In life (
|
||
<figureCitation id="439A2A668D47FFEC5454CA9B8A8EF959" box="[567,646,1677,1702]" captionStart="FIGURE 5" captionStartId="9.[151,250,1479,1501]" captionTargetBox="[176,1412,186,1446]" captionTargetId="figure@9.[163,1424,173,1458]" captionTargetPageId="9" captionText="FIGURE 5. Photographs in life of Tympanocryptis cephalus species-group members (left to right, top to bottom): a) T. cephalus (photo—G. Harold), b) T. gigas (M. Peterson), c) T. diabolicus sp. nov. (M. Peterson), d) T. fortescuensis sp. nov. (G. Harold), e) T. pseudopsephos sp. nov. — plain form (G. Harold), f) T. pseudopsephos sp. nov. — with charcoal wash on dorsum (G. Harold)." httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="24" pageNumber="109">Figs. 5</figureCitation>
|
||
E, 5F), ground color medium to dark reddish-brown, often with dark grayish black wash over head, dorsum, and limbs; in some individuals this is quite extensive (e.g.
|
||
<figureCitation id="439A2A668D47FFEC5290CAA78D3DF935" box="[1267,1333,1713,1738]" captionStart="FIGURE 5" captionStartId="9.[151,250,1479,1501]" captionTargetBox="[176,1412,186,1446]" captionTargetId="figure@9.[163,1424,173,1458]" captionTargetPageId="9" captionText="FIGURE 5. Photographs in life of Tympanocryptis cephalus species-group members (left to right, top to bottom): a) T. cephalus (photo—G. Harold), b) T. gigas (M. Peterson), c) T. diabolicus sp. nov. (M. Peterson), d) T. fortescuensis sp. nov. (G. Harold), e) T. pseudopsephos sp. nov. — plain form (G. Harold), f) T. pseudopsephos sp. nov. — with charcoal wash on dorsum (G. Harold)." httpUri="https://zenodo.org/record/236306/files/figure.png" pageId="24" pageNumber="109">Fig. 5</figureCitation>
|
||
F); limbs weakly banded; nuchal region dark brown with 3 pale stripes; tail with ~8 alternating bands of pale white and ground color, anterior edge of pale bands dark, pale bands sometimes coalescing to form continuous stripe along top of tail towards tip. In preservative, ground color usually a grayish-brown, dark blotching on dorsum and banding on limbs dark gray to brown (
|
||
<figureCitation id="439A2A668D47FFEC5425CB578A91F8A5" box="[582,665,1857,1882]" captionStart="FIGURE 15" captionStartId="23.[151,250,840,862]" captionTargetBox="[151,1436,193,818]" captionTargetId="figure@23.[151,1436,193,818]" captionTargetPageId="23" captionText="FIGURE 15. Paratypes of Tympanocryptis pseudopsephos sp. nov. (scale bar = 1 cm)." httpUri="https://zenodo.org/record/236316/files/figure.png" pageId="24" pageNumber="109">Fig. 15</figureCitation>
|
||
); some individuals with dark stippling near gular fold; pale bands on tail yellowy-white; ventrum pale with yellow or light orange hue (
|
||
<figureCitation id="439A2A668D47FFEC551FCB738BC7F881" box="[892,975,1893,1918]" captionStart="FIGURE 14" captionStartId="22.[151,250,1989,2011]" captionTargetBox="[151,1436,767,1966]" captionTargetId="figure@22.[151,1436,767,1966]" captionTargetPageId="22" captionText="FIGURE 14. Holotype of Tympanocryptis pseudopsephos sp. nov. (scale bar = 1 cm)." httpUri="https://zenodo.org/record/236315/files/figure.png" pageId="24" pageNumber="109">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D47FFEC56A4CB9F8CFFF85D" blockId="24.[151,1436,1677,2026]" box="[199,1271,1929,1954]" pageId="24" pageNumber="109">
|
||
<emphasis id="E9D5EAF18D47FFEC56A4CB9F8922F85D" bold="true" box="[199,298,1929,1954]" pageId="24" pageNumber="109">Habitat.</emphasis>
|
||
