1094 lines
88 KiB
XML
1094 lines
88 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/herpetozoa.34.e69176" ID-GBIF-Dataset="f66c5bd4-4589-4834-87b6-7357484635db" ID-GBIF-Taxon="190234780" ID-Pensoft-Pub="2682-955X-34-233" ID-Pensoft-UUID="FFCB4A62CCBA5446923C8F77B4D3A4DF" LSID="urn:lsid:zoobank.org:act:6D7EA79C-98E3-4287-92A9-E81C1831D6CA" ModsDocID="2682-955X-34-233" checkinTime="1636677274783" checkinUser="pensoft" docAuthor="Gruenwald, Christoph I., Toribio-Jimenez, Sarahi, Montano-Ruvalcaba, Carlos, Franz-Chavez, Hector, Penaloza-Montano, Miguel A., Barrera-Nava, Eduardo Y., Jones, Jason M., Rodriguez, Christopher M., Hughes, India M., Strickland, Jason L. & Reyes-Velasco, Jacobo" docDate="2021" docId="4ECD0BA497425AF2BBE6AB5DA8FC3608" docLanguage="en" docName="Herpetozoa 34: 233-257" docOrigin="Herpetozoa 34" docPubDate="2021-11-11" docSource="http://dx.doi.org/10.3897/herpetozoa.34.e69176" docTitle="Tropidodipsas papavericola Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco 2021, sp. nov." docType="treatment" docVersion="4" id="FFCB4A62CCBA5446923C8F77B4D3A4DF" lastPageNumber="233" masterDocId="FFCB4A62CCBA5446923C8F77B4D3A4DF" masterDocTitle="Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species" masterLastPageNumber="257" masterPageNumber="233" pageId="0" pageNumber="233" updateTime="1637284501310" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of snail-eating snakes of the genus Tropidodipsas (Serpentes, Dipsadidae) from southern Mexico, with notes on related species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Gruenwald, Christoph I.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5994-555X</mods:nameIdentifier>
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<mods:affiliation>Biencom Real Estate, Carretera Chapala-Jocotepec # 57 - 1, C. P. 45920, Ajijic, Jalisco, Mexico. & Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico.</mods:affiliation>
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<mods:nameIdentifier type="email">cgruenwald@switaki.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Toribio-Jimenez, Sarahi</mods:namePart>
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<mods:affiliation>Centro de Ciencias de Desarrollo Regional, Universidad Autonoma de Guerrero. Calle Pino s / n. Col. El Roble. C. P. 39640. Acapulco, Guerrero, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Montano-Ruvalcaba, Carlos</mods:namePart>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Franz-Chavez, Hector</mods:namePart>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Penaloza-Montano, Miguel A.</mods:namePart>
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<mods:affiliation>Facultad de Ciencias Quimico Biologicas, Universidad Autonoma de Guerrero. Avenida Lazaro Cardenas # 88, Ciudad Universitaria Sur, C. P. 39074, Chilpancingo de los Bravo, Guerrero, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barrera-Nava, Eduardo Y.</mods:namePart>
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<mods:affiliation>Facultad de Ciencias Quimico Biologicas, Universidad Autonoma de Guerrero. Avenida Lazaro Cardenas # 88, Ciudad Universitaria Sur, C. P. 39074, Chilpancingo de los Bravo, Guerrero, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jones, Jason M.</mods:namePart>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodriguez, Christopher M.</mods:namePart>
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<mods:affiliation>Los Angeles Zoo and Botanical Gardens, 45333 Zoo Drive, Los Angeles, P. C. 90027, California, USA.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hughes, India M.</mods:namePart>
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<mods:affiliation>Department of Biology, University of South Alabama, 5871 USA Dr. N., Mobile, P. C. 36688, Alabama, USA.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Strickland, Jason L.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-1927-7259</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, University of South Alabama, 5871 USA Dr. N., Mobile, P. C. 36688, Alabama, USA.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Reyes-Velasco, Jacobo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8787-9980</mods:nameIdentifier>
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<mods:affiliation>Biodiversa A. C., Avenida de la Ribera # 203, C. P. 45900, Chapala, Jalisco, Mexico. & Herp. mx A. C. C. P. 28989, Villa de Alvarez, Colima, Mexico.</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Herpetozoa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-11-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>233</mods:start>
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<mods:end>257</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/herpetozoa.34.e69176</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/herpetozoa.34.e69176</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2682-955X-34-233</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFCB4A62CCBA5446923C8F77B4D3A4DF</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="233" type="results">
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<treatment ID-GBIF-Taxon="190234780" LSID="urn:lsid:zoobank.org:act:6D7EA79C-98E3-4287-92A9-E81C1831D6CA" httpUri="http://treatment.plazi.org/id/4ECD0BA497425AF2BBE6AB5DA8FC3608" lastPageNumber="233" pageId="0" pageNumber="233">
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<subSubSection pageId="0" pageNumber="233" type="nomenclature">
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<paragraph pageId="0" pageNumber="233">
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<taxonomicName LSID="http://zoobank.org/6D7EA79C-98E3-4287-92A9-E81C1831D6CA" authority="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco, 2021" authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola" status="sp. nov.">Tropidodipsas papavericola</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="233">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="233" type="description">
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<paragraph pageId="0" pageNumber="233">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tropidodipsas papavericola sp. nov. Holotype (INIRENA 2801). Holotype (INIRENA 2801) in life (A); Ventral aspect of Holotype (INIRENA 2801) in preservative (B, E); Dorsal aspect of Holotype (INIRENA 2801) in preservative." figureDoi="10.3897/herpetozoa.34.e69176.figure4" httpUri="https://binary.pensoft.net/fig/609885" pageId="0" pageNumber="233">Figs 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="233" type="paratypes/">
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<paragraph pageId="0" pageNumber="233">Paratypes/</paragraph>
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<paragraph pageId="0" pageNumber="233">
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<emphasis bold="true" pageId="0" pageNumber="233">(5)</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5</figureCitation>
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) INIRENA 2802 (original field number CIG 1496, Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5C</figureCitation>
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). Sub-adult male, collected AOR 18.1 km S of Puerto del Gallo on road from Nuevo Dehli to Puerto del Gallo, Municipio de Atoyac de
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<normalizedToken originalValue="Álvarez">Alvarez</normalizedToken>
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(17.4342°, -100.191°, datum=WGS84, 1654 m a.s.l.), Guerrero, Mexico on June 29th, 2019 by Christoph I.
