plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/4F/1F/A8/4F1FA810C39BC570692482188FFC2F59.xml
maintenance 50c7799abe added 4F
2024-06-21 12:36:10 +02:00

316 lines
38 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2FC8257A46E4BF8148BDC131F830566E" ID-DOI="http://dx.doi.org/10.3897/zookeys.364.6109" ID-GBIF-Dataset="13f3f91c-6d08-48c0-986b-d2e530c0c06f" ID-PMC="PMC3890664" ID-Pensoft-Pub="1313-2970-364-47" ID-PubMed="24453545" ID-ZooBank="1D0850672D944404B0BD6E6615371BF6" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-364-47" ModsDocOrigin="ZooKeys 364" ModsDocTitle="Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae) with descriptions of three new species from Peru" checkinTime="1451246544016" checkinUser="pensoft" docAuthor="Aguilar, Cesar, Wood Jr, Perry L., Cusi, Juan C., Guzman, Alfredo, Huari, Frank, Lundberg, Mikael, Mortensen, Emma, Ramirez, Cesar, Robles, Daniel, Suarez, Juana, Ticona, Andres, Vargas, Victor J., Venegas, Pablo J. &amp; Sites Jr, Jack W." docDate="2013" docId="4F1FA810C39BC570692482188FFC2F59" docLanguage="en" docName="ZooKeys 364: 47-91" docOrigin="ZooKeys 364" docSource="http://dx.doi.org/10.3897/zookeys.364.6109" docTitle="Liolaemus chavin Aguilar, Jr, Cusi, Guzman, Huari, Lundberg, Mortensen, Ramirez, Robles, Suarez, Ticona, Vargas, Venegas &amp; Jr, 2013, sp. n." docType="treatment" docUuid="47B7926F-7D66-4C0B-9F25-9696C916E6C2" docUuidSource="ZooBank" docVersion="5" lastPageNumber="67" masterDocId="FFE2FFCFC41ABC38FFDEFFE3FF9EFFBB" masterDocTitle="Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae) with descriptions of three new species from Peru" masterLastPageNumber="91" masterPageNumber="47" pageNumber="64" updateTime="1701315944844" updateUser="plazi">
<mods:mods id="49CC7A8400CB48AD7E7163EB5780B370" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="52E85E0CD9B66C92C933EB5388249364">
<mods:title id="68EA13E179BA06496B271646B265D46B">Integrative taxonomy and preliminary assessment of species limits in the Liolaemus walkeri complex (Squamata, Liolaemidae) with descriptions of three new species from Peru</mods:title>
</mods:titleInfo>
<mods:name id="C4463D9DFEF5AA9DC7AA1D8FDEA03746" type="personal">
<mods:role id="923B2F9273D68BE3CFC515C7AB53E8DE">
<mods:roleTerm id="30FCCE5CE02003306D3DC98A7F6167A0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F16F363B638EF495E8E6510502D60C0">Aguilar, Cesar</mods:namePart>
</mods:name>
<mods:name id="238C15E34C6C0AE22E67BF2DA57AE68F" type="personal">
<mods:role id="B0465D2F884C3506F178A7411F258E5D">
<mods:roleTerm id="44DC504DE04567CA3D9F62B0632599B1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A3E20E2D5E07C29A5DCE6FD604694BB5">Wood Jr, Perry L.</mods:namePart>
</mods:name>
<mods:name id="012758FD5A7C9EC31E36B798A583F12E" type="personal">
<mods:role id="EBA5C67642441E10AC8E92045431EE69">
<mods:roleTerm id="99FD1619D0100490F08065A8DD3A5BD2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="ECCE05B7D041F7E870963482A22D5744">Cusi, Juan C.</mods:namePart>
</mods:name>
<mods:name id="EE3FFBBEAFA66FC650381CC54CF63535" type="personal">
<mods:role id="527F8620CD49BC4C3389AEC7975A2A4C">
<mods:roleTerm id="8125FBE0C59C8894DE32FCD23ED0DB05">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FBA3867EB37728A2327E74BE273F7C75">Guzman, Alfredo</mods:namePart>
</mods:name>
<mods:name id="894477E6B1C1D35CCE43E67AA19ED656" type="personal">
<mods:role id="6D5A1890BEA5A26EE37895F0C58583DB">
<mods:roleTerm id="109C08F47FB8D31F1281D9F3CF1B460C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F2E2E88C4C784C8AFC95951B7CFC1797">Huari, Frank</mods:namePart>