Recorded from Mallee and Mulga woodlands, often with stones strewn on the ground.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D47FFED56A4CBBB896AFF07" lastPageId="25" lastPageNumber="110" pageId="24" pageNumber="109" type="distribution">
|
||
<paragraph id="DB1E36E38D47FFED56A4CBBB896AFF07" blockId="24.[151,1436,1677,2026]" lastBlockId="25.[151,1436,151,1437]" lastPageId="25" lastPageNumber="110" pageId="24" pageNumber="109">
|
||
<emphasis id="E9D5EAF18D47FFEC56A4CBBB8968F839" bold="true" box="[199,352,1965,1990]" pageId="24" pageNumber="109">Distribution.</emphasis>
|
||
Widely distributed throughout the Goldfields region of Western
|
||
<collectingCountry id="A3B676738D47FFEC5230CBBB8CB2F839" box="[1107,1210,1965,1990]" name="Australia" pageId="24" pageNumber="109">Australia</collectingCountry>
|
||
(
|
||
<figureCitation id="439A2A668D47FFEC52AFCBBB8D19F839" box="[1228,1297,1965,1990]" captionStart="FIGURE 1" captionStartId="1.[151,250,2001,2023]" captionTargetBox="[177,1407,361,1954]" captionTargetId="figure@1.[151,1435,339,1979]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of the Tympanocryptis cephalus species-group in Western Australia." httpUri="https://zenodo.org/record/236303/files/figure.png" pageId="24" pageNumber="109">Fig. 1</figureCitation>
|
||
). From just outside the south-eastern edge of the Pilbara, then south to near Norseman. Two older western records are from Yalgoo (WAM R13639) and Dongara (WAM R13476), although the latter record is likely in error given the habitat preferences of this species. A recent eastern record (WAM R163342) is from Neale Junction towards the South Australian border.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D46FFED56A4CD12898AFC7F" pageId="25" pageNumber="110" type="etymology">
|
||
<paragraph id="DB1E36E38D46FFED56A4CD128C3BFEBF" blockId="25.[151,1436,151,1437]" pageId="25" pageNumber="110">
|
||
<emphasis id="E9D5EAF18D46FFED56A4CD128946FEE2" bold="true" box="[199,334,260,285]" pageId="25" pageNumber="110">Etymology.</emphasis>
|
||
<taxonomicName id="1CA14D608D46FFED573ACD128A0AFEE2" box="[345,514,260,285]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos">
|
||
<emphasis id="E9D5EAF18D46FFED573ACD128A0AFEE2" box="[345,514,260,285]" italics="true" pageId="25" pageNumber="110">pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
means ‘false pebble’, with
|
||
<emphasis id="E9D5EAF18D46FFED5533CD128B97FEE2" box="[848,927,260,285]" italics="true" pageId="25" pageNumber="110">pseudo</emphasis>
|
||
Latin for ‘false’ and
|
||
<emphasis id="E9D5EAF18D46FFED52C5CD128D08FEE2" box="[1190,1280,260,285]" italics="true" pageId="25" pageNumber="110">psephos</emphasis>
|
||
is Greek for ‘pebble’. The name is an allusion to the pebble-mimicking abilities of this species.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D46FFED56A4CD5A8B63FE78" blockId="25.[151,1436,151,1437]" pageId="25" pageNumber="110">
|
||
<emphasis id="E9D5EAF18D46FFED56A4CD5A8A56FE9A" bold="true" box="[199,606,332,357]" pageId="25" pageNumber="110">Comparisons with other species.</emphasis>
|
||
Based on its widespread distribution, we provide comparisons of
|
||
<taxonomicName id="1CA14D608D46FFED53E4CD5B8904FE77" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="diabolicus" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED53E4CD5B8904FE77" italics="true" pageId="25" pageNumber="110">T. diabolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5771CD798964FE77" box="[274,364,367,392]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5771CD798964FE77" bold="true" box="[274,364,367,392]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
to all species treated here except
|