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<normalizedToken originalValue="Grünwald">Gruenwald</normalizedToken>
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, Miguel A.
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<normalizedToken originalValue="Peñaloza-Montaño">Penaloza-Montano</normalizedToken>
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, Eduardo Y. Barrera-Nava, Karen I. Morales-Flores and Janelle Morales-Flores. INIRENA 2803 (original field number CIG 1502). Sub-adult male collected DOR at 5 km S of La Laguna, on road from San Luis La Loma to Bajitos de la Laguna, Municipio de
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<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
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de Galeana (17.5137°, -100.7737°, datum=WGS84, 1686 m a.s.l.), Guerrero, Mexico on June 29th, 2019 by Jason M. Jones and Carlos
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<normalizedToken originalValue="Montaño-Ruvalcaba">Montano-Ruvalcaba</normalizedToken>
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. INIRENA 2804 (original field number CIG 1632, Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5A-B</figureCitation>
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). Adult male, collected at Bajitos de la Laguna, Municipio de
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<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
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de Galeana, Guerrero, Mexico on October 30th, 2019 and retrieved by Alejandro Lara and donated to Jason M. Jones. INIRENA 2805 (original field number CIG 1457). Sub-adult male, collected at Jaguar Research Facility, Municipio de
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<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
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de Galeana, Guerrero, Mexico on June 10th, 2019 by Miguel A.
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<normalizedToken originalValue="Peñaloza-Montaño">Penaloza-Montano</normalizedToken>
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and Eduardo Y. Barrera-Nava. INIRENA 2810 (JRV 0362, Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Tropidodipsas tricolor sp. nov. Variation. Female from Municipio de Leonardo Bravo, Guerrero (A); Male from Municipio de Leonardo Bravo, Guerrero (B); Photos A-B by Peter Heimes. Female (INIRENA 2798) from Municipio de Malinaltepec, Guerrero (C-D)." figureDoi="10.3897/herpetozoa.34.e69176.figure2" httpUri="https://binary.pensoft.net/fig/609883" pageId="0" pageNumber="233">2D</figureCitation>
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). Juvenile male, collected DOR at 4.2 km S of La Laguna, on San Luis San Pedro - La Laguna Rd., Municipio de
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<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
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de Galeana (17.5173°, -100.7715°, datum=WGS84, 1620 m a.s.l.), Guerrero, Mexico on July 17th, 2020 by Jason M. Jones.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="233" type="diagnosis">
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<paragraph pageId="0" pageNumber="233">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="233">
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<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola">
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<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas papavericola</emphasis>
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</taxonomicName>
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sp. nov. is placed in the genus
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<taxonomicName authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas</emphasis>
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</taxonomicName>
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based on phylogenetic evidence (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Bayesian phylogenetic inference of members of the Dipsadidae based on two mitochondrial loci and two nuclear loci. All nodes with support of less than 0.5 are collapsed, while those with posterior support equal to 1 are marked with a black dot. New species described here in bold." figureDoi="10.3897/herpetozoa.34.e69176.figure6" httpUri="https://binary.pensoft.net/fig/609887" pageId="0" pageNumber="233">6</figureCitation>
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). It belongs to the
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<taxonomicName authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas fasciata" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="fasciata">
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<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas fasciata</emphasis>
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</taxonomicName>
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group as defined by
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<bibRefCitation DOI="https://doi.org/10.2307/1564485" author="Kofron, CP" journalOrPublisher="Journal of Herpetology" pageId="0" pageNumber="233" pagination="210 - 225" refId="B29" refString="Kofron, CP, 1987. Systematics of Neotropical gastropod-eating snakes: the fasciata group of the genus Sibon. Journal of Herpetology 21 (3): 210 - 225, DOI: https://doi.org/10.2307/1564485" title="Systematics of Neotropical gastropod-eating snakes: the fasciata group of the genus Sibon." url="https://doi.org/10.2307/1564485" volume="21" year="1987">Kofron (1987)</bibRefCitation>
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based on possessing a laterally compressed body, head distinctly wider than neck, protruding eyes of moderate to large diameter, vertebral and paravertebral scales not wider than other dorsal scales, 17 maxillary teeth, 22 dentary teeth, postmental scale absent. The species differs from all described species of
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<taxonomicName authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas</emphasis>
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</taxonomicName>
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recognized herein based on the following combination of characters: (1) 15/15/15 rows of smooth dorsal scales with no enlarged vertebral row; (2) prefrontal broader than long, not entering orbit; (3) loreal almost square, slightly longer than broad, not entering orbit; (4) 179-189 ventral scales in males, unknown in females; (5) 69-76 divided subcaudals in males, unknown in females; (6) 25-34 pale bands on body and 10-16 pale bands on tail; (7) 26-36 dark bands on body and 10-16 on tail, irregular with faded pale centers in adult individuals; (8) TL/SVL 0.30-0.35 in males, unknown in females; (9) iris dark chocolate-gray in life.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="233" type="comparisons">
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<paragraph pageId="0" pageNumber="233">Comparisons.</paragraph>
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<paragraph pageId="0" pageNumber="233">
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<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola">
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<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas papavericola</emphasis>
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</taxonomicName>
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sp. nov. is most similar to
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<taxonomicName genus="T." lsidName="T. philippii" pageId="0" pageNumber="233" rank="species" species="philippii">
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<emphasis italics="true" pageId="0" pageNumber="233">T. philippii</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="T." lsidName="T. fasciata" pageId="0" pageNumber="233" rank="species" species="fasciata">
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<emphasis italics="true" pageId="0" pageNumber="233">T. fasciata</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="T." lsidName="T. guerreroensis" pageId="0" pageNumber="233" rank="species" species="guerreroensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. guerreroensis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="T." lsidName="T. tricolor" pageId="0" pageNumber="233" rank="species" species="tricolor">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. tricolor</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. It is the only species of
|
|
<taxonomicName authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas</emphasis>
|
|
</taxonomicName>
|
|
in Mexico with a nebulated pattern of obscured body rings, although the nearly sympatric
|
|
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Dipsadidae" genus="Sibon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sibon nebulatus" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="nebulatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Sibon nebulatus</emphasis>
|
|
</taxonomicName>
|
|
exhibits a similar pattern as do several species of snail-suckers in northwestern South America (
|
|
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.766.24523" author="Arteaga, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="233" pagination="79 - 147" refId="B3" refString="Arteaga, A, Salazar-Valenzuela, D, Mebert, K, Penafiel, N, Aguiar, G, Sanchez-Nivicela, JC, Pyron, RA, Colston, TJ, Cisneros-Heredia, DF, Yanez-Munoz, MH, 2018. Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru. ZooKeys 766: 79 - 147, DOI: https://doi.org/10.3897/zookeys.766.24523" title="Systematics of South American snail-eating snakes (Serpentes, Dipsadini), with the description of five new species from Ecuador and Peru." url="https://doi.org/10.3897/zookeys.766.24523" volume="766" year="2018">Arteaga et al. 2018</bibRefCitation>
|
|
). It is distinguished from other Mexican snail-suckers such as the
|
|
<taxonomicName baseAuthorityName="Wagler" baseAuthorityYear="1830" class="Reptilia" family="Dipsadidae" genus="Geophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geophis chalybeus" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="chalybeus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Geophis chalybeus</emphasis>
|
|
</taxonomicName>
|
|
species group, the
|
|
<taxonomicName authorityName="Gunther" authorityYear="1893" class="Reptilia" family="Dipsadidae" genus="Geophis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Geophis omiltemanus" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="omiltemanus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Geophis omiltemanus</emphasis>
|
|
</taxonomicName>
|
|
species group,
|
|
<taxonomicName genus="G." lsidName="G. sartorii" pageId="0" pageNumber="233" rank="species" species="sartorii">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">G. sartorii</emphasis>
|
|
</taxonomicName>
|
|
new comb.,
|
|
<taxonomicName genus="G." lsidName="G. annuliferus" pageId="0" pageNumber="233" rank="species" species="annuliferus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">G. annuliferus</emphasis>
|
|
</taxonomicName>
|
|
new comb.,
|
|
<taxonomicName baseAuthorityName="Boulenger" baseAuthorityYear="1894" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas fischeri" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="fischeri">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas fischeri</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Similar looking but not closely related snail-suckers from southern Mexico. Sibon nebulatus from Municipio de Ixtlahuacan, Colima (A); Sibon nebulatus from Municipio de Las Margaritas, Chiapas (B); Tropidodipsas fischeri from Municipio de Rayon, Chiapas (C); Tropidodipsas fischeri from Municipio de Union Juarez, Chiapas (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure9" httpUri="https://binary.pensoft.net/fig/609890" pageId="0" pageNumber="233">9C,D</figureCitation>
|
|
),
|
|
<taxonomicName genus="S." lsidName="S. carri" pageId="0" pageNumber="233" rank="species" species="carri">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">S. carri</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="G." lsidName="G. sanniolus" pageId="0" pageNumber="233" rank="species" species="sanniolus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">G. sanniolus</emphasis>
|
|
</taxonomicName>
|
|
new comb. and
|
|
<taxonomicName genus="S." lsidName="S. linearis" pageId="0" pageNumber="233" rank="species" species="linearis">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">S. linearis</emphasis>
|
|
</taxonomicName>
|
|
by the laterally compressed body shape and and the nebulated pattern of broken up dorsal bands on a gray or brown background. It differs from
|
|
<taxonomicName genus="S." lsidName="S. anthracops" pageId="0" pageNumber="233" rank="species" species="anthracops">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">S. anthracops</emphasis>
|
|
</taxonomicName>
|
|
(character states in parenthesis) by possessing 15 scale rows at midbody (vs. 13), possessing a nebulated dorsal color pattern (vs. tricolor banded pattern), incomplete banding on venter (vs. complete), and lacking a loreal that enters the orbit (vs. loreal entering orbit). It may be distinguished from
|
|
<taxonomicName genus="S." lsidName="S. dimidiatus" pageId="0" pageNumber="233" rank="species" species="dimidiatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">S. dimidiatus</emphasis>
|
|
</taxonomicName>
|
|
by lacking a post-mental scale (usually one or two post-mental scales present), and possessing less than 80 subcaudals in males (vs. more than 95).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
Distinct from the superficially similar
|
|
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Reptilia" family="Dipsadidae" genus="Sibon" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sibon nebulatus" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="nebulatus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Sibon nebulatus</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Similar looking but not closely related snail-suckers from southern Mexico. Sibon nebulatus from Municipio de Ixtlahuacan, Colima (A); Sibon nebulatus from Municipio de Las Margaritas, Chiapas (B); Tropidodipsas fischeri from Municipio de Rayon, Chiapas (C); Tropidodipsas fischeri from Municipio de Union Juarez, Chiapas (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure9" httpUri="https://binary.pensoft.net/fig/609890" pageId="0" pageNumber="233">9A, B</figureCitation>
|
|
) by vertebral row of dorsal scales same size as other dorsal scales (vs. vertebral row of dorsal scales 1.25-1.35 times larger than other dorsal scales), smaller adult body size of 450-560 mm in males (vs. up to 890 mm in males), fewer dark body bands, 26-35 on body (vs. 36-47 on body), head predominately dark brown or black (vs. head pattern speckled with dark and light marking) and absence of pink speckling in the dorsal pattern (vs. pink speckling present).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
It differs from
|
|
<taxonomicName baseAuthorityName="Oliver" baseAuthorityYear="1937" class="Reptilia" family="Dipsadidae" genus="Dipsas" kingdom="Animalia" lsidName="Dipsas gaigeae" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="gaigeae">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Dipsas gaigeae</emphasis>
|
|
</taxonomicName>
|
|
by having 15 dorsal scale rows at midbody (vs. 13), possessing 179-189 ventral scales (vs. 155-169), loreal that does not enter orbit (vs. loreal enters orbit), and 26-35 pale body bands (vs. 7-12). Distinct from
|
|
<taxonomicName baseAuthorityName="Cope" baseAuthorityYear="1866" class="Reptilia" family="Dipsadidae" genus="Dipsas" kingdom="Animalia" lsidName="Dipsas brevifacies" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="brevifacies">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Dipsas brevifacies</emphasis>
|
|
</taxonomicName>
|
|
by possessing 179-189 ventral scakes (vs. 162-180), prefrontal and loreal not in contact with orbit (vs. variable in both), usually possessing 1+2 temporals (vs. usually 2+3), usually possessing one pair of infralabials in contact after the mental (vs. usually two pairs of infralabials in contact after the mental). Within
|
|
<taxonomicName authorityName="Gunther" authorityYear="1858" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName genus="T." lsidName="T. papavericola" pageId="0" pageNumber="233" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName genus="T." lsidName="T. fasciata" pageId="0" pageNumber="233" rank="species" species="fasciata">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. fasciata</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName genus="T." lsidName="T. guerreroensis" pageId="0" pageNumber="233" rank="species" species="guerreroensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. guerreroensis</emphasis>
|
|
</taxonomicName>
|
|
by possessing 15 smooth dorsal scale rows (vs. 17 keeled scale rows).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName genus="T." lsidName="T. tricolor" pageId="0" pageNumber="233" rank="species" species="tricolor">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. tricolor</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by lacking the tricolor outline in the dorsal bands (vs. possessing a tricolor outline in dorsal bands), 26-35 dorsal dark bands on body (vs. 19-22), by possessing a prefrontal which does not enter orbit (vs. prefrontal entering orbit), 2 preoculars (vs. 1), 5-7 supralabials (vs. 7-8), 6-7 infralabials (vs. 8-9), less subcaudal scales, 69-76 in males (vs. 78-79 in males), by possessing a pale ventral coloration of head with irregularly scattered dark spots, and a black and white mental (vs. pale ventral coloration of head with heavy dark stippling anteriorly and black mental), and by a smaller eye - head length ratio of 17-21% (vs. 25%). It is most similar to
|
|
<taxonomicName genus="T." lsidName="T. philippii" pageId="0" pageNumber="233" rank="species" species="philippii">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. philippii</emphasis>
|
|
</taxonomicName>
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Closely related species of Tropidodipsas from southern Mexico. Tropidodipsas fasciata from Municipio de Santo Domingo Zanatepec, Oaxaca (A-B); Tropidodipsas philippii from Municipio de Minatitlan, Colima (C-D)." figureDoi="10.3897/herpetozoa.34.e69176.figure8" httpUri="https://binary.pensoft.net/fig/609889" pageId="0" pageNumber="233">8C, D</figureCitation>
|
|
), from which it can be distinguished by having smooth dorsal scales (vs. slightly keeled), a longer tail which is 30-35% of SVL in males (vs. 25-29% in males), and less subcaudal scales, 69-76 in males (vs. 76-90 in males).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
Genetic divergence in a 1,072-bp long fragment of the mitochondrial
|
|
<emphasis italics="true" pageId="0" pageNumber="233">cytb</emphasis>
|
|
gene between
|
|
<taxonomicName genus="T." lsidName="T. papavericola" pageId="0" pageNumber="233" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName genus="T." lsidName="T. philippii" pageId="0" pageNumber="233" rank="species" species="philippii">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. philippii</emphasis>
|
|
</taxonomicName>
|
|
is 12-14%; between
|
|
<taxonomicName genus="T." lsidName="T. papavericola" pageId="0" pageNumber="233" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName genus="T." lsidName="T. guerreroensis" pageId="0" pageNumber="233" rank="species" species="guerreroensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. guerreroensis</emphasis>
|
|
</taxonomicName>
|
|
, 10-13%; between
|
|
<taxonomicName genus="T." lsidName="T. papavericola" pageId="0" pageNumber="233" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName genus="T." lsidName="T. fasciata" pageId="0" pageNumber="233" rank="species" species="fasciata">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. fasciata</emphasis>
|
|
</taxonomicName>
|
|
is 10-12%.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="description">
|
|
<paragraph pageId="0" pageNumber="233">Description of holotype.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tropidodipsas papavericola sp. nov. Holotype (INIRENA 2801). Holotype (INIRENA 2801) in life (A); Ventral aspect of Holotype (INIRENA 2801) in preservative (B, E); Dorsal aspect of Holotype (INIRENA 2801) in preservative." figureDoi="10.3897/herpetozoa.34.e69176.figure4" httpUri="https://binary.pensoft.net/fig/609885" pageId="0" pageNumber="233">4</figureCitation>
|
|
) INIRENA 2801. Adult male. SVL 421 mm; TL 137 mm; TotL 558 mm. HL 13.8 mm (from tip to posterior border of parietal) and 13.5 mm (from snout to posterior commissure of mouth); HW 9.8 mm, head distinct from body, approximately 2 times wider than neck. Snout long, SL 5.7 mm, contained 2.4 times in HL, rounded in dorsal and lateral profile. Loreal region tall, canthus rostralis angular. Rostral 1.2 times as broad as high (3.3 mm wide, 2.7 mm high); internasals broader than long (1.7 mm length, 2.8 mm width), rounded anteriorly, in lateral contact with anterior and posterior nasals; prefrontals longer than broad, (3.8 mm length, 2.9 mm width), rectangular, in median contact with each other and in lateral contact with postnasal, loreal, upper preocular and supraocular; frontal 1.1 times longer than broad, (4.4 mm long, 3.9 mm broad), pentagonal with angular tip posteriorly, in contact with prefrontals, supraoculars and parietals. Two moderately large preoculars on each side, upper larger than lower. Single supraocular large, in contact with upper preocular, prefrontal, frontal, parietal, and upper postocular. Two moderately large postoculars on each side, upper one larger than lower. Parietal 1.4 times as long as wide (6.0 mm long, 4.2 mm wide), the length of parietal 61% of HL, the common suture between parietals 4.4 mm, equal to frontal length. Nasal divided, prenasal slightly taller than postnasal, with combined length of both nasals 2.6 mm, longer than loreal. Loreal small, slightly longer than high and almost square, 1.4 high, 1.7 mm long on left side, 1.6 mm high, 1.8 mm long on right side, not reaching orbit. Eye moderately large, 3.1 mm, 23% of HL. Supralabials 7 on both sides, first in contact with nasals, second and third in contact with loreal, fourth and fifth entering orbit, sixth largest and in contact with lower postocular, anterior and posterior temporals. Temporals 1 + 2. Five nuchal scales in contact with parietals.
|
|
</paragraph>
|
|
<caption ID-Table-UUID="009715BE6CA3941E8945C20485E8A2FD" httpUri="http://table.plazi.org/id/009715BE6CA3941E8945C20485E8A2FD" pageId="0" pageNumber="233" start="Table 3" startId="T3">
|
|
<paragraph pageId="0" pageNumber="233">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">Table 3.</emphasis>
|
|
Meristic variation of
|
|
<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas tricolor" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="tricolor">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas tricolor</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. and
|
|
<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
<table pageId="0" pageNumber="233">
|
|
<tr pageId="0" pageNumber="233">
|
|
<th colspan="1" pageId="0" pageNumber="233" rowspan="1">Species</th>
|
|
<th colspan="2" pageId="0" pageNumber="233" rowspan="1">
|
|
<taxonomicName genus="T." lsidName="T. tricolor" pageId="0" pageNumber="233" rank="species" species="tricolor">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. tricolor</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
<th colspan="6" pageId="0" pageNumber="233" rowspan="1">
|
|
<taxonomicName genus="T." lsidName="T. papavericola" pageId="0" pageNumber="233" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov.
|
|
</th>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">Specimen</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2800</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2798</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2801</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2802</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2803</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2804</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2810</emphasis>
|
|
</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">INIRENA 2805</emphasis>
|
|
</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Sex</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Female</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Male</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Snout-vent Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">210</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">310</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">421</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">366</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">363</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">430</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">258</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">336</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Tail Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">67</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">93</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">137</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">119</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">113</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">127</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">91</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">112</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Total Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">277</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">403</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">558</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">485</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">476</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">557</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">349</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">448</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Head length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8.30</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">13.62</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">18.97</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15.25</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15.83</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">18.58</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">13.03</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">14.92</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Anterior Chinshield</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Posterior Chinshields</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Longer than wide</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Gulars</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Infras in contact with 1st CS</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Ventral Scales</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">183</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">183</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">180</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">180</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">188</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">189</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">182</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">179</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Subcaudal Scales</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">79 / 79</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">79 / 79</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">76</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">74 left / 74</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">73 / 73</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">69</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">74</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">73</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pre-oculars (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1*</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Post-oculars (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3. 3</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Anterior Temporals (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1. 1</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Posterior Temporals (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2. 2</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Supralabials (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8. 7</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">7. 8</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">7. 7</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 7</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5. 6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">7. 7</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Infralabials (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8. 8</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8. 9</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">7. 6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">7. 7</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6. 6</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Post-mentals</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pale Body Bands (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">19. 19</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">19. 20</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">32. 28</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">34. 35</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">29. 27</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">27. 27</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">25. 25</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pale Tail Bands (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">9. 9</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8. 8</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">14. 12</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">11. 11</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15. 16</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">13. 13</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Black Body Blotches (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">19. 19</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">19. 21</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">32. 33</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">30. 31</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">35. 35</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">27. 27</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">29. 28</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">26. 26</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Black Tail Blotches (Side)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8. 8</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">8. 8</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">14. 14</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15. 15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">-</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">10. 11</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">16. 14</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">13. 13</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Total Pale Body Blotch Count (Body & Tail)</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">28. 28</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">28. 28</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">46. 47</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">45. 46</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">42. 42</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">37. 38</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">45. 42</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">39. 39</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">TL/TotL</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.24</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.23</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.25</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.25</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.24</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.23</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.26</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.25</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">TL/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.32</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.30</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.33</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.33</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.31</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.30</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.35</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.33</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Internasal Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.00</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.34</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.70</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.34</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.50</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.46</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.90</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.44</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Internasal Width</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.60</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.62</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.48</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.53</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.58</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.74</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.96</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.52</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Loreal Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.9 / 1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.19 / 1.10</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.73 / 1.70</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.46 / 1.38</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.36 / 1.34</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.26 / 1.22</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.0 / 1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.40 / 1.38</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Loreal Width</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.0 / 1.0</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.24 / 1.12</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.84 / 1.82</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.45 / 1.2</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.38 / 1.42</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.40 / 1.46</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.92 / 0.94</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.30 / 1.26</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Loreal into orbit</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pre-frontal Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.70</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.34</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.39</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.81</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.89</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.26</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.48</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.36</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pre-frontal Width</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.00</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.64</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5.75</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.69</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.69</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5.06</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.84</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.33</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Prefrontal into orbit</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">No</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Frontal Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.80</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.88</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.23</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.58</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.71</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.93</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.31</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.60</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Frontal Width</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.40</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.70</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.81</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.73</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.35</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.35</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.71</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.08</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Parietal Length</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.30</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5.54</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6.58</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6.82</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5.51</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">6.26</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.61</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5.28</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Length Anterior Chinshields</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.4 / 2.6</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.49 / 3.63</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.73</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.03</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">4.31</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">5.72</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.33</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.58</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Length Posterior Chinshields</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.9 / 1.8</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.28 / 2.42</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.72</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.94</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.25</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.67</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">1.81</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.15</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Eye Diameter</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.10</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.80</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.20</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.02</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.11</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">3.56</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.71</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">2.85</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">ED/HL</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.25</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.21</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.17</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.20</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.20</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.19</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.21</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.19</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pre-Frontal Length/Width</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.85</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.89</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.59</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.60</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.62</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.64</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.65</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.55</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">HL/SVL</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.04</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.04</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.05</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.04</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.04</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.04</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.05</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">0.04</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Midbody scale rows</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Head scale rows</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Caudal scale rows</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">15</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Lateral compresion</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Extreme</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Extreme</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Oval</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Oval</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Oval</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Oval</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Oval</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes - Oval</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Anal plate</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Undivided</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Penultimate supralabial enlarged</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Fused</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">First infraliabals in contact</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
</tr>
|
|
<tr pageId="0" pageNumber="233">
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Pale nuchal band</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
<td colspan="1" pageId="0" pageNumber="233" rowspan="1">Yes</td>
|
|
</tr>
|
|
</table>
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">Mental 2.4 times as broad as long (2.6 mm broad, 1.1 mm long), flat anteriorly, triangular posteriorly and separated from the anterior chinshields by the first pair of enlarged infralabials which are in contact with each other. Infralabials 7/ 8; on the left 1-5 in contact with anterior chinshields and fifth in contact with posterior chinshield; on the right 1-6 in contact with anterior chinshields and sixth in contact with posterior chinshield. Anterior chinshields irregular, both 2.3 times as long as wide (left 4.6 mm long and 2.0 mm wide, right 4.4 mm long and 1.9 mm wide). Left posterior chinshield 1.6 times as long as wide (3.5 mm long, 2.0 mm wide), and right posterior chinshield 1.3 times as long as wide (2.6 mm long, 2.0 mm wide). Four gular scales. Infralabials and scales in the chin region smooth. Dorsal scales in 15-15-15 rows, smooth throughout body; apical pits not evident. Ventrals 180; anal plate single; 76 paired subcaudal scales. Body laterally compressed. Pupil elliptical.</paragraph>
|
|
<caption doi="10.3897/herpetozoa.34.e69176.figure4" httpUri="https://binary.pensoft.net/fig/609885" pageId="0" pageNumber="233" start="Figure 4" startId="F4">
|
|
<paragraph pageId="0" pageNumber="233">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">Figure 4.</emphasis>
|
|
<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. Holotype (INIRENA 2801). Holotype (INIRENA 2801) in life (A); Ventral aspect of Holotype (INIRENA 2801) in preservative (B,E); Dorsal aspect of Holotype (INIRENA 2801) in preservative.
|
|
</paragraph>
|
|
</caption>
|
|
<caption doi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233" start="Figure 5" startId="F5">
|
|
<paragraph pageId="0" pageNumber="233">
|
|
<emphasis bold="true" pageId="0" pageNumber="233">Figure 5.</emphasis>
|
|
<taxonomicName authorityName="Grünwald & Toribio-Jiménez & Montaño-Ruvalcaba & Franz-Chávez & Peñaloza-Montaño & Barrera-Nava & Jones & Rodriguez & Hughes & Strickland & Reyes-Velasco" authorityYear="2021" class="Reptilia" family="Colubridae" genus="Tropidodipsas" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tropidodipsas papavericola" order="Squamata" pageId="0" pageNumber="233" phylum="Chordata" rank="species" species="papavericola">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Tropidodipsas papavericola</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. Variation. Male (INIRENA 2804) from Municipio de
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Guerrero (D).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="coloration in life of the holotype">
|
|
<paragraph pageId="0" pageNumber="233">Coloration in life of the holotype</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
(Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tropidodipsas papavericola sp. nov. Holotype (INIRENA 2801). Holotype (INIRENA 2801) in life (A); Ventral aspect of Holotype (INIRENA 2801) in preservative (B, E); Dorsal aspect of Holotype (INIRENA 2801) in preservative." figureDoi="10.3897/herpetozoa.34.e69176.figure4" httpUri="https://binary.pensoft.net/fig/609885" pageId="0" pageNumber="233">4A</figureCitation>
|
|
). Dorsal coloration of head, body and tail brown with 32/33 irregular dark brown bands on the body and neck which all have pale centers that are stippled with paler brown and pale cream. Pale brown nuchal band two body scale rows long but reduced to one at the mid-dorsal line; not involving the posterior part of the parietals. Dark bands on body and tail 3-7 scale rows long, widest mid-dorsally, and alternating on posterior half of body. Head coloration predominately dark brown on dorsal surface and with light brown mottling and pale cream on rostral and anterior labials. First three supralabials bicolor pale cream and dark brown, posterior two supralabials bicolor dark brown with pale brown. Ventral coloration of head pale cream with irregular dark brown blotches. Ventral coloration of body pale yellowish cream, with dark and pale bands incomplete on venter. The ventral pattern consists of longitudinal rows of dark spots that connect to form irregular longitudinal lines. Anal plate bicolor, dark brown and pale cream. All 15 dark bands on tail incomplete ventrally, dark coloration forming longitudinal lines ventrally on tail. Tail tip pale cream with black tip. Pupil black, iris dark chocolate-gray.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="coloration in preservative of the holotype">
|
|
<paragraph pageId="0" pageNumber="233">Coloration in preservative of the holotype</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
(Figs
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Tropidodipsas papavericola sp. nov. Holotype (INIRENA 2801). Holotype (INIRENA 2801) in life (A); Ventral aspect of Holotype (INIRENA 2801) in preservative (B, E); Dorsal aspect of Holotype (INIRENA 2801) in preservative." figureDoi="10.3897/herpetozoa.34.e69176.figure4" httpUri="https://binary.pensoft.net/fig/609885" pageId="0" pageNumber="233">4B-E</figureCitation>
|
|
). General coloration bicolor. Dorsal surfaces of head, body and tail predominately dark brown with darker brown-black bands. Pale bands brown-gray, but whitish in a few places where
|
|
<emphasis italics="true" pageId="0" pageNumber="233">stratum corneum</emphasis>
|
|
has been removed (snake was going into shed when preserved). The dark bands remain brown-black and the pale cream faded to white.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="variation">
|
|
<paragraph pageId="0" pageNumber="233">Variation.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
Meristic variation of five available specimens given in Table
|
|
<tableCitation captionStart="Table 3" captionStartId="T3" captionText="Table 3. Meristic variation of Tropidodipsas tricolor sp. nov. and Tropidodipsas papavericola sp. nov." httpUri="http://table.plazi.org/id/009715BE6CA3941E8945C20485E8A2FD" pageId="0" pageNumber="233" tableUuid="009715BE6CA3941E8945C20485E8A2FD">3</tableCitation>
|
|
. Variation in color and pattern detailed below (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="color in life">
|
|
<paragraph pageId="0" pageNumber="233">Color in life.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
A sub-adult male paratype from above Nueva Dehli, Municipio de Atoyac de
|
|
<normalizedToken originalValue="Álvarez">Alvarez</normalizedToken>
|
|
, Guerrero (INIRENA 2802) had a pale brown body coloration with 31/29 dark brown bands on body and neck posterior to pale nuchal band and 14 on the tail. This specimen has light centers in all dark body bands, albeit limited to the lateral portions only. This implies that the irregular dark banding with light centers may be subject to ontogenetic variation, as this specimen is intermediate in size and pattern between the juvenile INIRENA 2810 and the large adult holotype INIRENA 2801. Neither the pale body bands, nor the dark body bands, are complete ventrally, reaching only to the lateral edges of the ventrals. The venter was cream or pale tan mid-ventrally with two dark brown irregular longitudinal stripes running down the length of the entire venter. The lines run into the remnants of the dark brown dorsal bands on the edges of the venter (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5C</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="color in preservative">
|
|
<paragraph pageId="0" pageNumber="233">Color in preservative.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
An adult male paratype from above Bajitos de la Laguna, Municipio de
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Guerrero (INIRENA 2804) has 29/27 dark bands on the body posterior to the pale nape band and 12 on the tail. This specimen differs from the holotype because it has a relatively well-defined pattern of dark and pale bands with regular edges. Furthermore, the first well-defined dark band on the neck does not have a pale center, and on the second dark body band the pale center is interrupted mid-dorsally. The venter is white and covered with irregular black spots that do not form longitudinal rows of spots nor are fused into lines and dashes (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5A, B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
A juvenile male paratype from above San Luis La Loma, Municipio de
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Guerrero (INIRENA 2810) has 28/29 dark bands on the body and neck posterior to the pale nuchal band and 16 on the tail. This specimen lacks any light centers in the dark body bands of the tail, suggesting that this color pattern may be subject to an ontogenetic shift from juveniles to adults. It has black stippling in the light interspaces reminiscent of
|
|
<taxonomicName genus="T." lsidName="T. tricolor" pageId="0" pageNumber="233" rank="species" species="tricolor">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">T. tricolor</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. Both the pale body bands and the dark body bands are not complete ventrally, reaching only onto the lateral edges of the ventrals. The venter (in preservative) is white mid-ventrally, with two irregular longitudinal stripes running down the length of the entire venter. The lines run into the remnants of the dark dorsal bands on the edges of the venter (Fig.
|
|
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Tropidodipsas papavericola sp. nov. Variation. Male (INIRENA 2804) from Municipio de Tecpan de Galeana, Guerrero (A-B); Male (INIRENA 2802) from Municipio de Leonardo Bravo, Guerrero (B); Juvenile male (INIRENA 2810) from Municipio de Tecpan de Galeana, Guerrero (D)." figureDoi="10.3897/herpetozoa.34.e69176.figure5" httpUri="https://binary.pensoft.net/fig/609886" pageId="0" pageNumber="233">5D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
An adult male paratype from above San Luis La Loma, Municipio de
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Guerrero (INIRENA 2803) has 35 dark bands on body and neck posterior to the pale nuchal band. None of the dark bands have pale centers mid-dorsally, some have the pale centers on the lateral portions of the body and others lack them altogether. Ventral coloration white with black blotches forming an irregular
|
|
<normalizedToken originalValue="“checkerboard”">"checkerboard"</normalizedToken>
|
|
pattern. No pale or dark body bands complete on venter.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
A sub-adult male paratype, from above
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Guerrero (INIRENA 2805) has 26 dark bands on body and neck posterior to the pale nuchal band, all with faded pale centers in the dark body bands, most of them extensive but incomplete mid-dorsally. It has 12 dark bands on the tail and a dark tail tip. Ventral coloration white with dark gray mottling, with no dark or pale dorsal bands complete on venter, but the dense mottling makes the dark dorsal bands on posterior portion of body connect in an alternating undulated pattern.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="dentition">
|
|
<paragraph pageId="0" pageNumber="233">Dentition.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">The holotype (INIRENA 2801) appears to have 14 maxillary teeth visible, however we did not remove the maxillary arch to avoid damaging the specimen.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
We observed
|
|
<emphasis italics="true" pageId="0" pageNumber="233">in situ</emphasis>
|
|
the maxillary arch and dentition of the holotype (INIRENA 2801): it has 14 maxillary teeth and 20-22 dentary teeth.
|
|
</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">An adult male paratype (INIRENA 2805) has 17 maxillary teeth (counting empty sockets) on the right side. We counted 22 dentary teeth on the right dentary of this same specimen.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="distribution">
|
|
<paragraph pageId="0" pageNumber="233">Distribution, habitat and ecology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
This species appears to be restricted to moderate elevations in the Sierra Madre del Sur in central Guerrero in two distinct areas (Fig.
|
|
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution map of species in the Tropidodipsas fasciata species group in Mexico. Triangles represent type localities, type localities are only mapped when known with certainty." figureDoi="10.3897/herpetozoa.34.e69176.figure10" httpUri="https://binary.pensoft.net/fig/609891" pageId="0" pageNumber="233">10</figureCitation>
|
|
). Habitats include cloud forest, mesic pine-oak woodland, tropical evergreen forest, tropical semi-deciduous forest and combinations of these assemblages. Known from around 1600-2200 m a.s.l. in the Sierra de
|
|
<normalizedToken originalValue="Técpan">Tecpan</normalizedToken>
|
|
de Galeana, Sierra de Atoyac and above Acapulco, in the foothills in the area encompassed between Acahuizotla and El Treinta. Specimens were found at night during the rainy season on low to medium (1.5-3.0 m) vegetation or crossing the road.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="233" type="etymology">
|
|
<paragraph pageId="0" pageNumber="233">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="233">
|
|
The specific epithet
|
|
<emphasis italics="true" pageId="0" pageNumber="233">papaver</emphasis>
|
|
+
|
|
<emphasis italics="true" pageId="0" pageNumber="233">cola</emphasis>
|
|
refers to the living among poppy plants (genus
|
|
<taxonomicName class="Magnoliopsida" family="Papaveraceae" genus="Papaver" kingdom="Plantae" lsidName="Papaver" order="Ranunculales" pageId="0" pageNumber="233" phylum="Tracheophyta" rank="genus">
|
|
<emphasis italics="true" pageId="0" pageNumber="233">Papaver</emphasis>
|
|
</taxonomicName>
|
|
) which are illegally planted throughout the range of this species for the extraction of opium gum.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</subSection>
|
|
</document> |