</mods:name>
<mods:name id="6EA533EE52F5E0BFFCB1C8B67C00F75D" type="personal">
<mods:role id="D40D32D9A09B17B1F60E9D4508FE0C2A">
<mods:roleTerm id="A5635E26576894106F0A7F2112F7D95E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4B34DCFF5E0F4B593FF3593C523AB975">Lundberg, Mikael</mods:namePart>
</mods:name>
<mods:name id="6A7F97CE67C3DBC7F5328B1EFB78B8AE" type="personal">
<mods:role id="23B2E6C946AF43EEA4B0F292EFDC374C">
<mods:roleTerm id="E561110C1A6921315E8744AB8D897029">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="29E1A89A11C963519150FD028E271D95">Mortensen, Emma</mods:namePart>
</mods:name>
<mods:name id="6B4F4ECD260214C8B2A15D65B35EFBBC" type="personal">
<mods:role id="D59A93F86AE770B83C8A53555B60D508">
<mods:roleTerm id="C295744C4EAF009599269C0279E31A44">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D83E6B4969B3E4DAE5902AB2DC2FC571">Ramirez, Cesar</mods:namePart>
</mods:name>
<mods:name id="B5A736454CE5BD6B8DD3A8B911256B85" type="personal">
<mods:role id="773B25CA1409DF75A39FE167C858EE98">
<mods:roleTerm id="FF4E4CE65EEBA7F179D80FA7FDA33731">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DC20F4ADBB1FC31E7F584E23D0DCB8AF">Robles, Daniel</mods:namePart>
</mods:name>
<mods:name id="D8DA796F75FC32878B6ADE267B804A09" type="personal">
<mods:role id="C3261E7E44E2B476CEDA94A67081095E">
<mods:roleTerm id="B3557481300CF0F564A480125498960E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="314EA5A6B26579A00D9FC4E427B7EA5C">Suarez, Juana</mods:namePart>
</mods:name>
<mods:name id="E4B65A2F0F333AFBFAA7C209AA8182C7" type="personal">
<mods:role id="1AF6EF3774857717A923C99EC4EA4EA1">
<mods:roleTerm id="522468B19FBDC2A7BDFB1D9D75631A99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="23F6DEDD5FA4C6D67B436B6B9A2DC329">Ticona, Andres</mods:namePart>
</mods:name>
<mods:name id="602F26DB7DD9BB89E53CB475B8BF8236" type="personal">
<mods:role id="E2441D084F8FA6D1EB706D37C1551848">
<mods:roleTerm id="B21847406BEF6FBAF3059F6CAF166720">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4C187C4ECC86BBC6EF96073ACF6D509E">Vargas, Victor J.</mods:namePart>
</mods:name>
<mods:name id="8115DD09EA9D5FB1D01F769B6D74523D" type="personal">
<mods:role id="3A98B2F94267E55C65D4455A98986290">
<mods:roleTerm id="94A0B77F035249F012F7375BBFDC0B83">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2A00DDD19AFFA09D7FF20E0CE54E995D">Venegas, Pablo J.</mods:namePart>
</mods:name>
<mods:name id="24C4F9533694B7874D59ED3A3FA17BB2" type="personal">
<mods:role id="A05B81154C7F35C13998D69F3A02A2C9">
<mods:roleTerm id="2F714088BEE0EA697966C2E5E496B92A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E3B2FCA72D132E4A124E49FF122D9654">Sites Jr, Jack W.</mods:namePart>
</mods:name>
<mods:typeOfResource id="09D6F42699394A92DEDC6004922F2CCE">text</mods:typeOfResource>
<mods:relatedItem id="E8C9A8739C9DD07A844366F22FBA830F" type="host">
<mods:titleInfo id="36BA8C8E0456AF050995B90D617DFBFF">
<mods:title id="A35E0E01D5969D19F0CDDABCDFD450FC">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="CAD9EB8D67701FA1AA52B609A91324AB">
<mods:date id="20E6F44176120C8CFBB69D4356210307">2013</mods:date>
<mods:detail id="E84F1921D1BCECBD7AC8554D03C9F86B" type="volume">
<mods:number id="16D5328A91EB084B8E4E13A17F56A691">364</mods:number>
</mods:detail>
<mods:extent id="19FAF9F3838539AA7DF02F24D76DE3BC" unit="page">
<mods:start id="CD697FCCDF07DD9CFB373E2DEC387D92">47</mods:start>
<mods:end id="97E6D48031E756F9359B7F791C4EAE4B">91</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="8FF6B2F3CCCCA6B1FC03B6F1B5F7304B">
<mods:url id="BDBDB9F8F63489A1A6491A6266AEA14A">http://dx.doi.org/10.3897/zookeys.364.6109</mods:url>
</mods:location>
<mods:classification id="0BB6298997B5670E8187859F7CFC138E">journal article</mods:classification>
<mods:identifier id="3C3D27DA19F81BEE424BF0B0731E40EE" type="DOI">http://dx.doi.org/10.3897/zookeys.364.6109</mods:identifier>
<mods:identifier id="4622497CAA08584890000DF3E30AD09B" type="Pensoft-Pub">1313-2970-364-47</mods:identifier>
<mods:identifier id="CEAD773230A5D551151E096AB6B585EA" type="ZooBank">1D0850672D944404B0BD6E6615371BF6</mods:identifier>
</mods:mods>
<treatment id="4F1FA810C39BC570692482188FFC2F59" ID-GBIF-Taxon="152050211" LSID="urn:lsid:zoobank.org:act:47B7926F-7D66-4C0B-9F25-9696C916E6C2" httpUri="http://treatment.plazi.org/id/4F1FA810C39BC570692482188FFC2F59" lastPageId="20" lastPageNumber="67" pageId="17" pageNumber="64">
<subSubSection id="9985C6508A61A48A2B0C47348CAAECEE" pageId="17" pageNumber="64" type="nomenclature">
<paragraph id="58BFC49C8E8419C6AE322E467DAFE28F" pageId="17" pageNumber="64">
<taxonomicName id="A96B430D5B61AEE84046CCBC534270F1" ID-CoL="72HJH" LSID="http://zoobank.org/47B7926F-7D66-4C0B-9F25-9696C916E6C2" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chavin" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="chavin">Liolaemus chavin</taxonomicName>
<taxonomicNameLabel id="B0ED85C61E7F2A08D38E01947E7764CB" pageId="17" pageNumber="64">sp. n.</taxonomicNameLabel>
Figure 8
</paragraph>
</subSubSection>
<subSubSection id="FE82B8759CD9FD785F8126FA2B118150" pageId="17" pageNumber="64" type="reference_group">
<paragraph id="2AC06717A230EEF438066C8D7F132FB4" pageId="17" pageNumber="64">
<taxonomicName id="055F0EC32F2F020848397EE65960C7C3" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus alticolor" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="alticolor">Liolaemus alticolor</taxonomicName>
Lehr 2002
</paragraph>
<paragraph id="C4E80A737162BFAA5BF23383EFE3AF74" pageId="17" pageNumber="64">
<taxonomicName id="1B9639681C58E863AB64258110FFDDFB" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus incaicus" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="incaicus">Liolaemus incaicus</taxonomicName>
Lobo, Quinteros and
<normalizedToken id="667FAF13575DF4E906C1AADD63EA3049" originalValue="Díaz">Diaz</normalizedToken>
<normalizedToken id="5C4DEADC8E808F8315714A16DF6A3D55" originalValue="Gómez">Gomez</normalizedToken>
2007
</paragraph>
<paragraph id="96F5516484B13B86D3F6249DF17D8269" pageId="17" pageNumber="64">
<taxonomicName id="5F3379C2E225C07DF34BB3928BEFDAE8" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus" pageId="17" pageNumber="64" phylum="Chordata" rank="genus">Liolaemus</taxonomicName>
aff. walkeri Langstroth 2011
</paragraph>
</subSubSection>
<subSubSection id="AB260C555D7C3F2FD98012C6D67D8983" pageId="17" pageNumber="64" type="holotype">
<paragraph id="7DFD9925CF7ED47287A294505430F389" pageId="17" pageNumber="64">Holotype.</paragraph>
<paragraph id="83B589F855D1CF956F51364BC6AD7880" pageId="17" pageNumber="64">
MUSM 25417, adult male collected at Conococha, Recuay Province, Ancash Department, Peru,
<geoCoordinate id="7CEA26FA6EC48D7123F0F1EC4067B32E" direction="south" orientation="latitude" precision="55" value="-10.123">10.123S</geoCoordinate>
,
<geoCoordinate id="962605D2A0541DD333140C15895ACACB" direction="west" orientation="longitude" precision="55" value="-77.293">77.293W</geoCoordinate>
, elevation 4100 m, on 31 March 2006 by Mikael Lundberg.
</paragraph>
</subSubSection>
<subSubSection id="0D21DB972D5509E8930BE2723F330B24" pageId="17" pageNumber="64" type="paratypes">
<paragraph id="0668D23BA574F10A96EA0E972B7E3797" pageId="17" pageNumber="64">Paratypes.</paragraph>
<paragraph id="7D784E74532E9892FBB81016EFB95F77" pageId="17" pageNumber="64">
Three males (MUSM 20141, 20143, 20146) and twelve females (MUSM 25324, 25327, 25328, 25331, 25333, 25334, 25340, 25423, 25412, 30812, 30813, BYU 50192) from the same locality as the holotype. One male (MUSM 20147) from Carpa, Recuay Province, Ancash Department, on 28 February 2001 by Edgar Lehr and
<normalizedToken id="08FD99070C69A56A49551A4AB248B2BC" originalValue="César">Cesar</normalizedToken>
Aguilar (see Data resources for elevation and coordinates). One female (MUSM 20201) from La
<normalizedToken id="FC508DC2E4F2C4F55A23FE9A12F93CD3" originalValue="Unión">Union</normalizedToken>
,
<normalizedToken id="74EEF9B22CBBC4BD51AB2B5F1A80280A" originalValue="Huánuco">Huanuco</normalizedToken>
Department, on 3 March 1997 by Edgar Lehr (see Data resources for elevation and coordinates). Seven males (CORBIDI 10439, 10450, 10452, 10442, 10441, 10443, 10437) and six females (CORBIDI 10444, 10451, 10440, 10438, 10445, 10449) from Pampas de Huamani, San Marcos District, Huari Province, Ancash Department, on 12 February 2012 by Pablo J. Venegas (see Data resources for elevation and coordinates).
</paragraph>
</subSubSection>
<subSubSection id="453658044418130108D397B9BE6A04DE" pageId="17" pageNumber="64" type="diagnosis">
<paragraph id="A09E9A1E6CF02D33E5563D3D7498F4F9" pageId="17" pageNumber="64">Diagnosis.</paragraph>
<paragraph id="421D91BC242E376FEC2EB1F4BAB7D005" pageId="17" pageNumber="64">
Small (61.7 mm maximum SVL), slender
<taxonomicName id="119C43D09A837F98695E21BA6DFFD0E3" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus" pageId="17" pageNumber="64" phylum="Chordata" rank="genus">Liolaemus</taxonomicName>
closely related to
<taxonomicName id="CE411842B650E9667579FF455740E142" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus walkeri" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="walkeri">Liolaemus walkeri</taxonomicName>
,
<taxonomicName id="E0E6B6C7F70E51B9E452B8BF7421475A" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus tacnae" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="tacnae">Liolaemus tacnae</taxonomicName>
,
<taxonomicName id="F43A028745AEAA007BDD11BA57D97A39" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus pachacutec" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="pachacutec">Liolaemus pachacutec</taxonomicName>
sp. n. and
<taxonomicName id="27A7919ED6E4F8B1E8441F7A685CE9AD" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus wari" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="wari">Liolaemus wari</taxonomicName>
sp. n. (described below) (Fig. 1). It differs from
<taxonomicName id="F1D93D688475ABD72236248AFCF7C6B6" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus walkeri" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="walkeri">Liolaemus walkeri</taxonomicName>
,
<taxonomicName id="080AA8A94269EAB53C60C8B0A3651992" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus pachacutec" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="pachacutec">Liolaemus pachacutec</taxonomicName>
sp. n. and
<taxonomicName id="DEF5D885953DD37CCD246A2D1059B261" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus wari" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="wari">Liolaemus wari</taxonomicName>
sp. n. in the absence of precloacal pores in males. It differs from
<taxonomicName id="9A853D4B2E01D213697A85558072C156" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus tacnae" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="tacnae">Liolaemus tacnae</taxonomicName>
in having a melanistic belly in adult males (not melanistic in adult
<taxonomicName id="8135C81B116F7CF7E60F6F7A8EB42380" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus tacnae" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="tacnae">Liolaemus tacnae</taxonomicName>
males).In comparison with other species assigned to the
<taxonomicName id="4823EA55EEF6A52B5391E22AC562B123" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus alticolor" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="alticolor">Liolaemus alticolor</taxonomicName>
group,
<taxonomicName id="CCDA9BE48CB3863DEB3213C06E5C1AAB" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chavin" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="chavin">Liolaemus chavin</taxonomicName>
sp. n. differs from
<taxonomicName id="11CCC55B6D65436460605A4F67E82130" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus bitaeniatus" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="bitaeniatus">Liolaemus bitaeniatus</taxonomicName>
and
<taxonomicName id="9286AAE815541EA94CA995DC02EF378B" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus pagaburoi" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="pagaburoi">Liolaemus pagaburoi</taxonomicName>
in having a smooth dorsal surface of the head (rough to slightly rough dorsal surface). It differs from
<taxonomicName id="449135149327172DAAE3DEE82FB5D904" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus alticolor" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="alticolor">Liolaemus alticolor</taxonomicName>
,
<taxonomicName id="DA0C345F31B1DB7B967536587DFD2290" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus aparicioi" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="aparicioi">Liolaemus aparicioi</taxonomicName>
,
<taxonomicName id="1AF7FCAD77B83CF3C3E433ACB5A47BA6" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus incaicus" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="incaicus">Liolaemus incaicus</taxonomicName>
,
<taxonomicName id="6A34F586DAA2914B678AEE840CB35BAA" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus paulinae" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="paulinae">Liolaemus paulinae</taxonomicName>
,
<taxonomicName id="CB3CFC29C927AF1C7F85A0873FE178E1" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus pyriphlogos" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="pyriphlogos">Liolaemus pyriphlogos</taxonomicName>
,
<taxonomicName id="FD87B479F2E5728BB5D8BC8E1288EE09" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus puna" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="puna">Liolaemus puna</taxonomicName>
, and
<taxonomicName id="82A825B8F1FAF7014FC67830FB142377" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus variegatus" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="variegatus">Liolaemus variegatus</taxonomicName>
in the absence of precloacal pores in males.
<taxonomicName id="A41C4013BB85F2FADF55049B00E088D1" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chaltin" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="chaltin">Liolaemus chaltin</taxonomicName>
also lacks precloacal pores in males, but
<taxonomicName id="CDD128107C1EE2E6AC14F3C4108B6C5C" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chavin" pageId="17" pageNumber="64" phylum="Chordata" rank="species" species="chavin">Liolaemus chavin</taxonomicName>
sp. n. differs in having also a melanistic belly in adult males.
</paragraph>
</subSubSection>
<subSubSection id="D25D64BFC546F021080886E84BBD7CD8" lastPageId="19" lastPageNumber="66" pageId="17" pageNumber="64" type="description of holotype">
<paragraph id="EDE94DCB30CAC0F805D2E3621965C2F5" pageId="17" pageNumber="64">Description of holotype.</paragraph>
<paragraph id="9CD4565ADF119ECD450E6AD9FB661F5D" pageId="17" pageNumber="64">Adult male; SVL 56.8 mm; head length 13.7 mm; head width 11.3 mm; head height 7.7 mm; axilla-groin 21.0 mm (37% of SVL); foot length 10.3 mm (18.3% of SVL); tail length (regenerated) 35.2 mm (0.6 times SVL).</paragraph>
<paragraph id="9B0611BE756F9F44C9DFB3BF6EBC9A4C" lastPageId="19" lastPageNumber="66" pageId="17" pageNumber="64">
Fifteen dorsal head scales (from a line drawn horizontally between anterior edges of external auditory meatus to anterior border of rostral). Dorsal head scales smooth except for the interparietal and surrounding scales, scale organs more abundant in prefrontal, internasal, and supralabial regions. Five scale organs on postrostral. Nasal scale in contact
<pageBreakToken id="A0F60F068159BD8B14AEDF1F5B1011C3" pageId="18" pageNumber="65" start="start">with</pageBreakToken>
rostral, separated from first supralabial by one scale, nasal bordered by eight scales; canthus separated from nasal by one scale. Six supralabials. Six lorilabial scales, three in contact with the subocular. Six infralabials. Auditory meatus oval (height 2.3 mm, width 1.2 mm), with three small, projecting scales on anterior margin. Seven convex, smooth temporals.
<normalizedToken id="6A95A487D054A0B3EA413086B96D0069" originalValue="Orbit–auditory">Orbit-auditory</normalizedToken>
meatus distance 4.9 mm.
<normalizedToken id="CE043664C85AFF46A56299443459479E" originalValue="Orbit–anterior">Orbit-anterior</normalizedToken>
margin of rostral distance 6.3 mm. Rostral almost three times wider than high (width 2.9 mm; height 1.2 mm). Mental subpentagonal, about two times as wide as high (width 3.2 mm; height 1.7 mm). Interparietal pentagonal with an elongated posterior apex, bordered by eight scales, the parietal slightly smaller. Frontal quadrangular. Supraorbital semicircles complete on both sides. Semicircles formed by 6 scales. Four enlarged supraoculars. Six distinctly imbricate superciliaries on both sides. Eleven upper and ten lower ciliaries.
<pageBreakToken id="B36CE853659F10B2EF2548C9D6CB134B" pageId="19" pageNumber="66" start="start">Subocular</pageBreakToken>
elongate, 3.8 mm, longer than eye diameter (2.9 mm), separated from supralabials by a single, but interrupted row of lorilabials. Second supralabial elongate, 1.9 mm. Six lorilabials with single and double rows of scale organs. Sixth, fifth and fourth lorilabials contacting subocular. Preocular small, separated from lorilabial row by one scale. Postocular as large as preocular. Mental in contact with four scales: first infralabials (on each side) and two enlarged chin shields. Chin shields forming a longitudinal row of three enlarged scales separated one from the other by seven smaller scales. Scales of throat round, flat, and imbricate. Twenty-four gulars between auditory meatus. Longitudinal neck fold without keeled scales, that are similar to dorsal in size scales. Antehumeral pocket and antehumeral neck fold well developed. Forty-two scales between auditory meatus and shoulder (counting along postauricular and longitudinal neck fold), thirty-two scales between auditory meatus and antehumeral neck fold. Gular folds absent.
</paragraph>
<paragraph id="1AEB230B36898C1CA357CEC8448BEE4D" pageId="19" pageNumber="66">Dorsal scales rhomboidal, keeled, and imbricate. Sixty-six dorsal scales between occiput and level of groin. Sixty-two scales around midbody. Thirty rows of keeled scales on dorsum at midtrunk. Scales become smooth along flank and toward belly. Ventral scales slightly wider than dorsals. Eighty-two ventral scales between mental and cloaca; no precloacal pores. Supracarpals laminar, round, and smooth. Subdigital lamellae of fingers with three keels, in number I: 6; II: 11; III: 14; IV: 15; V: 10 (right hand). Claws moderately long. Supradigital lamellae convex, smooth, and imbricate. Infracarpals and infratarsals keeled, distinctly imbricate. Supratarsals smooth. Subdigital lamellae of toes I: 13; II: 13; III: 13; IV: 12; V: 6 (right foot).</paragraph>
</subSubSection>
<subSubSection id="57D8BDE229F7F6FE92202D380F1F4BFB" pageId="19" pageNumber="66" type="color pattern in preservation">
<paragraph id="A85589AD806C07F75DB52B5F88E8D9D5" pageId="19" pageNumber="66">Color pattern in preservation.</paragraph>
<paragraph id="F4C9AA110C0CD1B51D6B972EB3BBD3B8" pageId="19" pageNumber="66">Dorsal background color from occiput to base of tail greenish brown. Black continuous vertebral stripe present. Dark paravertebral marks. Paravertebral and vertebral fields of same background color. Dorsolateral stripes distinctly cream-color. Small dark cream-colored markings scattered in lateral field. Cream ventrolateral stripe, beginning on the upper auricular meatus, continuing across the longitudinal neck fold, through the shoulders, ending in the groin. Dark and small cream-colored marks in the ventral field. Black ventral color from about second third of head to femur, tibia and first third of tail. Dark and cream-colored small markings in first third of ventral head and two posterior thirds of tail.</paragraph>
</subSubSection>
<subSubSection id="3823DB04BA98EDBB4FB24C03F6277A88" pageId="19" pageNumber="66" type="color pattern in life">
<paragraph id="B8CC5824C751122A7F45C1CF0344C46C" pageId="19" pageNumber="66">Color pattern in life.</paragraph>
<paragraph id="A75182DC79D9FFAFD087E18379DFD39E" pageId="19" pageNumber="66">Head dorsally brown with black and light brown dots. Subocular cream colored, dorsum bisected by a dark vertebral line. Vertebral field not conspicuous, bordering the vertebral line with a tenuous yellowish line. Paravertebral field with dark marks, bordered dorsally by a yellowish cream dorsolateral stripe. Lateral field with black and yellow reticulated pattern and white dots. Inconspicuous ventrolateral stripe, beginning on upper margin of auricular meatus, continuing from the longitudinal neck fold, through the shoulders, ending in the groin. Ventrolateral similar to lateral field but with more white dots. Fore and hind limbs same color as the paravertebral field, with diffuse dorsal markings. Dark, melanistic ventral color from about second third of head to femur, tibia and first third of tail. Dark and white dots in first third of ventral head and two posterior thirds of tail.</paragraph>
</subSubSection>
<subSubSection id="F7C1A4619702F475EC1F1BDB002D00CE" lastPageId="20" lastPageNumber="67" pageId="19" pageNumber="66" type="variation">
<paragraph id="8AB96888E8971139FBC0FF77E0E68618" pageId="19" pageNumber="66">Variation.</paragraph>
<paragraph id="FB2CEE6481105D69AE27CB486EDD1794" lastPageId="20" lastPageNumber="67" pageId="19" pageNumber="66">
Variation in characters is summarized in Tables 1-4. There is sexual dichromatism. Adult males exhibit melanistic belly, cloacal region and throat, or mela
<pageBreakToken id="A5D98BC4A045575A774CBA58B401F674" pageId="20" pageNumber="67" start="start">nistic</pageBreakToken>
belly only; adult females exhibit black and white spots on belly, cloacal region and throat, or yellowish belly and tail.
</paragraph>
</subSubSection>
<subSubSection id="BF79A07B5E0A3D9281CC05DC30DD4BB6" pageId="20" pageNumber="67" type="etymology">
<paragraph id="8E9F16867FC87E260681DAA0B187D76E" pageId="20" pageNumber="67">Etymology.</paragraph>
<paragraph id="1E4B6BFD1E05F92CFF50BEC77D0041A0" pageId="20" pageNumber="67">The specific epithet chavin refers to the pre-Inca culture Chavin, which had its center close to the type locality and frequently depicted reptile figures on some of its most remarkable sculptures. The species name is in the nominative singular.</paragraph>
</subSubSection>
<subSubSection id="FE3363DFFEE1033F6A2C24DD641099B3" pageId="20" pageNumber="67" type="distribution and natural history">
<paragraph id="E5C1BAABCA89551C914B293D748CD170" pageId="20" pageNumber="67">Distribution and natural history.</paragraph>
<paragraph id="4EC07BD47E99554D0D95B6CE561B40A0" pageId="20" pageNumber="67">
<taxonomicName id="5CC4D503C57968EE783A901095667E30" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chavin" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="chavin">Liolaemus chavin</taxonomicName>
sp. n. is known from four localities in the central Andes, at elevations of 3535-4450 m in Ancash and
<normalizedToken id="DB460718FAF28A921548951C61F0A767" originalValue="Huánuco">Huanuco</normalizedToken>
Departments in western central Peru (Fig. 11). It is the northernmost species of the subgenus
<taxonomicName id="A63E843316A9387D008EF98062BE1BF9" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus" pageId="20" pageNumber="67" phylum="Chordata" rank="genus">Liolaemus</taxonomicName>
.
</paragraph>
<paragraph id="B44E3D9CC4FF15CC5B06024C893B120F" pageId="20" pageNumber="67">
<taxonomicName id="4E3DA6735FE4EFF6BE7AA862A348E6A8" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chavin" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="chavin">Liolaemus chavin</taxonomicName>
sp. n. was found active and under rocks in grassland and shrubland habitats at higher and lower elevations respectively (Fig. 8). In Pampas de Huamani the new species was usually found basking on grass up to 60 cm above the ground, and when they were disturbed they escaped into the base of grass clumps. Individuals basking on rocks were very rare in all localities. On cloudy days we found this species inactive hidden in the base of grass clumps, although some individuals were also found inactive under rocks. This species is viviparous; one female showed two uterine chambers per side with developed embryos, yolk and no visible shell in either chamber, and three females showed two uterine chambers per side with yolk, without developed embryos and no visible shell in each chamber. At the type locality no sympatric species of reptiles were found, but four amphibians are known:
<taxonomicName id="E5B6C1820C748279E48E54CBD1AADF9A" class="Amphibia" family="Leptodactylidae" genus="Pleurodema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pleurodema marmoratum" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="marmoratum">Pleurodema marmoratum</taxonomicName>
(
<normalizedToken id="31821B3185308233C03121AAD50A046A" originalValue="Duméril">Dumeril</normalizedToken>
&amp; Bibron, 1840),
<taxonomicName id="B8AEEA8BBBA3AC5642586D6379F3CA6E" class="Amphibia" family="Telmatobiidae" genus="Telmatobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telmatobius mayoloi" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="mayoloi">Telmatobius mayoloi</taxonomicName>
Salas &amp; Sinsch, 1996,
<taxonomicName id="7DFCBD8D624A1C4CEE34F8D7D17027FB" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca peruana" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="peruana">Gastrotheca peruana</taxonomicName>
and
<taxonomicName id="B1AE86CD982DBF28E7CE920B3987C8A6" class="Amphibia" family="Bufonidae" genus="Rhinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinella (Bufo) spinulosa" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="spinulosa" subGenus="Bufo">Rhinella (Bufo) spinulosa</taxonomicName>
(Wiegmann, 1834) (
<bibRefCitation id="D60C18C042F79D26B76327BFB28D262B" author="Lehr, E" journalOrPublisher="Natur und Tier-Verlag, Muenster" pageId="35" pageNumber="82" title="Amphibien und Reptilien in Peru." year="2002">Lehr 2002</bibRefCitation>
; personal observations). Sympatric species at Catac include the anurans
<taxonomicName id="3E4F24BF3222136599304597EA0907FA" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca peruana" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="peruana">Gastrotheca peruana</taxonomicName>
,
<taxonomicName id="380F739BD0DF9671AA1CE370A9743F32" class="Amphibia" family="Bufonidae" genus="Rhinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinella (Bufo) spinulosa" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="spinulosa" subGenus="Bufo">Rhinella (Bufo) spinulosa</taxonomicName>
,
<taxonomicName id="A8639AF552E56AE5CA8EF80EAA638720" class="Amphibia" family="Telmatobiidae" genus="Telmatobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telmatobius rimac" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="rimac">Telmatobius rimac</taxonomicName>
Schmidt, 1954,
<taxonomicName id="71B40C7C013E01182C804C151530D761" class="Amphibia" family="Telmatobiidae" genus="Telmatobius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Telmatobius mayoloi" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="mayoloi">Telmatobius mayoloi</taxonomicName>
, and the lizard
<taxonomicName id="37459BED8C04E0E3C1BB4A602F0EB48F" class="Squamata" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus chrysopygus" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="chrysopygus">Stenocercus chrysopygus</taxonomicName>
Boulenger, 1900; at Carpa,
<taxonomicName id="5374D8F78612527FFF56CAA8FC934C88" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca peruana" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="peruana">Gastrotheca peruana</taxonomicName>
(Boulenger, 1900),
<taxonomicName id="15A4D9E76FA6316FAB58137DDDBB6B40" class="Amphibia" family="Bufonidae" genus="Rhinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinella (Bufo) spinulosa" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="spinulosa" subGenus="Bufo">Rhinella (Bufo) spinulosa</taxonomicName>
and
<taxonomicName id="9CE5542FF5029681A99B9096685A9534" class="Amphibia" family="Leptodactylidae" genus="Pleurodema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pleurodema marmoratum" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="marmoratum">Pleurodema marmoratum</taxonomicName>
; at Pampas de Huamani,
<taxonomicName id="E44B1FBF45F245EAFCDEFA720601D462" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca peruana" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="peruana">Gastrotheca peruana</taxonomicName>
,
<taxonomicName id="D5FC8231AB4F0E677108EC521A5D04E5" class="Amphibia" family="Leptodactylidae" genus="Pleurodema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pleurodema marmoratum" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="marmoratum">Pleurodema marmoratum</taxonomicName>
and
<taxonomicName id="79EEDEC9BF778682A3A7257FB702AE17" class="Amphibia" family="Bufonidae" genus="Rhinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinella (Bufo) spinulosa" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="spinulosa" subGenus="Bufo">Rhinella (Bufo) spinulosa</taxonomicName>
; and at La
<normalizedToken id="82486314B466898DCCDDB3716B91B420" originalValue="Unión">Union</normalizedToken>
,
<taxonomicName id="955DA84CEEB6D4E15F9BF2EF21B4B54E" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca griswoldi" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="griswoldi">Gastrotheca griswoldi</taxonomicName>
Shreve, 1941,
<taxonomicName id="4FF3B1A7249E70AB80FA7E34B218772D" class="Amphibia" family="Hemiphractidae" genus="Gastrotheca" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gastrotheca peruana" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="peruana">Gastrotheca peruana</taxonomicName>
,
<taxonomicName id="B3A13A6ECB600704D626FEC1CBE51487" class="Amphibia" family="Bufonidae" genus="Rhinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhinella (Bufo) spinulosa" order="Anura" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="spinulosa" subGenus="Bufo">Rhinella (Bufo) spinulosa</taxonomicName>
and
<taxonomicName id="1C385428820AF299B3150A89FA7EFFD8" class="Squamata" family="Tropiduridae" genus="Stenocercus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenocercus chrysopygus" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="chrysopygus">Stenocercus chrysopygus</taxonomicName>
(Lehr, 2002).
</paragraph>
<caption id="EBA37D7C2EC8F2422BD482ED730A184F" pageId="20" pageNumber="67">
<paragraph id="17E5902052090BBA2B34DAFB81F9DF6F" pageId="20" pageNumber="67">
Figure 8. Dorsal (A) and ventral (B) views of the holotype of
<taxonomicName id="67C55EADBFA8C763DAB5B40BA9009641" class="Squamata" family="Liolaemidae" genus="Liolaemus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liolaemus chavin" pageId="20" pageNumber="67" phylum="Chordata" rank="species" species="chavin">Liolaemus chavin</taxonomicName>
sp. n. (C) Type locality.
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink