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<mods:title id="F48F2A666ABB6837A965822B4729402D">Ommatoiulus malleatus n. sp., a new Tunisian millipede, with notes on the punicus species group of Ommatoiulus (Diplopoda, Julidae)</mods:title>
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<mods:date id="CBD1509970FF6432A079AA1919E78F43">2007</mods:date>
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<paragraph id="C77B36AAFFABFFCFFF2EF99BCC61F9DB" blockId="7.[151,559,1613,1639]" box="[151,559,1613,1639]" pageId="7" pageNumber="66">
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<emphasis id="F5B0EAB8FFABFFCFFF2EF99BCC61F9DB" bold="true" box="[151,559,1613,1639]" pageId="7" pageNumber="66">
Key to the
<taxonomicName id="00C44D29FFABFFCFFEA6F99BCF31F9DB" ID-CoL="49CR9" box="[287,383,1613,1639]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="punicus">
<emphasis id="F5B0EAB8FFABFFCFFEA6F99BCF31F9DB" bold="true" box="[287,383,1613,1639]" italics="true" pageId="7" pageNumber="66">punicus</emphasis>
</taxonomicName>
species group
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8FDE6521FFABFFC0FF2EF94ECBD7FEEE" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66" type="key">
<keyStep id="7C302E0FFFABFFCFFF2EF94ECBD4F8BE" pageId="7" pageNumber="66">
<paragraph id="C77B36AAFFABFFCFFF2EF94ECBD4F966" blockId="7.[151,1437,1688,2034]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF94ECBD4F966" pageId="7" pageNumber="66">1 Promerite strongly narrower on its distal third to half (may be reduced to a simple digit­shaped extension) ..................................................................................................................................................................... 2</keyLead>
</paragraph>
<paragraph id="C77B36AAFFABFFCFFF2EF93ECBD4F8BE" blockId="7.[151,1437,1688,2034]" box="[151,1434,1768,1794]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF93ECBD4F8BE" box="[151,1434,1768,1794]" pageId="7" pageNumber="66">1* Promerite not or barely narrower distally .................................................................................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C302E0FFFABFFCFFF2EF8C6CBD4F8EE" pageId="7" pageNumber="66">
<paragraph id="C77B36AAFFABFFCFFF2EF8C6CBD4F895" blockId="7.[151,1437,1688,2034]" box="[151,1434,1807,1834]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF8C6CBD4F895" box="[151,1434,1807,1834]" pageId="7" pageNumber="66">
2 Hammer­shaped mesomerite (
<figureCitation id="5FFF2A2FFFABFFCFFD9EF8C6CC23F896" box="[551,621,1808,1834]" captionStart="FIGURES 1 5" captionStartId="3.[151,269,1520,1544]" captionTargetBox="[222,1414,389,1492]" captionTargetId="figure@3.[86,1482,379,1516]" captionTargetPageId="3" captionText="FIGURES 1 5. Ommatoiulus malleatus n. sp., ♂, paratypes: 1. head and collum (lateral); 2. first right leg (lateral); 3. right promerite (anterior), fd folds, l lateral lobe, lpb lateral pointed bump, m median lobe, mrb median rounded bump, n notch, p 1 median protuberances, p 2 lateral protuberance; 4. left promerite (posterior); 5. posterior gonopod (posterior), hy hyaline sheet, ms mesomerite, op opisthomerite, px paracoxite, s solenomerite." httpUri="https://zenodo.org/record/175411/files/figure.png" pageId="7" pageNumber="66">Fig. 5</figureCitation>
)
<emphasis id="F5B0EAB8FFABFFCFFDC4F8C6CB1EF895" box="[637,1360,1808,1833]" italics="true" pageId="7" pageNumber="66">
............................................................
<taxonomicName id="00C44D29FFABFFCFFB83F8C6CB1EF895" ID-CoL="49CQT" box="[1082,1360,1808,1833]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="malleatus" status="sp. nov.">Ommatoiulus malleatus</taxonomicName>
</emphasis>
<taxonomicNameLabel id="EE8357C3FFABFFCFFAE1F8D9CBD4F895" box="[1368,1434,1807,1833]" pageId="7" pageNumber="66" rank="species">
<emphasis id="F5B0EAB8FFABFFCFFAE1F8D9CBD4F895" bold="true" box="[1368,1434,1807,1833]" pageId="7" pageNumber="66">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="C77B36AAFFABFFCFFF2EF8EECBD4F8EE" blockId="7.[151,1437,1688,2034]" box="[151,1434,1848,1874]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF8EECBD4F8EE" box="[151,1434,1848,1874]" pageId="7" pageNumber="66">2* Different shaped mesomerite ....................................................................................................................... 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C302E0FFFABFFCFFF2EF8B6CBD7F81E" pageId="7" pageNumber="66">
<paragraph id="C77B36AAFFABFFCFFF2EF8B6CBD7F8C6" blockId="7.[151,1437,1688,2034]" box="[151,1433,1888,1914]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF8B6CBD7F8C6" box="[151,1433,1888,1914]" pageId="7" pageNumber="66">
3 Solenomerite long (
<figureCitation id="5FFF2A2FFFABFFCFFE17F8B6CC4DF8C6" box="[430,515,1888,1914]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1428,1452]" captionTargetBox="[187,1348,182,1427]" captionTargetId="figure@7.[171,1417,173,1438]" captionTargetPageId="7" captionText="FIGURES 12 17. Posterior gonopod: 12. O. punicus; 13. O. seurati (redrawn after Brolemann 1925 a); 14. O. aumalensis (redrawn after Brolemann 1925 b); 15. O. brandti (redrawn after Berlese 1885); 16. O. atlanteus (redrawn after Schubart 1960); 17. O. quadridentatus (redrawn after Schubart 1960)." httpUri="https://zenodo.org/record/175413/files/figure.png" pageId="7" pageNumber="66">Fig. 12</figureCitation>
)
<emphasis id="F5B0EAB8FFABFFCFFDB6F8B6CDFFF8C5" box="[527,945,1888,1913]" italics="true" pageId="7" pageNumber="66">.........................................................</emphasis>
<taxonomicName id="00C44D29FFABFFCFFC0AF8B6CBD7F8C6" ID-CoL="49CR9" authority="(Brolemann, 1894)" authorityName="Brolemann" authorityYear="1894" baseAuthorityName="Brolemann" baseAuthorityYear="1894" box="[947,1433,1888,1914]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="punicus">
<emphasis id="F5B0EAB8FFABFFCFFC0AF8B6CAFAF8C5" box="[947,1204,1888,1913]" italics="true" pageId="7" pageNumber="66">Ommatoiulus punicus</emphasis>
(
<bibRefCitation id="A3554B5BFFABFFCFFB7BF8B6CBDFF8C6" author="Brolemann" box="[1218,1425,1888,1914]" pageId="7" pageNumber="67" refString="Brolemann, H. W. (1894) Contribution a la faune myriapodologique mediterraneenne. Memoires de la Societe Zooogique de France, VII, 437 - 462." type="book chapter" year="1894">Brölemann, 1894</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C77B36AAFFABFFCFFF2EF85ECBD7F81E" blockId="7.[151,1437,1688,2034]" box="[151,1433,1928,1954]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF85ECBD7F81E" box="[151,1433,1928,1954]" pageId="7" pageNumber="66">
3* Solenomerite short (
<figureCitation id="5FFF2A2FFFABFFCFFE0DF85ECC47F81E" box="[436,521,1928,1954]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1428,1452]" captionTargetBox="[187,1348,182,1427]" captionTargetId="figure@7.[171,1417,173,1438]" captionTargetPageId="7" captionText="FIGURES 12 17. Posterior gonopod: 12. O. punicus; 13. O. seurati (redrawn after Brolemann 1925 a); 14. O. aumalensis (redrawn after Brolemann 1925 b); 15. O. brandti (redrawn after Berlese 1885); 16. O. atlanteus (redrawn after Schubart 1960); 17. O. quadridentatus (redrawn after Schubart 1960)." httpUri="https://zenodo.org/record/175413/files/figure.png" pageId="7" pageNumber="66">Fig. 14</figureCitation>
)
<emphasis id="F5B0EAB8FFABFFCFFDAFF85ECDCAF81D" box="[534,900,1928,1953]" italics="true" pageId="7" pageNumber="66">..................................................</emphasis>
<taxonomicName id="00C44D29FFABFFCFFC32F85ECBD7F81E" ID-CoL="49CPX" authority="(Brolemann, 1925)" authorityName="Brolemann" authorityYear="1925" baseAuthorityName="Brolemann" baseAuthorityYear="1925" box="[907,1433,1928,1954]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="aumalensis">
<emphasis id="F5B0EAB8FFABFFCFFC32F85ECAFDF81D" box="[907,1203,1928,1953]" italics="true" pageId="7" pageNumber="66">Ommatoiulus aumalensis</emphasis>
(Brolemann, 1925)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C302E0FFFABFFC0FF2EF866CBD4FF0E" lastPageId="8" lastPageNumber="67" pageId="7" pageNumber="66">
<paragraph id="C77B36AAFFABFFCFFF2EF866CBD4F84E" blockId="7.[151,1437,1688,2034]" pageId="7" pageNumber="66">
<keyLead id="7C35959FFFABFFCFFF2EF866CBD4F84E" pageId="7" pageNumber="66">
4 Mesomerite particularly broad distally with fan­shaped apex (
<figureCitation id="5FFF2A2FFFABFFCFFC20F866CDA4F876" box="[921,1002,1968,1994]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1428,1452]" captionTargetBox="[187,1348,182,1427]" captionTargetId="figure@7.[171,1417,173,1438]" captionTargetPageId="7" captionText="FIGURES 12 17. Posterior gonopod: 12. O. punicus; 13. O. seurati (redrawn after Brolemann 1925 a); 14. O. aumalensis (redrawn after Brolemann 1925 b); 15. O. brandti (redrawn after Berlese 1885); 16. O. atlanteus (redrawn after Schubart 1960); 17. O. quadridentatus (redrawn after Schubart 1960)." httpUri="https://zenodo.org/record/175413/files/figure.png" pageId="7" pageNumber="66">Fig. 16</figureCitation>
)........................................................
<emphasis id="F5B0EAB8FFABFFCFFF75F80ECD80F84D" box="[204,974,2008,2033]" italics="true" pageId="7" pageNumber="66">.........................................................................................................</emphasis>
<taxonomicName id="00C44D29FFABFFCFFC68F80ECBD4F84E" ID-CoL="49CPW" authority="Schubart, 1960" authorityName="Schubart" authorityYear="1960" box="[977,1434,2008,2034]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="7" pageNumber="66" phylum="Arthropoda" rank="species" species="atlanteus">
<emphasis id="F5B0EAB8FFABFFCFFC68F80ECAAFF84D" box="[977,1249,2008,2033]" italics="true" pageId="7" pageNumber="66">Ommatoiulus atlanteus</emphasis>
<bibRefCitation id="A3554B5BFFABFFCFFB51F80ECBD4F84E" author="Schubart" box="[1256,1434,2008,2034]" pageId="7" pageNumber="68" refString="Schubart, O. (1960) Ein weiterer Beitrag zur Diplopoden-Fauna Marokkos. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 40 (3), 159 - 232." type="journal article" year="1960">Schubart, 1960</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C77B36AAFFA4FFC0FF2EFF4ECBD4FF0E" blockId="8.[151,1434,152,338]" box="[151,1434,152,178]" pageId="8" pageNumber="67">
<keyLead id="7C35959FFFA4FFC0FF2EFF4ECBD4FF0E" box="[151,1434,152,178]" pageId="8" pageNumber="67">4* Mesomerite not or less broad distally without fan­shaped apex.................................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C302E0FFFA4FFC0FF2EFF16CBD4FEBE" pageId="8" pageNumber="67">
<paragraph id="C77B36AAFFA4FFC0FF2EFF16CBD4FF66" blockId="8.[151,1434,152,338]" box="[151,1434,192,218]" pageId="8" pageNumber="67">
<keyLead id="7C35959FFFA4FFC0FF2EFF16CBD4FF66" box="[151,1434,192,218]" pageId="8" pageNumber="67">5 Mesomerite large, slightly broader distally ................................................................................................. 6</keyLead>
</paragraph>
<paragraph id="C77B36AAFFA4FFC0FF2EFF3ECBD4FEBE" blockId="8.[151,1434,152,338]" box="[151,1434,232,258]" pageId="8" pageNumber="67">
<keyLead id="7C35959FFFA4FFC0FF2EFF3ECBD4FEBE" box="[151,1434,232,258]" pageId="8" pageNumber="67">
5* Mesomerite slender, slightly narrower distally (
<figureCitation id="5FFF2A2FFFA4FFC0FD4BFF3ECD09FEBE" box="[754,839,232,258]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1428,1452]" captionTargetBox="[187,1348,182,1427]" captionTargetId="figure@7.[171,1417,173,1438]" captionTargetPageId="7" captionText="FIGURES 12 17. Posterior gonopod: 12. O. punicus; 13. O. seurati (redrawn after Brolemann 1925 a); 14. O. aumalensis (redrawn after Brolemann 1925 b); 15. O. brandti (redrawn after Berlese 1885); 16. O. atlanteus (redrawn after Schubart 1960); 17. O. quadridentatus (redrawn after Schubart 1960)." httpUri="https://zenodo.org/record/175413/files/figure.png" pageId="8" pageNumber="67">Fig. 17</figureCitation>
)
<emphasis id="F5B0EAB8FFA4FFC0FCE8FF3ECDCBFEBD" box="[849,901,232,257]" italics="true" pageId="8" pageNumber="67">.......</emphasis>
<taxonomicName id="00C44D29FFA4FFC0FC32FF3ECBD4FEBE" ID-CoL="74M55" authority="Schubart, 1960" authorityName="Schubart" authorityYear="1960" box="[907,1434,232,258]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="quadridentatus">
<emphasis id="F5B0EAB8FFA4FFC0FC32FF3ECAAFFEBD" box="[907,1249,232,257]" italics="true" pageId="8" pageNumber="67">Ommatoiulus quadridentatus</emphasis>
<bibRefCitation id="A3554B5BFFA4FFC0FB51FF3ECBD4FEBE" author="Schubart" box="[1256,1434,232,258]" pageId="8" pageNumber="68" refString="Schubart, O. (1960) Ein weiterer Beitrag zur Diplopoden-Fauna Marokkos. Bulletin de la Societe des Sciences Naturelles et Physiques du Maroc, 40 (3), 159 - 232." type="journal article" year="1960">Schubart, 1960</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="7C302E0FFFA4FFC0FF2EFEC6CBD7FEEE" pageId="8" pageNumber="67">
<paragraph id="C77B36AAFFA4FFC0FF2EFEC6CBD7FE96" blockId="8.[151,1434,152,338]" box="[151,1433,272,298]" pageId="8" pageNumber="67">
<keyLead id="7C35959FFFA4FFC0FF2EFEC6CBD7FE96" box="[151,1433,272,298]" pageId="8" pageNumber="67">
6 Paracoxite long with hook­shaped apex (
<figureCitation id="5FFF2A2FFFA4FFC0FD1DFEC6CCB7FE96" box="[676,761,272,298]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1428,1452]" captionTargetBox="[187,1348,182,1427]" captionTargetId="figure@7.[171,1417,173,1438]" captionTargetPageId="7" captionText="FIGURES 12 17. Posterior gonopod: 12. O. punicus; 13. O. seurati (redrawn after Brolemann 1925 a); 14. O. aumalensis (redrawn after Brolemann 1925 b); 15. O. brandti (redrawn after Berlese 1885); 16. O. atlanteus (redrawn after Schubart 1960); 17. O. quadridentatus (redrawn after Schubart 1960)." httpUri="https://zenodo.org/record/175413/files/figure.png" pageId="8" pageNumber="67">Fig. 15</figureCitation>
)
<emphasis id="F5B0EAB8FFA4FFC0FCB1FEC6CD93FE95" box="[776,989,272,297]" italics="true" pageId="8" pageNumber="67">.............................</emphasis>
<taxonomicName id="00C44D29FFA4FFC0FC5AFEC6CBD7FE96" ID-CoL="7YYJJ" authority="(Berlese, 1885)" authorityName="Berlese" authorityYear="1885" baseAuthorityName="Berlese" baseAuthorityYear="1885" box="[995,1433,272,298]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="brandti">
<emphasis id="F5B0EAB8FFA4FFC0FC5AFEC6CA90FE95" box="[995,1246,272,297]" italics="true" pageId="8" pageNumber="67">Ommatoiulus brandti</emphasis>
(
<bibRefCitation id="A3554B5BFFA4FFC0FB54FEC6CBDFFE96" author="Berlese" box="[1261,1425,272,298]" pageId="8" pageNumber="67" refString="Berlese, A. (1885) Acari, Myriapoda et Scorpiones hucusque in Italia reperta. Patavini, Vol. III, Fasc. XXI, No. 3." type="book" year="1885">Berlese, 1885</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="C77B36AAFFA4FFC0FF2EFEEECBD7FEEE" blockId="8.[151,1434,152,338]" box="[151,1433,312,338]" pageId="8" pageNumber="67">
<keyLead id="7C35959FFFA4FFC0FF2EFEEECBD7FEEE" box="[151,1433,312,338]" pageId="8" pageNumber="67">
6* Paracoxite short with rounded apex (
<figureCitation id="5FFF2A2FFFA4FFC0FDCCFEEECC84FEEE" box="[629,714,312,338]" captionStart="FIGURES 12 17" captionStartId="7.[151,269,1428,1452]" captionTargetBox="[187,1348,182,1427]" captionTargetId="figure@7.[171,1417,173,1438]" captionTargetPageId="7" captionText="FIGURES 12 17. Posterior gonopod: 12. O. punicus; 13. O. seurati (redrawn after Brolemann 1925 a); 14. O. aumalensis (redrawn after Brolemann 1925 b); 15. O. brandti (redrawn after Berlese 1885); 16. O. atlanteus (redrawn after Schubart 1960); 17. O. quadridentatus (redrawn after Schubart 1960)." httpUri="https://zenodo.org/record/175413/files/figure.png" pageId="8" pageNumber="67">Fig. 13</figureCitation>
)
<emphasis id="F5B0EAB8FFA4FFC0FD6CFEEECDF7FEED" box="[725,953,312,337]" italics="true" pageId="8" pageNumber="67">...............................</emphasis>
<taxonomicName id="00C44D29FFA4FFC0FC07FEEECBD7FEEE" ID-CoL="49CRM" authority="(Brolemann, 1925)" authorityName="Brolemann" authorityYear="1925" baseAuthorityName="Brolemann" baseAuthorityYear="1925" box="[958,1433,312,338]" class="Diplopoda" family="Julidae" genus="Ommatoiulus" kingdom="Animalia" order="Julida" pageId="8" pageNumber="67" phylum="Arthropoda" rank="species" species="seurati">
<emphasis id="F5B0EAB8FFA4FFC0FC07FEEECAFAFEED" box="[958,1204,312,337]" italics="true" pageId="8" pageNumber="67">Ommatoiulus seurati</emphasis>
(Brolemann, 1925)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>