||
<taxonomicName id="1CA14D608D46FFED5486CD678B6CFE77" box="[741,868,367,392]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D46FFED5486CD678B6CFE77" box="[741,868,367,392]" italics="true" pageId="25" pageNumber="110">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D46FFED56A4CD838AC7FDE7" blockId="25.[151,1436,151,1437]" pageId="25" pageNumber="110">
|
||
<taxonomicName id="1CA14D608D46FFED56A4CD838A22FE52" box="[199,554,404,429]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED56A4CD838A22FE52" box="[199,554,404,429]" italics="true" pageId="25" pageNumber="110">Tympanocryptis pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5453CD828A81FE52" box="[560,649,404,429]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5453CD828A81FE52" bold="true" box="[560,649,404,429]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName id="1CA14D608D46FFED55E1CD838BD2FE53" box="[898,986,405,428]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="gigas">
|
||
<emphasis id="E9D5EAF18D46FFED55E1CD838BD2FE53" box="[898,986,405,428]" italics="true" pageId="25" pageNumber="110">T. gigas</emphasis>
|
||
</taxonomicName>
|
||
by smaller body size, more rotund body shape, concave (
|
||
<emphasis id="E9D5EAF18D46FFED5734CDAF8997FE2F" box="[343,415,441,464]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
convex) snout, possessing enlarged scales on dorsum arranged in short transverse rows (
|
||
<emphasis id="E9D5EAF18D46FFED56C3CDCB88E0FE0B" box="[160,232,477,500]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
slightly enlarged scales scattered on dorsum), conspicuous ridge on front of thigh formed by enlarged row of scales, and scales on top of thigh homogeneous.
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D46FFED56A4CE338BD2FD57" blockId="25.[151,1436,151,1437]" pageId="25" pageNumber="110">
|
||
<taxonomicName id="1CA14D608D46FFED56A4CE338A24FDC2" box="[199,556,548,573]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED56A4CE338A24FDC2" box="[199,556,548,573]" italics="true" pageId="25" pageNumber="110">Tympanocryptis pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5456CE328A99FDC2" box="[565,657,548,573]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5456CE328A99FDC2" bold="true" box="[565,657,548,573]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName id="1CA14D608D46FFED55F6CE338C20FDC2" box="[917,1064,548,573]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="diabolicus" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED55F6CE338C20FDC2" box="[917,1064,548,573]" italics="true" pageId="25" pageNumber="110">T. diabolicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5253CE328C84FDC2" box="[1072,1164,548,573]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5253CE328C84FDC2" bold="true" box="[1072,1164,548,573]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by possessing enlarged dorsal scales in short transverse rows in rows of 4–7 scales (
|
||
<emphasis id="E9D5EAF18D46FFED5555CE5F8B76FD9F" box="[822,894,585,608]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
2–5), snout concave (versus straight or convex), rostral scale width 2 times height (
|
||
<emphasis id="E9D5EAF18D46FFED547DCE7B8A6DFD7B" box="[542,613,621,644]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
3 times), keels of scales on upper arm not aligned, scales on upper thigh not aligned, and ventrals smooth or slightly raised (
|
||
<emphasis id="E9D5EAF18D46FFED54B4CE878B16FD57" box="[727,798,657,680]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
with low keels).
|
||
</paragraph>
|
||
<paragraph id="DB1E36E38D46FFED56A4CEA3898AFC7F" blockId="25.[151,1436,151,1437]" pageId="25" pageNumber="110">
|
||
<taxonomicName id="1CA14D608D46FFED56A4CEA38A21FD32" box="[199,553,692,717]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED56A4CEA38A21FD32" box="[199,553,692,717]" italics="true" pageId="25" pageNumber="110">Tympanocryptis pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5452CEA28A82FD32" box="[561,650,692,717]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5452CEA28A82FD32" bold="true" box="[561,650,692,717]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is distinguished from
|
||
<taxonomicName id="1CA14D608D46FFED55E6CEA38C3AFD33" box="[901,1074,692,716]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="fortescuensis" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED55E6CEA38C3AFD33" box="[901,1074,692,716]" italics="true" pageId="25" pageNumber="110">T. fortescuensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5259CEA28C9BFD32" box="[1082,1171,692,717]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5259CEA28C9BFD32" bold="true" box="[1082,1171,692,717]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by possessing enlarged dorsal scales in short transverse rows in rows of 4–7 scales (
|
||
<emphasis id="E9D5EAF18D46FFED5554CECF8B76FD0F" box="[823,894,729,752]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
2–5), concave snout (
|
||
<emphasis id="E9D5EAF18D46FFED5216CECF8CB4FD0F" box="[1141,1212,729,752]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
straight to convex), scales on snout rugose with feeble keels (
|
||
<emphasis id="E9D5EAF18D46FFED5415CEEB8AB5FCEB" box="[630,701,765,788]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
with low keels), rostral scale 3 times wider than high (
|
||
<emphasis id="E9D5EAF18D46FFED535ECEEB8D8DFCEB" box="[1341,1413,765,788]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
2 times), keels on dorsal surface of upper arm not aligned, keels of scales on top of thigh not aligned (
|
||
<emphasis id="E9D5EAF18D46FFED529FCF378D4CFCC7" box="[1276,1348,801,824]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
usually aligned), ventrals smooth or slightly raised (
|
||
<emphasis id="E9D5EAF18D46FFED54F2CF538AD0FCA3" box="[657,728,837,860]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
with low keels), and reddish-brown color overlain with black (
|
||
<emphasis id="E9D5EAF18D46FFED56C3CF7F88E0FC7F" box="[160,232,873,896]" italics="true" pageId="25" pageNumber="110">versus</emphasis>
|
||
light brown).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="93BB65688D46FFED56A4CF9A8A72FA62" pageId="25" pageNumber="110" type="discussion">
|
||
<paragraph id="DB1E36E38D46FFED56A4CF9A8A72FA62" blockId="25.[151,1436,151,1437]" pageId="25" pageNumber="110">
|
||
<emphasis id="E9D5EAF18D46FFED56A4CF9A8933FC5A" bold="true" box="[199,315,908,933]" pageId="25" pageNumber="110">Remarks.</emphasis>
|
||
<taxonomicName id="1CA14D608D46FFED5720CF9B8AA0FC5A" box="[323,680,908,933]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED5720CF9B8AA0FC5A" box="[323,680,908,933]" italics="true" pageId="25" pageNumber="110">Tympanocryptis pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED54D3CF9A8B03FC5A" box="[688,779,908,933]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED54D3CF9A8B03FC5A" bold="true" box="[688,779,908,933]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is usually the taxon depicted in field guides (e.g. Storr
|
||
<emphasis id="E9D5EAF18D46FFED53EBCF9B88BCFC37" italics="true" pageId="25" pageNumber="110">et al.</emphasis>
|
||
1983; Wilson & Knowles 1988; Wilson & Swan 2003; Cogger 2014), as this species is the most widelydistributed of the
|
||
<taxonomicName id="1CA14D608D46FFED571FCFC38A0FFC12" box="[380,519,980,1005]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D46FFED571FCFC38A0FFC12" box="[380,519,980,1005]" italics="true" pageId="25" pageNumber="110">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
species-group members and therefore most likely to be encountered and photographed. This also extends to diagnoses and descriptions in such guides and keys, and also often in taxonomic treatments as well. For example, in Mitchell’s 1948 revision of the group, the specimen WAM R7067 figured under the
|
||
<taxonomicName id="1CA14D608D46FFED56A3C85789A2FBA7" box="[192,426,1087,1112]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="subSpecies" species="cephalus" subSpecies="cephalus">
|
||
<emphasis id="E9D5EAF18D46FFED56A3C85789A2FBA7" box="[192,426,1087,1112]" italics="true" pageId="25" pageNumber="110">T. cephalus cephalus</emphasis>
|
||
</taxonomicName>
|
||
section is a
|
||
<taxonomicName id="1CA14D608D46FFED5457C8578AF0FBA7" box="[564,760,1087,1112]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED5457C8578AF0FBA7" box="[564,760,1087,1112]" italics="true" pageId="25" pageNumber="110">T. pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED5563C8298B51FBA7" box="[768,857,1087,1112]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED5563C8298B51FBA7" bold="true" box="[768,857,1087,1112]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
from
|
||
<quantity id="1C599B068D46FFED55FEC8568BECFBA7" box="[925,996,1087,1112]" metricMagnitude="4" metricUnit="m" metricValue="4.8" pageId="25" pageNumber="110" unit="km" value="48.0">48 km</quantity>
|
||
east of Kalgoorlie. Moreover, Mitchell commented that his description of the ‘inland form’ (i.e.
|
||
<taxonomicName id="1CA14D608D46FFED5540C8728BC5FB82" box="[803,973,1124,1149]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos">
|
||
<emphasis id="E9D5EAF18D46FFED5540C8728BC5FB82" box="[803,973,1124,1149]" italics="true" pageId="25" pageNumber="110">pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
) differed slightly from that in Günther’s (1867) original description, Boulenger’s (1885) figure, and photographs of the
|
||
<typeStatus id="041A88418D46FFED5245C89E8C69FB5F" box="[1062,1121,1160,1184]" pageId="25" pageNumber="110">types</typeStatus>
|
||
sent to him by the Natural History Museum: ‘…the [two
|
||
<typeStatus id="041A88418D46FFED5794C8BB8A59FB3A" box="[503,593,1197,1221]" pageId="25" pageNumber="110" type="syntype">syntype</typeStatus>
|
||
] specimens...differ in scalation detail from inland specimens. It has been suggested to me by a fellow worker that this inland form should be described as a new race. However, after considering that no similar specimens have been taken, it is assumed that the
|
||
<typeStatus id="041A88418D46FFED5244C8E38C5FFAF2" box="[1063,1111,1269,1293]" pageId="25" pageNumber="110">type</typeStatus>
|
||
specimens are merely local variants, and not typical representatives of the species which Gunther in fact described.’ (p. 64). We now know that the
|
||
<typeStatus id="041A88418D46FFED56A4C92B8921FAAA" box="[199,297,1341,1365]" pageId="25" pageNumber="110" type="syntype">syntypes</typeStatus>
|
||
were of true
|
||
<taxonomicName id="1CA14D608D46FFED57BBC92B8A56FAAA" box="[472,606,1340,1365]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="cephalus">
|
||
<emphasis id="E9D5EAF18D46FFED57BBC92B8A56FAAA" box="[472,606,1340,1365]" italics="true" pageId="25" pageNumber="110">T. cephalus</emphasis>
|
||
</taxonomicName>
|
||
and that the ‘inland form’ is the distinct
|
||
<taxonomicName id="1CA14D608D46FFED5200C92B8D27FAAA" box="[1123,1327,1340,1365]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="species" species="pseudopsephos" status="sp. nov.">
|
||
<emphasis id="E9D5EAF18D46FFED5200C92B8D27FAAA" box="[1123,1327,1340,1365]" italics="true" pageId="25" pageNumber="110">T. pseudopsephos</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F2E6578A8D46FFED535FC92A8D94FAAA" box="[1340,1436,1340,1365]" pageId="25" pageNumber="110" rank="species">
|
||
<emphasis id="E9D5EAF18D46FFED535FC92A8D94FAAA" bold="true" box="[1340,1436,1340,1365]" pageId="25" pageNumber="110">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Historically, a lack of specimens has hindered taxonomic understanding in
|
||
<taxonomicName id="1CA14D608D46FFED558FC9778C96FA87" box="[1004,1182,1377,1400]" class="Reptilia" family="Agamidae" genus="Tympanocryptis" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="110" phylum="Chordata" rank="genus">
|
||
<emphasis id="E9D5EAF18D46FFED558FC9778C96FA87" box="[1004,1182,1377,1400]" italics="true" pageId="25" pageNumber="110">Tympanocryptis</emphasis>
|
||
</taxonomicName>
|
||
, and indeed continues to hinder revision of other arid zone forms.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |