treatments-xml/data/4F/24/83/4F248367DB5D5C319ADBFEA636B98710.xml
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<document id="F2D5D1B8ABD55E34E35C6BE24EF779AD" ID-DOI="10.11646/zootaxa.3957.1.8" ID-GBIF-Dataset="d97787ca-3db9-4e8f-956f-d26b955e71d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="236432" ID-ZooBank="EBFD444B-8315-4908-866D-22D30AFB5557" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461059621670" checkinUser="plazi" docAuthor="Özbek, Hasan Hüseyin &amp; Halliday, Bruce" docDate="2015" docId="4F248367DB5D5C319ADBFEA636B98710" docLanguage="en" docName="zt03957p108.pdf" docOrigin="Zootaxa 3957 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachyseius destitutus Özbek &amp; Halliday, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="101" masterDocId="B31DFB1FDB5C5C329A4CFFC934468300" masterDocTitle="Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species" masterLastPageNumber="108" masterPageNumber="98" pageNumber="99" updateTime="1698581885510" updateUser="plazi">
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<mods:title id="493042F5796AAC5CF7B94DE076B79D2F">Two new species of Pachyseius Berlese (Acari: Pachylaelapidae) from Turkey, with a key to the world species</mods:title>
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<mods:name id="3F62368E22BCD6DCC2B7D238AF73DA89" type="personal">
<mods:role id="8366D0A305DF1D47F4EC3F51EAFBDACB">
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<mods:namePart id="924E8523D2487A24615F88AE3F80FD39">Özbek, Hasan Hüseyin</mods:namePart>
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<mods:namePart id="499D521BEEED1374B29F7E4EF1E558CB">Halliday, Bruce</mods:namePart>
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<mods:title id="CC056779F019227EE1F9E4B3835AD29F">Zootaxa</mods:title>
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<mods:date id="BF090BA06D6CEF8769553F09A6448F09">2015</mods:date>
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<treatment id="4F248367DB5D5C319ADBFEA636B98710" ID-DOI="http://doi.org/10.5281/zenodo.6093707" ID-GBIF-Taxon="119613792" ID-Zenodo-Dep="6093707" LSID="urn:lsid:plazi:treatment:4F248367DB5D5C319ADBFEA636B98710" httpUri="http://treatment.plazi.org/id/4F248367DB5D5C319ADBFEA636B98710" lastPageId="3" lastPageNumber="101" pageId="1" pageNumber="99">
<subSubSection id="8F9761FADB5D5C339ADBFEA6351982AB" pageId="1" pageNumber="99" type="nomenclature">
<paragraph id="C7323271DB5D5C339ADBFEA635B8828A" blockId="1.[151,510,367,427]" box="[151,510,367,394]" pageId="1" pageNumber="99">
<heading id="9C7A851DDB5D5C339ADBFEA635B8828A" bold="true" box="[151,510,367,394]" fontSize="11" level="1" pageId="1" pageNumber="99" reason="1">
<emphasis id="F5F9EE63DB5D5C339ADBFEA635B8828A" bold="true" box="[151,510,367,394]" pageId="1" pageNumber="99">
<taxonomicName id="008D49F2DB5D5C339ADBFEA635DF8289" ID-CoL="8Y7N9" box="[151,409,367,393]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="destitutus" status="sp. nov.">
<emphasis id="F5F9EE63DB5D5C339ADBFEA635DF8289" bold="true" box="[151,409,367,393]" italics="true" pageId="1" pageNumber="99">Pachyseius destitutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EECA5318DB5D5C339BECFEB935B8828A" box="[416,510,368,394]" pageId="1" pageNumber="99" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C7323271DB5D5C339ADBFE5B351982AB" blockId="1.[151,510,367,427]" box="[151,351,402,427]" pageId="1" pageNumber="99">
(
<figureCitation id="5FB62EF4DB5D5C339AD3FE5B354282AB" box="[159,260,402,427]" captionStart-0="FIGURES 1 2" captionStart-1="FIGURES 3 6" captionStartId-0="1.[151,264,1900,1923]" captionStartId-1="2.[151,264,1361,1384]" captionTargetBox-0="[163,1418,772,1869]" captionTargetBox-1="[151,1429,201,1325]" captionTargetId-0="figure@1.[151,1436,767,1878]" captionTargetId-1="figure@2.[151,1436,193,1340]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURES 1 2. Pachyseius destitutus sp. nov., female. 1. Dorsal shield; 2. Ventral idiosoma." captionText-1="FIGURES 3 6. Pachyseius destitutus sp. nov., female. 3. Ventral gnathosoma; 4. Epistome; 5. Chelicera; 6. Tarsus II." httpUri-0="https://zenodo.org/record/236433/files/figure.png" httpUri-1="https://zenodo.org/record/236434/files/figure.png" pageId="1" pageNumber="99">Figs 16</figureCitation>
,
<figureCitation id="5FB62EF4DB5D5C339B43FE5A351182AB" box="[271,343,403,427]" captionStart="FIGURES 19 22" captionStartId="8.[151,264,1740,1763]" captionTargetBox="[327,1259,501,1719]" captionTargetId="figure@8.[327,1259,501,1720]" captionTargetPageId="8" captionText="FIGURES 19 22. Pachyseius spp., females. 19. P. destitutus sp. nov., anterior region of dorsal idiosoma; 20. P. destitutus sp. nov., ventri-anal shield; 21. P. destitutus sp. nov., ventral gnathosoma; 22. P. quadrigeminus sp. nov., ventri-anal shield. Not scaled." httpUri="https://zenodo.org/record/236438/files/figure.png" pageId="1" pageNumber="99">1921</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="8F9761FADB5D5C309ADBFE1330C986CB" lastPageId="2" lastPageNumber="100" pageId="1" pageNumber="99" type="diagnosis">
<paragraph id="C7323271DB5D5C339ADBFE1331DB81CA" blockId="1.[151,1437,474,715]" pageId="1" pageNumber="99">
<emphasis id="F5F9EE63DB5D5C339ADBFE13355782F3" bold="true" box="[151,273,474,499]" pageId="1" pageNumber="99">Diagnosis.</emphasis>
Dorsal shield bearing 29 pairs of long and pointed setae,
<emphasis id="F5F9EE63DB5D5C3399DEFE1237ED82F3" box="[914,939,475,499]" italics="true" pageId="1" pageNumber="99">z1</emphasis>
absent; anterior margin of ventri-anal shield conspicuously concave, with three pairs of pre-anal setae, post-anal seta longer than other ventri-anal setae; anterolateral margins of ventrianal shield closely adjacent to relatively long and narrow metapodal shield; presternal shields absent; peritrematal shields fused with exopodal shields, their posterior ends reaching beyond coxae IV; soft integument between peritrematal shields and anterolateral margins of ventri-anal shield with one pair of small oval platelets; epigynal shield reduced in size, relatively small, posterior margin with a transverse row of four elongate platelets and a pair of small oval platelets; lateral and opisthogastric soft integument with 11 pairs of setae.
</paragraph>
<caption id="93F262F9DB5D5C339ADBF8A5300F8482" box="[151,1097,1900,1923]" httpUri="https://zenodo.org/record/236433/files/figure.png" pageId="1" pageNumber="99" targetBox="[163,1418,772,1869]" targetPageId="1">
<paragraph id="C7323271DB5D5C339ADBF8A5300F8482" blockId="1.[151,1097,1900,1923]" box="[151,1097,1900,1923]" pageId="1" pageNumber="99">
<emphasis id="F5F9EE63DB5D5C339ADBF8A5357F8482" bold="true" box="[151,313,1900,1923]" pageId="1" pageNumber="99">FIGURES 12.</emphasis>
<taxonomicName id="008D49F2DB5D5C339B0CF8A4365F8482" box="[320,537,1901,1922]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="1" pageNumber="99" phylum="Arthropoda" rank="species" species="destitutus" status="sp. nov.">
<emphasis id="F5F9EE63DB5D5C339B0CF8A4365F8482" box="[320,537,1901,1922]" italics="true" pageId="1" pageNumber="99">Pachyseius destitutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EECA5318DB5D5C33986CF8A436368482" box="[544,624,1901,1922]" pageId="1" pageNumber="99" rank="species">
<emphasis id="F5F9EE63DB5D5C33986CF8A436368482" bold="true" box="[544,624,1901,1922]" pageId="1" pageNumber="99">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 1. Dorsal shield; 2. Ventral idiosoma.
</paragraph>
</caption>
<caption id="93F262F9DB5E5C309ADBFA9831008667" box="[151,1350,1361,1384]" httpUri="https://zenodo.org/record/236434/files/figure.png" pageId="2" pageNumber="100" targetBox="[151,1429,201,1325]" targetPageId="2">
<paragraph id="C7323271DB5E5C309ADBFA9831008667" blockId="2.[151,1350,1361,1384]" box="[151,1350,1361,1384]" pageId="2" pageNumber="100">
<emphasis id="F5F9EE63DB5E5C309ADBFA98357F8666" bold="true" box="[151,313,1361,1384]" pageId="2" pageNumber="100">FIGURES 36.</emphasis>
<taxonomicName id="008D49F2DB5E5C309B0CFA9B365F8667" box="[320,537,1362,1383]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="2" pageNumber="100" phylum="Arthropoda" rank="species" species="destitutus" status="sp. nov.">
<emphasis id="F5F9EE63DB5E5C309B0CFA9B365F8667" box="[320,537,1362,1383]" italics="true" pageId="2" pageNumber="100">Pachyseius destitutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EECA5318DB5E5C30986CFA9B36368667" box="[544,624,1362,1383]" pageId="2" pageNumber="100" rank="species">
<emphasis id="F5F9EE63DB5E5C30986CFA9B36368667" bold="true" box="[544,624,1362,1383]" pageId="2" pageNumber="100">sp. nov.</emphasis>
</taxonomicNameLabel>
, female. 3. Ventral gnathosoma; 4. Epistome; 5. Chelicera; 6. Tarsus II.
</paragraph>
</caption>
<paragraph id="C7323271DB5E5C309A8BFA4730C986CB" blockId="2.[151,1437,1422,2023]" pageId="2" pageNumber="100">
<emphasis id="F5F9EE63DB5E5C309A8BFA47358186A7" bold="true" box="[199,455,1422,1447]" pageId="2" pageNumber="100">Specimens examined.</emphasis>
<typeStatus id="18368CD3DB5E5C309B81FA47367386A7" box="[461,565,1422,1447]" pageId="2" pageNumber="100" type="holotype">Holotype</typeStatus>
female, Trabzon Province, Demirkapı Village,
<geoCoordinate id="A2B954B6DB5E5C309E00FA46309F86A7" box="[1100,1241,1423,1447]" direction="north" orientation="latitude" pageId="2" pageNumber="100" precision="15" value="40.573055">40°34'23'' N</geoCoordinate>
,
<geoCoordinate id="A2B954B6DB5E5C309EA8FA47312886A7" box="[1252,1390,1422,1447]" direction="east" orientation="longitude" pageId="2" pageNumber="100" precision="15" value="40.40167">40°24'06'' E</geoCoordinate>
, alt.
<quantity id="00759F94DB5E5C309ADBFA7A34A886CA" box="[151,238,1459,1483]" metricMagnitude="3" metricUnit="m" metricValue="1.7399999999999998" pageId="2" pageNumber="100" unit="m" value="1740.0">1740 m</quantity>
,
<date id="B33314B1DB5E5C309AB6FA7A35D486CB" box="[250,402,1459,1483]" pageId="2" pageNumber="100" value="2014-06-17">17 June 2014</date>
, litter under
<taxonomicName id="008D49F2DB5E5C309853FA7A361986CA" box="[543,607,1459,1482]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="2" pageNumber="100" phylum="Tracheophyta" rank="genus">
<emphasis id="F5F9EE63DB5E5C309853FA7A361986CA" box="[543,607,1459,1482]" italics="true" pageId="2" pageNumber="100">Pinus</emphasis>
</taxonomicName>
sp.
<typeStatus id="18368CD3DB5E5C3098C1FA7A36B986CB" box="[653,767,1459,1483]" pageId="2" pageNumber="100" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="D18BF9F8DB5E5C309945FA7B37C086CB" box="[777,902,1458,1483]" pageId="2" pageNumber="100" type="female">16 females</specimenCount>
, same data as
<typeStatus id="18368CD3DB5E5C309E6AFA7B30CA86CB" box="[1062,1164,1458,1483]" pageId="2" pageNumber="100" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="8F9761FADB5E5C319A8BFA1F315081A8" lastPageId="3" lastPageNumber="101" pageId="2" pageNumber="100" type="description">
<paragraph id="C7323271DB5E5C309A8BFA1F37C8857F" blockId="2.[151,1437,1422,2023]" pageId="2" pageNumber="100">
<emphasis id="F5F9EE63DB5E5C309A8BFA1F356386EF" bold="true" box="[199,293,1494,1519]" pageId="2" pageNumber="100">Female.</emphasis>
<emphasis id="F5F9EE63DB5E5C309B62FA1135CE86EF" box="[302,392,1496,1519]" italics="true" pageId="2" pageNumber="100">Dorsum</emphasis>
(
<figureCitation id="5FB62EF4DB5E5C309BD5FA1F359B86EF" box="[409,477,1494,1519]" captionStart="FIGURES 1 2" captionStartId="1.[151,264,1900,1923]" captionTargetBox="[163,1418,772,1869]" captionTargetId="figure@1.[151,1436,767,1878]" captionTargetPageId="1" captionText="FIGURES 1 2. Pachyseius destitutus sp. nov., female. 1. Dorsal shield; 2. Ventral idiosoma." httpUri="https://zenodo.org/record/236433/files/figure.png" pageId="2" pageNumber="100">Fig. 1</figureCitation>
). Dorsal shield 485535 long, 275300 wide at widest point, oblong (length/width, 1.701.78), ornamented with net-like pattern, and bearing 29 pairs of mostly long and pointed needle-like setae, setae
<emphasis id="F5F9EE63DB5E5C309A9BF9D634B68537" box="[215,240,1567,1591]" italics="true" pageId="2" pageNumber="100">z1</emphasis>
absent (
<figureCitation id="5FB62EF4DB5E5C309B1DF9D735E08537" box="[337,422,1566,1591]" captionStart="FIGURES 19 22" captionStartId="8.[151,264,1740,1763]" captionTargetBox="[327,1259,501,1719]" captionTargetId="figure@8.[327,1259,501,1720]" captionTargetPageId="8" captionText="FIGURES 19 22. Pachyseius spp., females. 19. P. destitutus sp. nov., anterior region of dorsal idiosoma; 20. P. destitutus sp. nov., ventri-anal shield; 21. P. destitutus sp. nov., ventral gnathosoma; 22. P. quadrigeminus sp. nov., ventri-anal shield. Not scaled." httpUri="https://zenodo.org/record/236438/files/figure.png" pageId="2" pageNumber="100">Fig. 19</figureCitation>
); setae
<emphasis id="F5F9EE63DB5E5C309BB1F9D636528537" box="[509,532,1567,1591]" italics="true" pageId="2" pageNumber="100">j1</emphasis>
shortest, 1012;
<emphasis id="F5F9EE63DB5E5C309895F9D636B68537" box="[729,752,1567,1591]" italics="true" pageId="2" pageNumber="100">j2</emphasis>
3450;
<emphasis id="F5F9EE63DB5E5C309902F9D637228537" box="[846,868,1567,1591]" italics="true" pageId="2" pageNumber="100">j3</emphasis>
,
<emphasis id="F5F9EE63DB5E5C30993FF9D637CA8537" box="[883,908,1567,1591]" italics="true" pageId="2" pageNumber="100">s4</emphasis>
,
<emphasis id="F5F9EE63DB5E5C3099D0F9D637F28536" box="[924,948,1567,1590]" italics="true" pageId="2" pageNumber="100">s5</emphasis>
, and
<emphasis id="F5F9EE63DB5E5C3099B9F9D630558537" box="[1013,1043,1567,1591]" italics="true" pageId="2" pageNumber="100">Z1</emphasis>
7075;
<emphasis id="F5F9EE63DB5E5C309E3EF9D630CD8537" box="[1138,1163,1567,1591]" italics="true" pageId="2" pageNumber="100">z6</emphasis>
6065;
<emphasis id="F5F9EE63DB5E5C309EA6F9D631428536" box="[1258,1284,1567,1590]" italics="true" pageId="2" pageNumber="100">
J
<date id="B33314B1DB5E5C309EBBF9D631428536" box="[1271,1284,1567,1590]" pageId="2" pageNumber="100" value="1922-05-28">5</date>
</emphasis>
2228, other dorsal setae 3560; a pair of lyrifissures near bases of
<emphasis id="F5F9EE63DB5E5C3098B6F98A3756855B" box="[762,784,1603,1627]" italics="true" pageId="2" pageNumber="100">j2</emphasis>
; slit-like gland pores
<emphasis id="F5F9EE63DB5E5C309E49F98A3071855B" box="[1029,1079,1603,1627]" italics="true" pageId="2" pageNumber="100">gdj3</emphasis>
present between seta
<emphasis id="F5F9EE63DB5E5C309F62F98A3102855B" box="[1326,1348,1603,1627]" italics="true" pageId="2" pageNumber="100">j2</emphasis>
and
<emphasis id="F5F9EE63DB5E5C309F37F98A31D2855B" box="[1403,1428,1603,1627]" italics="true" pageId="2" pageNumber="100">s2</emphasis>
; anterior ends of peritremes extending forward beyond level of
<emphasis id="F5F9EE63DB5E5C309919F9AE37C1857F" box="[853,903,1639,1663]" italics="true" pageId="2" pageNumber="100">gdj3</emphasis>
.
</paragraph>
<paragraph id="C7323271DB5E5C319A8BF942372E821D" blockId="2.[151,1437,1422,2023]" lastBlockId="3.[151,1436,151,1040]" lastPageId="3" lastPageNumber="101" pageId="2" pageNumber="100">
<emphasis id="F5F9EE63DB5E5C309A8BF942354A85A2" box="[199,268,1675,1698]" italics="true" pageId="2" pageNumber="100">Venter</emphasis>
(
<figureCitation id="5FB62EF4DB5E5C309B51F943352485A3" box="[285,354,1674,1699]" captionStart="FIGURES 1 2" captionStartId="1.[151,264,1900,1923]" captionTargetBox="[163,1418,772,1869]" captionTargetId="figure@1.[151,1436,767,1878]" captionTargetPageId="1" captionText="FIGURES 1 2. Pachyseius destitutus sp. nov., female. 1. Dorsal shield; 2. Ventral idiosoma." httpUri="https://zenodo.org/record/236433/files/figure.png" pageId="2" pageNumber="100">Fig. 2</figureCitation>
). Pre-sternal shields absent. Sternal shield 135145 long, 6575 wide at level of coxae II; central region ornamented with punctate polygonal lines; carrying three pairs of pointed needle-like setae and two pairs of lyrifissures, edges of shield strongly sclerotised. Length of sternal setae
<emphasis id="F5F9EE63DB5E5C309996F91A305485EB" box="[986,1042,1747,1771]" italics="true" pageId="2" pageNumber="100">
st
<date id="B33314B1DB5E5C3099A3F91A305485EB" box="[1007,1042,1747,1771]" pageId="2" pageNumber="100" value="1920-01-28" valueMax="1920-03-28" valueMin="1920-01-28">1-3</date>
</emphasis>
2028. Metasternal shields small, each bearing a metasternal seta (1822) and a poroid. Epigynal shield 5865 long, 7085 wide at widest point, with punctate ornamentation, and a row of four elongate platelets and a pair of small oval platelets along its posterior margin; genital pores situated outside postero-lateral margins of shield, genital setae length 1822. Ventri-anal shield with a concave anterior margin (
<figureCitation id="5FB62EF4DB5E5C30981EF8AB36E3847B" box="[594,677,1890,1915]" captionStart="FIGURES 19 22" captionStartId="8.[151,264,1740,1763]" captionTargetBox="[327,1259,501,1719]" captionTargetId="figure@8.[327,1259,501,1720]" captionTargetPageId="8" captionText="FIGURES 19 22. Pachyseius spp., females. 19. P. destitutus sp. nov., anterior region of dorsal idiosoma; 20. P. destitutus sp. nov., ventri-anal shield; 21. P. destitutus sp. nov., ventral gnathosoma; 22. P. quadrigeminus sp. nov., ventri-anal shield. Not scaled." httpUri="https://zenodo.org/record/236438/files/figure.png" pageId="2" pageNumber="100">Fig. 20</figureCitation>
) 182210 long, 182200 wide at widest point (length/width, 0.95 1.07), with polygonal ornamentation, bearing three pairs of pointed pre-anal setae (
<emphasis id="F5F9EE63DB5E5C309E79F84E301A849F" box="[1077,1116,1927,1951]" italics="true" pageId="2" pageNumber="100">Jv1</emphasis>
and
<emphasis id="F5F9EE63DB5E5C309EDEF84E3086849F" box="[1170,1216,1927,1951]" italics="true" pageId="2" pageNumber="100">Jv2,</emphasis>
2025;
<emphasis id="F5F9EE63DB5E5C309F57F84E3104849F" box="[1307,1346,1927,1951]" italics="true" pageId="2" pageNumber="100">Jv3</emphasis>
shorter, 1416), a pair of longer para-anal setae (2830), and a long postanal seta (3038). Peritrematal shields fused with exopodal shields, with four pairs of small post-stigmatal pores, their posterior ends rounded, enclosing posterior margins of coxae IV and extending medially almost to genital setae. Opisthogastric soft integument between peritrematal shields and anterolateral margins of ventri-anal shield with one pair of oval platelets. Metapodal shields elongated (6062 long) and very close to anterolateral margins of ventri-anal shield. Lateral and opisthogastric soft integument bearing 11 pairs of setae (1522).
</paragraph>
<paragraph id="C7323271DB5F5C319A8BFEE134F382F5" blockId="3.[151,1436,151,1040]" pageId="3" pageNumber="101">
<emphasis id="F5F9EE63DB5F5C319A8BFEE135108240" box="[199,342,296,320]" italics="true" pageId="3" pageNumber="101">Gnathosoma</emphasis>
(
<figureCitation id="5FB62EF4DB5F5C319B2AFEEE35ED8240" box="[358,427,295,320]" captionStart="FIGURES 3 6" captionStartId="2.[151,264,1361,1384]" captionTargetBox="[151,1429,201,1325]" captionTargetId="figure@2.[151,1436,193,1340]" captionTargetPageId="2" captionText="FIGURES 3 6. Pachyseius destitutus sp. nov., female. 3. Ventral gnathosoma; 4. Epistome; 5. Chelicera; 6. Tarsus II." httpUri="https://zenodo.org/record/236434/files/figure.png" pageId="3" pageNumber="101">Fig. 3</figureCitation>
). Corniculi short and horn-like; hypostomal seta
<emphasis id="F5F9EE63DB5F5C31999AFEE137B48240" box="[982,1010,296,320]" italics="true" pageId="3" pageNumber="101">h3</emphasis>
longest (4552),
<emphasis id="F5F9EE63DB5F5C319EF6FEE130908240" box="[1210,1238,296,320]" italics="true" pageId="3" pageNumber="101">h2</emphasis>
shortest (1014), and
<emphasis id="F5F9EE63DB5F5C319A8AFE8534A48264" box="[198,226,332,356]" italics="true" pageId="3" pageNumber="101">
h
<date id="B33314B1DB5F5C319A9AFE8534A48264" box="[214,226,332,356]" pageId="3" pageNumber="101" value="1923-01-27">1</date>
</emphasis>
2327 long;
<emphasis id="F5F9EE63DB5F5C319B39FE8435D68264" box="[373,400,333,356]" italics="true" pageId="3" pageNumber="101">pc</emphasis>
1519 long; deutosternal groove with five rows of denticles and two smooth transverse lines (
<figureCitation id="5FB62EF4DB5F5C319AD3FEA634B58288" box="[159,243,367,392]" captionStart="FIGURES 19 22" captionStartId="8.[151,264,1740,1763]" captionTargetBox="[327,1259,501,1719]" captionTargetId="figure@8.[327,1259,501,1720]" captionTargetPageId="8" captionText="FIGURES 19 22. Pachyseius spp., females. 19. P. destitutus sp. nov., anterior region of dorsal idiosoma; 20. P. destitutus sp. nov., ventri-anal shield; 21. P. destitutus sp. nov., ventral gnathosoma; 22. P. quadrigeminus sp. nov., ventri-anal shield. Not scaled." httpUri="https://zenodo.org/record/236438/files/figure.png" pageId="3" pageNumber="101">Fig. 21</figureCitation>
). Palp with 3-tined apotele. Anterior and lateral margins of epistome denticulate (
<figureCitation id="5FB62EF4DB5F5C319EC5FEA630898288" box="[1161,1231,367,392]" captionStart="FIGURES 3 6" captionStartId="2.[151,264,1361,1384]" captionTargetBox="[151,1429,201,1325]" captionTargetId="figure@2.[151,1436,193,1340]" captionTargetPageId="2" captionText="FIGURES 3 6. Pachyseius destitutus sp. nov., female. 3. Ventral gnathosoma; 4. Epistome; 5. Chelicera; 6. Tarsus II." httpUri="https://zenodo.org/record/236434/files/figure.png" pageId="3" pageNumber="101">Fig. 4</figureCitation>
). Cheliceral digits slender, movable digit about 45 long, with two subdistal teeth and an terminal hook, fixed digit about 40 long (to dorsal seta), medially edentate, terminally with bidentate hook; pilus dentilis minute; arthrodial brush short (
<figureCitation id="5FB62EF4DB5F5C319F23FE7E34E182F5" captionStart="FIGURES 3 6" captionStartId="2.[151,264,1361,1384]" captionTargetBox="[151,1429,201,1325]" captionTargetId="figure@2.[151,1436,193,1340]" captionTargetPageId="2" captionText="FIGURES 3 6. Pachyseius destitutus sp. nov., female. 3. Ventral gnathosoma; 4. Epistome; 5. Chelicera; 6. Tarsus II." httpUri="https://zenodo.org/record/236434/files/figure.png" pageId="3" pageNumber="101">Fig. 5</figureCitation>
).
</paragraph>
<paragraph id="C7323271DB5F5C319A8BFDC930328118" blockId="3.[151,1436,151,1040]" box="[199,1140,511,536]" pageId="3" pageNumber="101">
<taxonomicName id="008D49F2DB5F5C319A8BFDC9359C8118" box="[199,474,512,536]" class="Arachnida" family="Pachylaelapidae" genus="Spermathecal" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="apparatus">
<emphasis id="F5F9EE63DB5F5C319A8BFDC9359C8118" box="[199,474,512,536]" italics="true" pageId="3" pageNumber="101">Spermathecal apparatus</emphasis>
</taxonomicName>
. Spermathecal tubes not visible in the available specimens.
</paragraph>
<paragraph id="C7323271DB5F5C319A8BFDEC315081A8" blockId="3.[151,1436,151,1040]" pageId="3" pageNumber="101">
<emphasis id="F5F9EE63DB5F5C319A8BFDEC34BA813C" box="[199,252,549,572]" italics="true" pageId="3" pageNumber="101">Legs</emphasis>
. Chaetotaxy: Leg I. Coxa 0 0/2 0, trochanter 1 1/3 1, femur 2 5/4 2, genu 2 5/3 2, tibia 2 5/3 2. Leg II. Coxa 0 0/2 0, trochanter 1 0/3 1, femur 2 5/3 1, genu 2 5/2 2, tibia 2 4/2 2, tarsus 3 7/5 3. Leg III. Coxa 0 0/2 0, trochanter 1 0/3 1, femur 1 3/1 1, genu 2 4/1 1, tibia 1 3/2 1, tarsus 3 7/5 3. Leg IV. Coxa 0 0/1 0, trochanter 1 0/3 1, femur 1 3/1 1, genu 2 4/1 1, tibia 1 3/2 1, tarsus 3 7/5 3. Tarsus II with one large spur-like seta (
<figureCitation id="5FB62EF4DB5F5C319E8FFD46314E81A8" box="[1219,1288,655,680]" captionStart="FIGURES 3 6" captionStartId="2.[151,264,1361,1384]" captionTargetBox="[151,1429,201,1325]" captionTargetId="figure@2.[151,1436,193,1340]" captionTargetPageId="2" captionText="FIGURES 3 6. Pachyseius destitutus sp. nov., female. 3. Ventral gnathosoma; 4. Epistome; 5. Chelicera; 6. Tarsus II." httpUri="https://zenodo.org/record/236434/files/figure.png" pageId="3" pageNumber="101">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="8F9761FADB5F5C319A8BFD7D36B98710" pageId="3" pageNumber="101" type="etymology">
<paragraph id="C7323271DB5F5C319A8BFD7D314881CD" blockId="3.[151,1436,151,1040]" box="[199,1294,692,717]" pageId="3" pageNumber="101">
<emphasis id="F5F9EE63DB5F5C319A8BFD7D350881CD" bold="true" box="[199,334,692,717]" pageId="3" pageNumber="101">Etymology.</emphasis>
The name
<taxonomicName id="008D49F2DB5F5C319B87FD7D367081CC" box="[459,566,692,716]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="F5F9EE63DB5F5C319B87FD7D367081CC" box="[459,566,692,716]" italics="true" pageId="3" pageNumber="101">destitutus</emphasis>
</taxonomicName>
, lacking, refers to the loss of dorsal shield seta
<emphasis id="F5F9EE63DB5F5C319E06FD7D302481CC" box="[1098,1122,692,716]" italics="true" pageId="3" pageNumber="101">
z
<quantity id="00759F94DB5F5C319E1AFD7D302481CC" box="[1110,1122,692,716]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="101" unit="in" value="1.0">1</quantity>
</emphasis>
in this species.
</paragraph>
<paragraph id="C7323271DB5F5C319A8BFD1E36B98710" blockId="3.[151,1436,151,1040]" pageId="3" pageNumber="101">
<emphasis id="F5F9EE63DB5F5C319A8BFD1E355781F0" bold="true" box="[199,273,727,752]" pageId="3" pageNumber="101">Notes.</emphasis>
The new species may be easily distinguished from the other congeners by the dorsal shield with 29 pairs of setae,
<emphasis id="F5F9EE63DB5F5C319B4FFD35355D8014" box="[259,283,764,788]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
absent, and the peritrematal shields fused with all the exopodal shields. The only other species of
<taxonomicName id="008D49F2DB5F5C319ADBFCE935538038" box="[151,277,800,824]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="genus">
<emphasis id="F5F9EE63DB5F5C319ADBFCE935538038" box="[151,277,800,824]" italics="true" pageId="3" pageNumber="101">Pachyseius</emphasis>
</taxonomicName>
that lacks seta
<emphasis id="F5F9EE63DB5F5C319B8DFCE9359F8038" box="[449,473,800,824]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
is
<taxonomicName id="008D49F2DB5F5C319BB7FCE836DD8038" box="[507,667,800,824]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="F5F9EE63DB5F5C319BB7FCE8364F8038" box="[507,521,801,824]" italics="true" pageId="3" pageNumber="101">P</emphasis>
.
<emphasis id="F5F9EE63DB5F5C31985BFCE936DD8038" box="[535,667,800,824]" italics="true" pageId="3" pageNumber="101">cavernicola</emphasis>
</taxonomicName>
(we have examined the
<typeStatus id="18368CD3DB5F5C3199E1FCD630498038" box="[941,1039,799,824]" pageId="3" pageNumber="101" type="holotype">holotype</typeStatus>
of
<taxonomicName id="008D49F2DB5F5C319E78FCE830928038" box="[1076,1236,800,824]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="F5F9EE63DB5F5C319E78FCE830928038" box="[1076,1236,800,824]" italics="true" pageId="3" pageNumber="101">P. cavernicola</emphasis>
</taxonomicName>
, and confirm that
<emphasis id="F5F9EE63DB5F5C319ADBFC8D34F6805C" box="[151,176,836,860]" italics="true" pageId="3" pageNumber="101">z1</emphasis>
setae are absent), but these two species are clearly distinguished by the number of pre-anal setae (two pairs in
<taxonomicName id="008D49F2DB5F5C319FCBFC8C355D8080" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="F5F9EE63DB5F5C319FCBFC8C355D8080" italics="true" pageId="3" pageNumber="101">P. cavernicola</emphasis>
</taxonomicName>
, three in
<taxonomicName id="008D49F2DB5F5C319BD4FCA036638080" box="[408,549,872,896]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="F5F9EE63DB5F5C319BD4FCA036638080" box="[408,549,872,896]" italics="true" pageId="3" pageNumber="101">P. destitutus</emphasis>
</taxonomicName>
), the different length and form of the dorsal shield setae (spatulate in
<taxonomicName id="008D49F2DB5F5C319FCBFCA0355D80A4" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="F5F9EE63DB5F5C319FCBFCA0355D80A4" italics="true" pageId="3" pageNumber="101">P. cavernicola</emphasis>
</taxonomicName>
, pointed in
<taxonomicName id="008D49F2DB5F5C319BE9FC44366880A4" box="[421,558,908,932]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="F5F9EE63DB5F5C319BE9FC44366880A4" box="[421,558,908,932]" italics="true" pageId="3" pageNumber="101">P. destitutus</emphasis>
</taxonomicName>
), the shape of the epistome (broadly arched in
<taxonomicName id="008D49F2DB5F5C319E07FC4430AD80A4" box="[1099,1259,908,932]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="F5F9EE63DB5F5C319E07FC4430AD80A4" box="[1099,1259,908,932]" italics="true" pageId="3" pageNumber="101">P. cavernicola</emphasis>
</taxonomicName>
, narrower in
<taxonomicName id="008D49F2DB5F5C319FCBFC44354480C8" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="F5F9EE63DB5F5C319FCBFC44354480C8" italics="true" pageId="3" pageNumber="101">P. destitutus</emphasis>
</taxonomicName>
), and the number of spur-like seta on tarsus II (two in
<taxonomicName id="008D49F2DB5F5C319928FC78304580C8" box="[868,1027,944,968]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="cavernicola">
<emphasis id="F5F9EE63DB5F5C319928FC78304580C8" box="[868,1027,944,968]" italics="true" pageId="3" pageNumber="101">P. cavernicola</emphasis>
</taxonomicName>
, one in
<taxonomicName id="008D49F2DB5F5C319E11FC7830A280C8" box="[1117,1252,944,968]" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="F5F9EE63DB5F5C319E11FC7830A280C8" box="[1117,1252,944,968]" italics="true" pageId="3" pageNumber="101">P. destitutus</emphasis>
</taxonomicName>
). The collection of
<specimenCount id="D18BF9F8DB5F5C319AFBFC1D357580ED" box="[183,307,980,1005]" pageId="3" pageNumber="101" type="female">17 females</specimenCount>
and no males, and the apparent absence of insemination structures in the females, suggests that
<taxonomicName id="008D49F2DB5F5C319FCBFC1C35458710" class="Arachnida" family="Pachylaelapidae" genus="Pachyseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="101" phylum="Arthropoda" rank="species" species="destitutus">
<emphasis id="F5F9EE63DB5F5C319FCBFC1C35458710" italics="true" pageId="3" pageNumber="101">P. destitutus</emphasis>
</taxonomicName>
may be thelytokous, but that is unconfirmed.
</paragraph>
</subSubSection>
</treatment>
</document>