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<document ID-CLB-Dataset="55511" ID-DOI="http://dx.doi.org/10.3897/zookeys.1100.81910" ID-GBIF-Dataset="242e0609-936b-4f2f-9807-3f8ccbb76fcc" ID-PMC="PMC9848845" ID-Pensoft-Pub="1313-2970-1100-165" ID-Pensoft-UUID="E417C237A77155CF9B6AA7E492C73F73" ID-PubMed="36760389" ID-ZooBank="363D45514CC04BDA97A14A4AC1BEE469" ModsDocID="1313-2970-1100-165" checkinTime="1652785889463" checkinUser="pensoft" docAuthor="Kolesnikov, Vasiliy B., Christophoryova, Jana, Przhiboro, Andrey A. &amp; Turbanov, Ilya S." docDate="2022" docId="4DC9B50A46B756A2BA2498403F8C8405" docLanguage="en" docName="ZooKeys 1100: 165-190" docOrigin="ZooKeys 1100" docPubDate="2022-05-16" docSource="http://dx.doi.org/10.3897/zookeys.1100.81910" docTitle="Neobisium (Neobisium) adjaricum Kolesnikov &amp; Christophoryová &amp; Przhiboro &amp; Turbanov 2022, sp. nov." docType="treatment" docUuid="A66BABF5-698C-4B29-888D-12945CE6FADB" docUuidSource="ZooBank" docVersion="6" id="E417C237A77155CF9B6AA7E492C73F73" lastPageNumber="165" masterDocId="E417C237A77155CF9B6AA7E492C73F73" masterDocTitle="The pseudoscorpions of the Caucasian Sphagnum bogs: part I. Description of Neobisium (Neobisium) adjaricum sp. nov. and redescription of the holotype of N. (N.) vilcekii Krumpal, 1983 (Arachnida, Pseudoscorpiones, Neobisiidae)" masterLastPageNumber="190" masterPageNumber="165" pageNumber="165" updateTime="1732848613000" updateUser="ExternalLinkService">
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<mods:title id="577AFB0AC35048D27AE442FDDB84A4FA">The pseudoscorpions of the Caucasian Sphagnum bogs: part I. Description of Neobisium (Neobisium) adjaricum sp. nov. and redescription of the holotype of N. (N.) vilcekii Krumpal, 1983 (Arachnida, Pseudoscorpiones, Neobisiidae)</mods:title>
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<mods:namePart id="A7C41C0E3BA3ADE76F65B77000CFA728">Kolesnikov, Vasiliy B.</mods:namePart>
<mods:nameIdentifier id="1A88247776B867225C52AF642F4592FF" type="ORCID">https://orcid.org/0000-0001-6177-7858</mods:nameIdentifier>
<mods:affiliation id="270F33E09605F2436C942CE9FC61D47C">Federal Public Budgetary Scientific Institution All-Russian Research Institute of Protection of Plants, VNIISS, Voronezh Province, 396030, Russia &amp; Tyumen State University, 6 Volodarskogo Str., Tyumen, 625003, Russia</mods:affiliation>
</mods:name>
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<mods:role id="944FF113A319675AFEECCD5C3A1636DE">
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<mods:namePart id="1252ACCF30C43044416DDDFE63D9B88D">Christophoryova, Jana</mods:namePart>
<mods:nameIdentifier id="7D212D6986379CB40021567E0F639463" type="ORCID">https://orcid.org/0000-0002-3746-1367</mods:nameIdentifier>
<mods:affiliation id="545996CE604F064CBE7DA0D2D1FB296C">Department of Zoology, Faculty of Natural Sciences, Comenius University, Mlynska dolina, Ilkovicova 6, SK- 842 15 Bratislava, Slovakia</mods:affiliation>
</mods:name>
<mods:name id="965911D85C7FF5ABB365898BE39CB128" type="personal">
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<mods:namePart id="B26DFCF2C93C118D67787E4A24A4CB50">Przhiboro, Andrey A.</mods:namePart>
<mods:affiliation id="5789C387A4F35D523FCC165E1F1AD068">Zoological Institute, Russian Academy of Sciences, Universitetskaya nab. 1, Saint Petersburg, 199034, Russia</mods:affiliation>
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<mods:namePart id="6BD03E5EC85984BED65B1A7267EFFD9F">Turbanov, Ilya S.</mods:namePart>
<mods:affiliation id="836A06482A1C4FDBD37F2F27BC90C6C1">I. D. Papanin Institute of Biology of Inland Waters, Russian Academy of Sciences, Borok, Yaroslavl Province, 152742, Russia &amp; Cherepovets State University, Vologda Province, Cherepovets, 162600, Russia</mods:affiliation>
<mods:nameIdentifier id="6846AD35AA2BC013434D3730FF92F025" type="email">turba13@mail.ru</mods:nameIdentifier>
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<mods:date id="1ED2672BA32610249BB93DBB3D306D15">2022</mods:date>
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<mods:number id="0CF3959B7A704253F094DC5C3CF17ACA">2022-05-16</mods:number>
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<mods:number id="0DCE97A1B522A3DAA3C1338C696E5E8E">1100</mods:number>
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<treatment id="4DC9B50A46B756A2BA2498403F8C8405" ID-GBIF-Taxon="195350942" LSID="urn:lsid:zoobank.org:act:A66BABF5-698C-4B29-888D-12945CE6FADB" httpUri="http://treatment.plazi.org/id/4DC9B50A46B756A2BA2498403F8C8405" lastPageNumber="165" pageId="0" pageNumber="165" scope_class="Arachnida" scope_family="Neobisiidae" scope_order="Pseudoscorpiones" scope_subGenus="Neobisium">
<subSubSection id="63139030E3625066FDE0B08C20720BE1" pageId="0" pageNumber="165" type="nomenclature">
<paragraph id="7776EBE07109682CAC664D87A9CC0733" pageId="0" pageNumber="165">
<taxonomicName id="E04C4DF891767624594C994C227F9FBD" LSID="http://zoobank.org/A66BABF5-698C-4B29-888D-12945CE6FADB" authority="Kolesnikov &amp; Christophoryová &amp; Przhiboro &amp; Turbanov, 2022" authorityName="Kolesnikov &amp; Christophoryová &amp; Przhiboro &amp; Turbanov" authorityYear="2022" class="Arachnida" family="Neobisiidae" genus="Neobisium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisium (Neobisium) adjaricum" order="Pseudoscorpiones" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" status="sp. nov." subGenus="Neobisium">Neobisium (Neobisium) adjaricum</taxonomicName>
<taxonomicNameLabel id="7537BB28F03C5408CEDC610211C10C44" pageId="0" pageNumber="165">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C0FE7B237F640141B649D28F29CFE9C1" pageId="0" pageNumber="165" type="description">
<paragraph id="C689B2353C0C676ABDBD45FD708984AC" pageId="0" pageNumber="165">
<figureCitation id="A4F8E397AD33BF16BF6DC372BFAE0013" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. A habitus of Neobisium (Neobisium) adjaricum sp. nov., holotype male, dorsal view B permanent slide mount with Neobisium (Neobisium) vilcekii Krumpal, 1983, holotype male. Scale bar: 1 mm." figureDoi="10.3897/zookeys.1100.81910.figure3" httpUri="https://binary.pensoft.net/fig/684537" pageId="0" pageNumber="165">Figs 3A</figureCitation>
<figureCitation id="016F75CD7E24FA00C04C8A9AAB9646F8" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">, 4</figureCitation>
<figureCitation id="86C346A7540A9E2E430E0341047DF50D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Neobisium (Neobisium) adjaricum sp. nov., holotype male (A-D) and paratype male (E, F) A left chela, dorsal view B right chela, lateral view C right pedipalp (without chela), dorsal view D left chelicera, dorsal view E fixed finger of left chelicera, partial dorsal view F movable finger of left chelicera, partial dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, est - exterior sub-terminal, et - exterior terminal, ib - interior basal, isb - interior sub-basal, ist - interior sub-terminal, it - interior terminal; trichobothria on movable chelal finger: b - basal, sb - sub-basal, st - sub-terminal, t - terminal; gs - galeal seta; pc - coupled sensilla. Scale bars: 1.0 mm (A-C); 0.1 mm (D-F)." figureDoi="10.3897/zookeys.1100.81910.figure5" httpUri="https://binary.pensoft.net/fig/684539" pageId="0" pageNumber="165">, 5</figureCitation>
<figureCitation id="4363CEBE6C2C75C55CC6A12EF3F84B5F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">, 6</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="665CEA68CE2BEA9989CD98D48E87AC49" pageId="0" pageNumber="165" type="materials_examined">
<paragraph id="B5027F884FB5FFBD3BF8BE494AE69427" pageId="0" pageNumber="165">Material examined.</paragraph>
<paragraph id="C7564B6A0859DD506FDD4E789E61B45E" pageId="0" pageNumber="165">
<materialsCitation id="F29CF12B3F6B9F157763AA412F318C36" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. Przhiboro" country="Georgia" elevation="600" latitude="41.86461" location="Ispani" longLatPrecision="1" longitude="41.79763" municipality="Kobuleti" specimenCount="♂" stateProvince="Adjara" typeStatus="Holotype">
<emphasis id="1C38AF0EEFFC575F2FDA2E1B573C23AF" bold="true" pageId="0" pageNumber="165">
<emphasis id="7B99EC9B69B86ED564AC4A4789B450F2" italics="true" pageId="0" pageNumber="165">
<typeStatus id="931DBD7EA0304431DA9093BADF35AC24">Holotype</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="7E785A638FA14143A84204EB4168044C"></specimenCount>
(ZISP 1437),
<emphasis id="EA1F2CF82C41E9B8B4B5AEAC4730279C" bold="true" pageId="0" pageNumber="165">
<collectingCountry id="A2242959A107700B6F7D431C2837FAD0" name="Georgia">Georgia</collectingCountry>
</emphasis>
,
<collectingRegion id="D0062175EA4917A0612054C43D2E6B93" country="Georgia" name="Ajaria">Adjara</collectingRegion>
, near
<collectingMunicipality id="CCDA44F1D67E42939D4CA84ABCA918FA">Kobuleti</collectingMunicipality>
,
<location id="2FB8E1E65E491C60822F8FC8F8F31A64" LSID="urn:lsid:plazi:treatment:4DC9B50A46B756A2BA2498403F8C8405:2FB8E1E65E491C60822F8FC8F8F31A64" country="Georgia" latitude="41.86461" longLatPrecision="1" longitude="41.79763" municipality="Kobuleti" name="Ispani" stateProvince="Adjara">Ispani</location>
2 bog,
<geoCoordinate id="24F191709C44847279F8E762B28F8392" degrees="41.86461" direction="north" orientation="latitude" precision="1" value="41.86461">41.86461°N</geoCoordinate>
,
<geoCoordinate id="4E5259904D19396F6B0C7DCEA0E42E96" degrees="41.79763" direction="east" orientation="longitude" precision="1" value="41.79763">41.79763°E</geoCoordinate>
, main bog, site 3 (
<quantity id="8A1E0AFB081A1359241BBC80B9F56634" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">
<elevation id="36E5B0DCC5AA5D78C9F29415914B0E93" metricMagnitude="2" metricUnit="m" metricValue="6.0" unit="m" value="600.0">600 m</elevation>
</quantity>
E of W
<taxonomicName id="1D0145B145DFA83EE8141C7148101B82" class="Magnoliopsida" family="Betulaceae" genus="Alnus" kingdom="Plantae" lsidName="Alnus" order="Fagales" pageId="0" pageNumber="165" phylum="Tracheophyta" rank="genus">
<emphasis id="4A0603394A787A601A932FC437BBC4E4" italics="true" pageId="0" pageNumber="165">Alnus</emphasis>
</taxonomicName>
belt), flat
<taxonomicName id="F3418F14CC1360CC72CB12ABC5486D27" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="C575AD12E8AD379853E759AA25BDDF7E" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
carpet, sample I-11, 10.V.2019,
<collectorName id="788F2171110D440ACB057C7CD6F194EA">A. Przhiboro</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="437641B170E60DE76C99FFA9FDE9E921" pageId="0" pageNumber="165">
<emphasis id="E90B4BAF1B8A40ACD392A1535D7DE056" bold="true" pageId="0" pageNumber="165">
<emphasis id="828113CBADD6975C5F023337C5D331B6" italics="true" pageId="0" pageNumber="165">
<typeStatus id="8F476716CB12C5C0EFEF1B354ADF57DA">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<specimenCount id="D16DFA956DBD6DC1023B02F5D48C47BA" type="male">1 ♂</specimenCount>
(ZMMU TI-65),
<materialsCitation id="623755DCA8036C83DD470807DB47A1DD" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. Przhiboro" country="Georgia" elevation="150" latitude="41.85805" location="Ispani" longLatPrecision="1" longitude="41.7875" municipality="Kobuleti" specimenCount="1" specimenCount-female="1" stateProvince="Adjara" typeStatus="paratype">
<specimenCount id="FAFC7F86AFB8282DF30EE58374B9F304" type="female">1 ♀</specimenCount>
(ZISP 1438),
<emphasis id="CBA55C2ECEB51C2DE93B63427349977D" bold="true" pageId="0" pageNumber="165">
<collectingCountry id="CF232AC7333453E7003DA362BC0484A2" name="Georgia">Georgia</collectingCountry>
</emphasis>
,
<collectingRegion id="061305673810181AA036B53B2216D633" country="Georgia" name="Ajaria">Adjara</collectingRegion>
, near
<collectingMunicipality id="F71672EAD8E9E5A9855F74ED38BA803C">Kobuleti</collectingMunicipality>
,
<location id="E922E9AAD651AC799CB9D3036BB6A378" LSID="urn:lsid:plazi:treatment:4DC9B50A46B756A2BA2498403F8C8405:E922E9AAD651AC799CB9D3036BB6A378" country="Georgia" latitude="41.85805" longLatPrecision="1" longitude="41.7875" municipality="Kobuleti" name="Ispani" stateProvince="Adjara">Ispani</location>
1 bog,
<geoCoordinate id="910EE9AB1C48349981657B58827121F6" degrees="41.85805" direction="north" orientation="latitude" precision="1" value="41.85805">41.85805°N</geoCoordinate>
,
<geoCoordinate id="CD7C50DD1EC129798F13F53F4F0F2479" degrees="41.78750" direction="east" orientation="longitude" precision="1" value="41.7875">41.78750°E</geoCoordinate>
, site 1 (
<quantity id="AEF2AE1F8B369E3AE4A296D589682F05" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">
<elevation id="C94D88D7DD69DBD38620762B7A430BDD" metricMagnitude="2" metricUnit="m" metricValue="1.5" unit="m" value="150.0">150 m</elevation>
</quantity>
E of NW bog edge), dry
<taxonomicName id="67BFB19D962EBB7D313968855CCC2308" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="FAAF90E9DAAFB11C4AEF85784C44DC8D" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
hummocks, sample I-18, 11.V.2019,
<collectorName id="33A3DB24D09D745616C83C0578AD27CE">A. Przhiboro</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="BF511F6BD8A3CBA9E34CB83F1DE46632" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="A. Przhiboro" country="Georgia" elevation="150" latitude="41.85805" location="Ispani" longLatPrecision="1" longitude="41.7875" municipality="Kobuleti" specimenCount="1" specimenCount-male="1" stateProvince="Ajaria" typeStatus="paratype">
<specimenCount id="09EAE2670E84B7633CDC014205357011" type="male">1 ♂</specimenCount>
(ISEA Ps. 001.0037), the same locality, habitat, and collector, sample I-37, 10.X.2019
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE9BA869AC07653E48A593CCD6FB9E1" pageId="0" pageNumber="165" type="diagnosis">
<paragraph id="86E0612F6BB7B1A8D3116BC30AA7FF33" pageId="0" pageNumber="165">Diagnosis.</paragraph>
<paragraph id="FE8C0A894AB1AA5736CB603DCF9629EF" pageId="0" pageNumber="165">
Carapace with moderately long and apically rounded epistome; two pairs of eyes present, lenses of posterior eyes with low convexity; movable cheliceral finger without large median tooth; pedipalpal femur and patella smooth (sometimes very rare and weak granules on outer side of femur); chelal hand with faint reticulate ornament; notch on median side of patella reaching almost middle of patellar club length; movable finger distinctly longer than hand+ but almost equal to femur in length; trichobothrium
<emphasis id="749EBCD1F11C25C4C6413D6F377973A1" italics="true" pageId="0" pageNumber="165">ist</emphasis>
situated distal to middle of fixed chelal finger; distal half of fixed chelal finger with teeth almost equal in size and shape; anterolateral process of coxa of leg I pointed and slightly enlarged, its mediolateral process slightly prominent, denticulate; sternite II with 5 setae; pedipalpal femur 4.5-5.1
<normalizedToken id="6E26FF80D9117D248092BDF6C04765BB" originalValue="×">x</normalizedToken>
as long as broad (0.77-1.00/0.15-0.22), chelal hand+ 1.62
<normalizedToken id="8C909FF3DA7307D9A0C10550039DD0A9" originalValue="×">x</normalizedToken>
as long as broad (0.62-0.71/0.38-40); movable chelal finger length 0.79-0.95.
</paragraph>
</subSubSection>
<subSubSection id="6DE363265F3F4C332E5965BDB017A15C" pageId="0" pageNumber="165" type="description">
<paragraph id="43AB48601F47E5F5FA0F736C6D829FF6" pageId="0" pageNumber="165">Description.</paragraph>
<paragraph id="194EA5FF0EEB58CB7D494B60C6EA1DBA" pageId="0" pageNumber="165">♂ and ♀. Coloration: carapace distinctly darker than tergite I and slightly paler than pedipalp; pedipalpal segments uniformly coloured.</paragraph>
<paragraph id="1C33C004FF65DA8860C07819BEECA4AD" pageId="0" pageNumber="165">
Carapace (Figs
<figureCitation id="6FB4AA82EAE32C3A8DF387131D073AD2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4A</figureCitation>
,
<figureCitation id="1BA3C239959B974D43A292CFC9221805" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6C, H, J</figureCitation>
): with posterior border partially unsclerotised and pale, covered with fine reticulate ornament (Fig.
<figureCitation id="B17615973027D4770C525C1E8D449356" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6H</figureCitation>
), wider than long, 0.79
<normalizedToken id="EFB9E3A6AB40A93728F2A703C301FF80" originalValue="×">x</normalizedToken>
(0.83-0.9
<normalizedToken id="E27C1A851D62091FCD256481D60F476F" originalValue="×">x</normalizedToken>
) as long as broad, with two pairs of corneate eyes, anterior eyes slightly larger than posterior ones; distance between anterior margin of anterior eye and anterior margin of carapace 0.05, diameter of anterior eye 0.10 (0.08-0.09), diameter of posterior eye 0.09 (0.07-0.09), distance between eyes 0.02; carapace with 27 (23-24) setae, its anterior margin with 6 (4-6) setae: 4 macrosetae and 2 marginal microsetae (one paratype without these microsetae); posterior margin with 9 (6-7) setae, chaetotaxy: m4m:6:6:9 (m4m:6:6:6, 4:6:6:7); epistome prominent, short and apically rounded, 0.025/0.040; glandular pores present; anterolateral corners with two small protuberances located below surface of carapace; carapace with 6 (5-6) microlyrifissures: one pair situated in ocular zone, close to anterior eyes, and two pairs located at posterior margin.
</paragraph>
<caption id="64F9BC55E1569095C93DCB840514F47B" doi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165" start="Figure 4" startId="F4">
<paragraph id="213F77DDF7E1FA6B29BA557AFCFE0405" pageId="0" pageNumber="165">
<emphasis id="3DDCA3CA100D685DFEBEE5F8916CAFE9" bold="true" pageId="0" pageNumber="165">Figure 4.</emphasis>
<taxonomicName id="3CBE5ABB3E3BEDB6146C94A4CA4EE571" authorityName="Kolesnikov &amp; Christophoryová &amp; Przhiboro &amp; Turbanov" authorityYear="2022" class="Arachnida" family="Neobisiidae" genus="Neobisium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisium (Neobisium) adjaricum" order="Pseudoscorpiones" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="Neobisium">Neobisium (Neobisium) adjaricum</taxonomicName>
sp. nov., holotype male
<emphasis id="27610D6371500AA792E366F03B47F59B" bold="true" pageId="0" pageNumber="165">A</emphasis>
carapace, dorsal view
<emphasis id="41192D2C81707DDD430AE27BD28439B3" bold="true" pageId="0" pageNumber="165">B</emphasis>
coxae of pedipalp and legs I-IV, ventral view
<emphasis id="8CEB94CDB091A21A8DEC43FE898EF2E7" bold="true" pageId="0" pageNumber="165">C</emphasis>
sternites II-IV, ventral view
<emphasis id="F6C610A18087A8E8521782B4A1965096" bold="true" pageId="0" pageNumber="165">D</emphasis>
male genitalia, ventral view
<emphasis id="F355C3BEBCABA326EA9BA5A6E8712669" bold="true" pageId="0" pageNumber="165">E</emphasis>
right leg IV (without trochanter), dorsal view
<emphasis id="81AE5BD59F123C9DC2F006D0BCF58626" bold="true" pageId="0" pageNumber="165">F</emphasis>
sub-terminal seta on pedal tarsus IV. Abbreviation:
<emphasis id="7A93800015D9C64166931BC13917E79D" italics="true" pageId="0" pageNumber="165">dat</emphasis>
- dorsal accessory tooth. Scale bars: 1.0 mm (
<emphasis id="17ED87A4B2B8EC90A8C631FF26C0524A" bold="true" pageId="0" pageNumber="165">A-C, E</emphasis>
); 0.1 mm (
<emphasis id="7C853AE90E79E5D9F1F345DB57770E97" bold="true" pageId="0" pageNumber="165">D, F</emphasis>
).
</paragraph>
</caption>
<paragraph id="493A5A6C5115DA6119A69A6612507EE6" pageId="0" pageNumber="165">Tergites weakly sclerotised; all setae simple; tergite X with 2 (2-4) pseudotactile and 2 tactile setae; tergite XI with 4 tactile setae; chaetotaxy: 6:9:11:11:11:12:12:13:11:4T2T4:T2T2T2T:2 (6:8-9:10-11:11-12:11:12:11:10-12:11:4T1-2T4:T2T2T2T:2).</paragraph>
<paragraph id="FA1AF0281E7FC6BD752701B18EC8E2B1" pageId="0" pageNumber="165">
Sternites entirely smooth, weakly sclerotised; anterior operculum of males with 5 and posterior operculum of males with 17 (16-17) setae, of which 7 (6-7) setae located close to genital aperture (Fig.
<figureCitation id="DB37F7C05F6B1ECDEC15D4A3DE7BF06F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4C</figureCitation>
); sternite II of female with 6 setae, sternite III of female: (3)18(3). All setae simple; those on sternites IV-XI uniseriate; chaetotaxy: 5:(3)17(3):(3)12(3):16:16:16:16:16:6T1T6:T2T3T2T:2 (5-6:(3)16-18(3):(3)12-13(3):15-17:14-15:15:14-16:13-15:4-6T1T4-6:T3T2T3T:2).
</paragraph>
<paragraph id="E20EC0A03C9B0627F59771EE3AD9652A" pageId="0" pageNumber="165">
Internal genitalia with moderately long lateral and median genital sacs, median sac wrinkled, same length as lateral sacs; genital opening with 4+4 internal setae (Fig.
<figureCitation id="9DDA88A62A7C08F25A06BC0B674AE2CD" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4D</figureCitation>
).
</paragraph>
<paragraph id="BE3832431217DE6B761F7631BB64B8A6" pageId="0" pageNumber="165">
Pleural membrane granulated. Chelicera (Figs
<figureCitation id="64E2A54F7BAAE4A8861F90B674267BB2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Neobisium (Neobisium) adjaricum sp. nov., holotype male (A-D) and paratype male (E, F) A left chela, dorsal view B right chela, lateral view C right pedipalp (without chela), dorsal view D left chelicera, dorsal view E fixed finger of left chelicera, partial dorsal view F movable finger of left chelicera, partial dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, est - exterior sub-terminal, et - exterior terminal, ib - interior basal, isb - interior sub-basal, ist - interior sub-terminal, it - interior terminal; trichobothria on movable chelal finger: b - basal, sb - sub-basal, st - sub-terminal, t - terminal; gs - galeal seta; pc - coupled sensilla. Scale bars: 1.0 mm (A-C); 0.1 mm (D-F)." figureDoi="10.3897/zookeys.1100.81910.figure5" httpUri="https://binary.pensoft.net/fig/684539" pageId="0" pageNumber="165">5D-F</figureCitation>
,
<figureCitation id="94725DF01E69B34530616D561E55A690" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6F</figureCitation>
): hand with 7 acuminate setae; galea knob-like, with poorly developed hyaline convexity, sub-galeal seta situated distal to middle (T = 0.7); hand covered with faint reticulate pattern with pointed corners (Fig.
<figureCitation id="936CE95E603C590B2054E4444777D34E" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Neobisium (Neobisium) adjaricum sp. nov., holotype male (A-D) and paratype male (E, F) A left chela, dorsal view B right chela, lateral view C right pedipalp (without chela), dorsal view D left chelicera, dorsal view E fixed finger of left chelicera, partial dorsal view F movable finger of left chelicera, partial dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, est - exterior sub-terminal, et - exterior terminal, ib - interior basal, isb - interior sub-basal, ist - interior sub-terminal, it - interior terminal; trichobothria on movable chelal finger: b - basal, sb - sub-basal, st - sub-terminal, t - terminal; gs - galeal seta; pc - coupled sensilla. Scale bars: 1.0 mm (A-C); 0.1 mm (D-F)." figureDoi="10.3897/zookeys.1100.81910.figure5" httpUri="https://binary.pensoft.net/fig/684539" pageId="0" pageNumber="165">5D</figureCitation>
); small tooth (
<emphasis id="080C94591F7596D203EA5B032601835F" italics="true" pageId="0" pageNumber="165">tis</emphasis>
) under interior seta (
<emphasis id="E68B5BA897DCB01DA6D58E6E94B6D469" italics="true" pageId="0" pageNumber="165">is</emphasis>
) present, triangular (Fig.
<figureCitation id="08DA42E95385382D382CC9759053F2C5" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6F</figureCitation>
); fixed finger with 18 (17-19) teeth reaching finger base; movable finger with 7 (8-10) teeth reaching distal to middle of segment, large median tooth absent; serrula interior with 22 (22-25) blades, serrula exterior with 26 (25-28) blades; rallum with 8 blades, two distal ones denticulate, 6 posterior blades simple, smooth and acuminate, 2 or 3 proximal blades smallest.
</paragraph>
<caption id="9CD421AE69F7ECA6EE49009BC3F14CBB" doi="10.3897/zookeys.1100.81910.figure5" httpUri="https://binary.pensoft.net/fig/684539" pageId="0" pageNumber="165" start="Figure 5" startId="F5">
<paragraph id="6C079285DF4C98990054225B61F794DD" pageId="0" pageNumber="165">
<emphasis id="D222BD958C4B32DC6C999DB299E1077A" bold="true" pageId="0" pageNumber="165">Figure 5.</emphasis>
<taxonomicName id="9149F34E379DC4A9DFB56B6A484EA474" authorityName="Kolesnikov &amp; Christophoryová &amp; Przhiboro &amp; Turbanov" authorityYear="2022" class="Arachnida" family="Neobisiidae" genus="Neobisium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisium (Neobisium) adjaricum" order="Pseudoscorpiones" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="Neobisium">Neobisium (Neobisium) adjaricum</taxonomicName>
sp. nov., holotype male (
<emphasis id="D9DDB44A833A5F500D6EE2E2CA3B2117" bold="true" pageId="0" pageNumber="165">A-D</emphasis>
) and paratype male (
<emphasis id="5BB5BC0BA4750456D9B394A78B273D29" bold="true" pageId="0" pageNumber="165">E, F</emphasis>
)
<emphasis id="563A8D78D551DD3EA17B406C70CE0194" bold="true" pageId="0" pageNumber="165">A</emphasis>
left chela, dorsal view
<emphasis id="C5748E25ACBE5305CF9A2F51A69CEB54" bold="true" pageId="0" pageNumber="165">B</emphasis>
right chela, lateral view
<emphasis id="8B04BB201988008B076091ED06F9621D" bold="true" pageId="0" pageNumber="165">C</emphasis>
right pedipalp (without chela), dorsal view
<emphasis id="C862054358D44CB0D5162CFAD1CAFD2F" bold="true" pageId="0" pageNumber="165">D</emphasis>
left chelicera, dorsal view
<emphasis id="C383E0617134B7D30C910AF79ED04E76" bold="true" pageId="0" pageNumber="165">E</emphasis>
fixed finger of left chelicera, partial dorsal view
<emphasis id="0390198DB7A919246534180EDBD2769F" bold="true" pageId="0" pageNumber="165">F</emphasis>
movable finger of left chelicera, partial dorsal view. Abbreviations: trichobothria on fixed chelal finger:
<emphasis id="90B59CE183215691B3FC25CD79B6B706" italics="true" pageId="0" pageNumber="165">eb</emphasis>
- exterior basal,
<emphasis id="E740AF847A27F859DD6B348DE6F83096" italics="true" pageId="0" pageNumber="165">esb</emphasis>
- exterior sub-basal,
<emphasis id="3B8746D5D5E457783A76089219250105" italics="true" pageId="0" pageNumber="165">est</emphasis>
- exterior sub-terminal,
<emphasis id="214E91F38CB50C7AB4CA013B3242FA10" italics="true" pageId="0" pageNumber="165">et</emphasis>
- exterior terminal,
<emphasis id="86AE7F1595D6B2744FFDE0C8425B408D" italics="true" pageId="0" pageNumber="165">ib</emphasis>
- interior basal,
<emphasis id="37AE02605EF8081C2E8DB39FC0823B80" italics="true" pageId="0" pageNumber="165">isb</emphasis>
- interior sub-basal,
<emphasis id="CD0D282E1BCBF75562C94E079A0B5D93" italics="true" pageId="0" pageNumber="165">ist</emphasis>
- interior sub-terminal,
<emphasis id="FDFB88FFCEBB119804F3E7BF21DB7D91" italics="true" pageId="0" pageNumber="165">it</emphasis>
- interior terminal; trichobothria on movable chelal finger:
<emphasis id="C9AF8978C97A9E16193C34FEBCF6A4C2" italics="true" pageId="0" pageNumber="165">b</emphasis>
- basal,
<emphasis id="D321ED30EB126C82C977FCA4E4BB7458" italics="true" pageId="0" pageNumber="165">sb</emphasis>
- sub-basal,
<emphasis id="9659A73310973DCC579793BCFA7B853A" italics="true" pageId="0" pageNumber="165">st</emphasis>
- sub-terminal,
<emphasis id="D4137F7604B4BA4FF753D69E73446FE5" italics="true" pageId="0" pageNumber="165">t</emphasis>
- terminal;
<emphasis id="ADAD18A5B9AA9B366E8F303516227C32" italics="true" pageId="0" pageNumber="165">gs</emphasis>
- galeal seta;
<emphasis id="709C1E00C082E314F55C0052622B5335" italics="true" pageId="0" pageNumber="165">pc</emphasis>
- coupled sensilla. Scale bars: 1.0 mm (
<emphasis id="FDE2E540BF89BFF64FB23A0DB5106F81" bold="true" pageId="0" pageNumber="165">A-C</emphasis>
); 0.1 mm (
<emphasis id="0F3F9CA92146F77184336388760D728D" bold="true" pageId="0" pageNumber="165">D-F</emphasis>
).
</paragraph>
</caption>
<paragraph id="591231C854998B6AE49F38AF5C76F806" pageId="0" pageNumber="165">
Coxae (Fig.
<figureCitation id="AD2E7984BDA79C9E55C412BBE434FA39" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4B</figureCitation>
): pedipalpal coxa excluding manducatory process with 7 or 8 setae, manducatory process with 4 acuminate setae, seta at base of manducatory process longest; coxa I with moderately long triangular, sclerotised, apically pointed anterolateral process (0.06/0.04), and with denticulate mediolateral process; coxal chaetotaxy of legs: 6:6-8:7:10-11 (5-6:6-8:7:10-11). Each coxa of legs with one lyrifissure, pedipalpal coxa with one maxillary lyrifissure.
</paragraph>
<paragraph id="D014C67EE6B96E2AE611DE9825F25CFB" pageId="0" pageNumber="165">
Pedipalp (Figs
<figureCitation id="A75106EA974CD396CB1EDC7193D4BEA9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Neobisium (Neobisium) adjaricum sp. nov., holotype male (A-D) and paratype male (E, F) A left chela, dorsal view B right chela, lateral view C right pedipalp (without chela), dorsal view D left chelicera, dorsal view E fixed finger of left chelicera, partial dorsal view F movable finger of left chelicera, partial dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, est - exterior sub-terminal, et - exterior terminal, ib - interior basal, isb - interior sub-basal, ist - interior sub-terminal, it - interior terminal; trichobothria on movable chelal finger: b - basal, sb - sub-basal, st - sub-terminal, t - terminal; gs - galeal seta; pc - coupled sensilla. Scale bars: 1.0 mm (A-C); 0.1 mm (D-F)." figureDoi="10.3897/zookeys.1100.81910.figure5" httpUri="https://binary.pensoft.net/fig/684539" pageId="0" pageNumber="165">5A-C</figureCitation>
,
<figureCitation id="C4F6B75336DC4128CFC49AFB8EE84125" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6B, D, E, G, I</figureCitation>
): femur and patella smooth (in one paratype, outer side of femur finely granulate), hand of chela covered with fine reticulate ornament (Fig.
<figureCitation id="1968740A48392C7215A1A15764BC96F8" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6D</figureCitation>
), pedicel of chela with distinct ornament; trochanter with small dorsal tubercle, 2.0
<normalizedToken id="5DC8787A6D2342BE8A9AFF7E595EDCFE" originalValue="×">x</normalizedToken>
(1.60-1.85
<normalizedToken id="CFB0117FFAE3B29A13F4A83BBAD39402" originalValue="×">x</normalizedToken>
) as long as broad; femur with short pedicel, its margins of femur without tubercles, some setae mostly located in basal half of segment, without enlarged alveoli, one glandular pore located dorsodistally, 4.5
<normalizedToken id="44B232B16C3DEA4BAA9F5EBE8AF8AF0C" originalValue="×">x</normalizedToken>
(5-5.1
<normalizedToken id="85AE12540C97C22165A6AE0F21FDB8BD" originalValue="×">x</normalizedToken>
) as long as broad; patella with short and stout pedicel (0.20-0.22), patella distinctly shorter than femur, 1.43
<normalizedToken id="D2465CFF7C70872C0F4138C992548B9A" originalValue="×">x</normalizedToken>
(1.54-1.66
<normalizedToken id="5CA6FA559DF4D277D3BD6693F60ADF4C" originalValue="×">x</normalizedToken>
) as long as broad, with 3 lyrifissures basally and 2 lyrifissures distally, notch on median side reaching very close to middle of club length. Chela+ 3.8
<normalizedToken id="90E78011349CAC6EF59E2252DF55EC5E" originalValue="×">x</normalizedToken>
(3.68-3.87
<normalizedToken id="7F8A2E45DCE83DB69C04012897EBF9EF" originalValue="×">x</normalizedToken>
) and chela- 3.7
<normalizedToken id="77C8FC07D87B74F999BF5A88E627F2E7" originalValue="×">x</normalizedToken>
(3.52-3.75
<normalizedToken id="FBAD2FFEAA25B761BCA0D205420D729E" originalValue="×">x</normalizedToken>
) as long as broad; movable finger distinctly longer than hand+, 1.31
<normalizedToken id="BD373C79A4A5E7D3CDEA9398BFA31311" originalValue="×">x</normalizedToken>
(1.27-1.33
<normalizedToken id="40FD1B115E27CF1AEDF20B7F7750290A" originalValue="×">x</normalizedToken>
), but almost equal to femur in length, 0.95
<normalizedToken id="18032FF971D2358F5E5632FFE01B84DC" originalValue="×">x</normalizedToken>
(0.95-1.02
<normalizedToken id="5FD0D17217EBCCFF1F54A39C62FB4090" originalValue="×">x</normalizedToken>
); hand+ 1.62
<normalizedToken id="5CECEC1FE44E5FD1FE2E829E69A91EAE" originalValue="×">x</normalizedToken>
(1.63-1.77
<normalizedToken id="9143E11E6A5D98A5D3A3FEE94E937F00" originalValue="×">x</normalizedToken>
) and hand- 1.52
<normalizedToken id="A532A7199C991C01BAE68B8EDEE478EE" originalValue="×">x</normalizedToken>
(1.57-1.62
<normalizedToken id="D72B6BCF7D789BB9FAA0399092BA3D57" originalValue="×">x</normalizedToken>
) as long as broad; retrolateral surface of hand with 3 glandular pores located around trichobothria
<emphasis id="C201B717D9666979CE24029207C442EC" italics="true" pageId="0" pageNumber="165">eb</emphasis>
and
<emphasis id="803A5CFF7B5DB6DB3877A61CC074A2D6" italics="true" pageId="0" pageNumber="165">esb</emphasis>
(two pores close to each other, one distant); fixed finger with 3 lyrifissures:
<emphasis id="6258DEB1B42EB9C0C2DE68C7283E1057" italics="true" pageId="0" pageNumber="165">fa</emphasis>
located close to base in retrolateral view,
<emphasis id="8BDBCB69E55440A8A76E5A1AFADA1CD4" italics="true" pageId="0" pageNumber="165">fb</emphasis>
located slightly proximal to
<emphasis id="D923F1604F3DC238BD334216AD87AFE9" italics="true" pageId="0" pageNumber="165">ib</emphasis>
,
<emphasis id="4F07D08F325CB3A380174BB447299A15" italics="true" pageId="0" pageNumber="165">fd</emphasis>
located distal to
<emphasis id="E45866F30D3F8067833128B3A772FF67" italics="true" pageId="0" pageNumber="165">it</emphasis>
; movable finger with 3 lyrifissures in retrolateral view:
<emphasis id="388E21F99E82A3629A474387467DD864" italics="true" pageId="0" pageNumber="165">ma</emphasis>
2 located closer to trichobothrium
<emphasis id="380FC513A0A49F7808B39C7B798AA97B" italics="true" pageId="0" pageNumber="165">b</emphasis>
than to
<emphasis id="83D8619B2CFF5C5CECE4BF5774C07BAC" italics="true" pageId="0" pageNumber="165">sb</emphasis>
,
<emphasis id="AF711085527CBA31F13CD72195211344" italics="true" pageId="0" pageNumber="165">ma</emphasis>
1 nearly at same level with
<emphasis id="0D7AFA7EB3C7050C040445C0F9185BD4" italics="true" pageId="0" pageNumber="165">sb</emphasis>
(slightly proximal to
<emphasis id="D0355F7BF0024F9279A44DA4115102C0" italics="true" pageId="0" pageNumber="165">sb</emphasis>
),
<emphasis id="B8DEBAEA3056A243856B4BAF9EBC4405" italics="true" pageId="0" pageNumber="165">ma</emphasis>
3 closer to
<emphasis id="6DB5D5E173DB5B186696B80885F1FF24" italics="true" pageId="0" pageNumber="165">st</emphasis>
than to
<emphasis id="D404951D1AA50F9E0050A499FB4A4448" italics="true" pageId="0" pageNumber="165">sb</emphasis>
,
<emphasis id="789694ADC00BC1A2D4E9B851CE7FAC0E" italics="true" pageId="0" pageNumber="165">ma</emphasis>
1 slightly closer to
<emphasis id="26522F7DAB5B3D1D857EB67D0784F694" italics="true" pageId="0" pageNumber="165">ma</emphasis>
3 than to
<emphasis id="4803653EFDA003FC08C579C599FDA0A6" italics="true" pageId="0" pageNumber="165">ma</emphasis>
2; one or two sensilla
<emphasis id="97782D524A2E42E0C4C6EAA6AF8C583A" italics="true" pageId="0" pageNumber="165">pc</emphasis>
located between trichobothria
<emphasis id="E97F29B2770AE80C98A3E172ECC27B96" italics="true" pageId="0" pageNumber="165">sb</emphasis>
and
<emphasis id="9B6F590399781C04ACC814311F8A18B6" italics="true" pageId="0" pageNumber="165">st</emphasis>
, close to dental canals (one sensillum closer to
<emphasis id="44455B46022E09A5911C28F2F93F34F3" italics="true" pageId="0" pageNumber="165">sb</emphasis>
, other closer to
<emphasis id="D08118DAD40619ABCA0BC556D1D47EE4" italics="true" pageId="0" pageNumber="165">st</emphasis>
; latter not present in all specimens); fixed finger with 48 (48-51) contiguous triangular teeth reaching level of trichobothrium
<emphasis id="1D441FBD4F2A83FD6E8CFC7AF4BBEF01" italics="true" pageId="0" pageNumber="165">ib</emphasis>
, two outer distal teeth smallest, other teeth almost equal in size, all teeth with dental canal; movable finger with 45 (44-47) contiguous teeth not reaching level of trichobothrium
<emphasis id="7C6ED74772115F22E26DCC79A3774596" italics="true" pageId="0" pageNumber="165">b</emphasis>
, teeth slightly reduced in size from tip of base of, distal 10-12 teeth triangular, other blunt and apically rounded, teeth with dental canal (except basal 6 or 7 small teeth); nodus ramosus of venom duct in fixed chelal finger short, situated very close to finger tip.
</paragraph>
<caption id="D5A247AC4A75F82C89E551FC74471097" doi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165" start="Figure 6" startId="F6">
<paragraph id="48A7BEC4B0CD24C6098F752ECCFBC6E1" pageId="0" pageNumber="165">
<emphasis id="95F9805F33C1D9EDD34C88B0E3E9661B" bold="true" pageId="0" pageNumber="165">Figure 6.</emphasis>
<taxonomicName id="457EB0A6A5115A74AD9E4FD317535B0A" authorityName="Kolesnikov &amp; Christophoryová &amp; Przhiboro &amp; Turbanov" authorityYear="2022" class="Arachnida" family="Neobisiidae" genus="Neobisium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisium (Neobisium) adjaricum" order="Pseudoscorpiones" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="Neobisium">Neobisium (Neobisium) adjaricum</taxonomicName>
sp. nov., holotype male, light microscope images
<emphasis id="12BFD380246F218E0338FE211EB2BF7A" bold="true" pageId="0" pageNumber="165">A</emphasis>
distal part of left pedal tarsus IV, dorsal view
<emphasis id="FA2EA92B9D8C30D9A5F8A685C0D03723" bold="true" pageId="0" pageNumber="165">B</emphasis>
basal part of pedipalpal patella, dorsal view
<emphasis id="795FD8C5C8880023ADC30B69E80D5D4F" bold="true" pageId="0" pageNumber="165">C</emphasis>
epistome, dorsal view
<emphasis id="EAA4219DF039F035B9D113CDD0AFF7D9" bold="true" pageId="0" pageNumber="165">D</emphasis>
surface of chelal hand, dorsal view
<emphasis id="AD3D38A67ABE1C5BE85CBE3631DA40C1" bold="true" pageId="0" pageNumber="165">E</emphasis>
basal part of right chelal fixed finger, lateral view
<emphasis id="F807379B8DD1F4D87ABBF37DD4942CB4" bold="true" pageId="0" pageNumber="165">F</emphasis>
part of left chelicera (part), dorsal view
<emphasis id="D490EB904432DEE3DFEF8600047AB000" bold="true" pageId="0" pageNumber="165">G</emphasis>
part of chelal fingers, lateral view
<emphasis id="6B0DDAE6D233E0881BCB8248CED3EA57" bold="true" pageId="0" pageNumber="165">H</emphasis>
surface of carapace, dorsal view
<emphasis id="9CE726D3EEB224E26A08C6702FF20C0D" bold="true" pageId="0" pageNumber="165">I</emphasis>
pedicel of chela, lateral view
<emphasis id="28DC00F475017369CDE86DE4A1FCB9A5" bold="true" pageId="0" pageNumber="165">J</emphasis>
right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger:
<emphasis id="38F7498D7B928B6AF834FF4FFB97ACD8" italics="true" pageId="0" pageNumber="165">eb</emphasis>
- exterior basal,
<emphasis id="47C30F810A6BB2D88EBD72BB30195476" italics="true" pageId="0" pageNumber="165">esb</emphasis>
- exterior sub-basal,
<emphasis id="BF8B171833ADAE0369B516424E263EC1" italics="true" pageId="0" pageNumber="165">isb</emphasis>
- interior sub-basal; trichobothria on movable chelal finger:
<emphasis id="9392411C8FB4C3F2CC740FE30CEA2C27" italics="true" pageId="0" pageNumber="165">st</emphasis>
- sub-terminal;
<emphasis id="5880A7F605A36962F88FBB3E5356C27C" italics="true" pageId="0" pageNumber="165">dat</emphasis>
- dorsal accessory tooth;
<emphasis id="0E818CB95860AAB4BB4F6A5D6809FB12" italics="true" pageId="0" pageNumber="165">fa</emphasis>
,
<emphasis id="E4D2BA502747733EBEE920CCE3D7905B" italics="true" pageId="0" pageNumber="165">fb</emphasis>
- lyrifissures;
<emphasis id="BBABD94AF1235ECD7A11B8961409E1E8" italics="true" pageId="0" pageNumber="165">gs</emphasis>
,
<emphasis id="0ECC4AE62F3ED85191BC810734CFFFF9" italics="true" pageId="0" pageNumber="165">ls</emphasis>
,
<emphasis id="4020865CCD79E7036237CF28CF87D1F0" italics="true" pageId="0" pageNumber="165">is</emphasis>
,
<emphasis id="EF369F941BD931722DDB286893C09550" italics="true" pageId="0" pageNumber="165">sbs</emphasis>
- cheliceral setae;
<emphasis id="278492B5D4F56C4D210DA7E6352F452D" italics="true" pageId="0" pageNumber="165">pc</emphasis>
- coupled sensilla;
<emphasis id="B4AD2C4EFC1FA0DFF44FFF4B33ECD428" italics="true" pageId="0" pageNumber="165">tis</emphasis>
- tooth close to seta
<emphasis id="D94DE9F7254B75857691FC27F7B88560" italics="true" pageId="0" pageNumber="165">is</emphasis>
on chelicera; arrows show glandular pores. Scale bar: 0.1 mm.
</paragraph>
</caption>
<paragraph id="F9FEFBAA3008E8A1B305CB4152251AF4" pageId="0" pageNumber="165">
Trichobothriotaxy: fixed finger with 8 trichobothria, movable finger with 4 trichobothria; fixed finger with trichobothria
<emphasis id="8B8BA2479E04BCC2B72B4B564ECE02F9" italics="true" pageId="0" pageNumber="165">esb</emphasis>
and
<emphasis id="B89E83E674F52B1836D1A1223413260E" italics="true" pageId="0" pageNumber="165">eb</emphasis>
located close to each other and slightly proximal to
<emphasis id="C04932FD0AEA4F576D3CE6C7B9F37CE1" italics="true" pageId="0" pageNumber="165">ib</emphasis>
, with
<emphasis id="04371179334E26AEA3C340E1C7F61C58" italics="true" pageId="0" pageNumber="165">isb</emphasis>
on retrolateral surface, with
<emphasis id="1A7EED4E022AE975B77DDBD37E58FA2F" italics="true" pageId="0" pageNumber="165">ib</emphasis>
closer to
<emphasis id="FC3F3020C266FDA9D0D6542CE9CE4640" italics="true" pageId="0" pageNumber="165">isb</emphasis>
than to
<emphasis id="979B4003CB29BE714133D52E84A2D6C0" italics="true" pageId="0" pageNumber="165">esb</emphasis>
(in lateral and dorsal views), trichobothrium
<emphasis id="A0B1BB7A9783DB2523EAC8C69302D02B" italics="true" pageId="0" pageNumber="165">ist</emphasis>
distinctly closer to
<emphasis id="7AA18F30E30A1EAEF13C4C8D7BAC63B0" italics="true" pageId="0" pageNumber="165">est</emphasis>
than to
<emphasis id="76CD80199520062CAB1BBFA879763943" italics="true" pageId="0" pageNumber="165">isb</emphasis>
, distance
<emphasis id="75D2A8EDF3EDAC2CACE6A5886534DCBC" italics="true" pageId="0" pageNumber="165">ist</emphasis>
-
<emphasis id="2AF59629EF6733F97064C9EFE3184FA3" italics="true" pageId="0" pageNumber="165">est</emphasis>
similar to distance
<emphasis id="DFF8201A53DC0EB7CBA40AC3F183B382" italics="true" pageId="0" pageNumber="165">est</emphasis>
-
<emphasis id="9C9BDE8B5ADA2A5D449C6C80B2DDDAF1" italics="true" pageId="0" pageNumber="165">it</emphasis>
, but less than twice as long as distance
<emphasis id="0239161F607358C21893265C4F0371A9" italics="true" pageId="0" pageNumber="165">est</emphasis>
-
<emphasis id="01424B7B7A2F03467EC62377F4375655" italics="true" pageId="0" pageNumber="165">et</emphasis>
in lateral view, trichobothrium
<emphasis id="CB853CB54C348DDB7ECF0E9852D1B2EE" italics="true" pageId="0" pageNumber="165">ist</emphasis>
situated distal to middle of finger (T = 0.57-0.60),
<emphasis id="0B92DDBEC067A2696CB25E5D69FA5AD7" italics="true" pageId="0" pageNumber="165">ist</emphasis>
located distinctly proximal to
<emphasis id="769518B23299FBD0BDA4FFDC1F3CF80F" italics="true" pageId="0" pageNumber="165">t</emphasis>
,
<emphasis id="43857F4B3C0D2E1571F22E1CE5B89B1F" italics="true" pageId="0" pageNumber="165">et</emphasis>
located posterior to
<emphasis id="EF7ECED787CF576AC18CBCE3AD3A53A4" italics="true" pageId="0" pageNumber="165">it</emphasis>
, at approximately the same distance between
<emphasis id="FCA3247E00B99C2612299DBB6C90919B" italics="true" pageId="0" pageNumber="165">est</emphasis>
and
<emphasis id="05DC03A6392CCDD97F06AE3ADDD131CF" italics="true" pageId="0" pageNumber="165">it</emphasis>
(in lateral and dorsal views),
<emphasis id="9754DCCE23B8A41C63A8EEDB6E744207" italics="true" pageId="0" pageNumber="165">ist</emphasis>
situated distal to middle of finger; movable finger with trichobothrium
<emphasis id="B889973513597BBB6C3C1A02F874BBDD" italics="true" pageId="0" pageNumber="165">st</emphasis>
situated slightly closer to
<emphasis id="FA46BC0880F1A75AA75F32F994DAAF5B" italics="true" pageId="0" pageNumber="165">t</emphasis>
than to
<emphasis id="27F04610432CFAC8F6820CF5BB5898C2" italics="true" pageId="0" pageNumber="165">sb</emphasis>
, trichobothrium
<emphasis id="70E7C508BCFA60820327FA7AEDEFE71C" italics="true" pageId="0" pageNumber="165">sb</emphasis>
slightly closer to
<emphasis id="FBF37813063BF72A90A8CDAD62B40B7D" italics="true" pageId="0" pageNumber="165">b</emphasis>
than to
<emphasis id="5BF43EFF39646AFC9B59126C7931C311" italics="true" pageId="0" pageNumber="165">st</emphasis>
(at almost equal distances), distance
<emphasis id="4A5BC651E01DFD6F371BF3314BFFB05E" italics="true" pageId="0" pageNumber="165">b</emphasis>
-
<emphasis id="61161E38F037F39E88C18A1F5A86C25B" italics="true" pageId="0" pageNumber="165">sb</emphasis>
almost equal to distance
<emphasis id="4AEE00F9E965F22386E077C61A195A34" italics="true" pageId="0" pageNumber="165">st</emphasis>
-
<emphasis id="069E695F6BA6237767DF797038BF9AF3" italics="true" pageId="0" pageNumber="165">t</emphasis>
.
</paragraph>
<paragraph id="6607BE440D4CE40EC46361047F6FD057" pageId="0" pageNumber="165">
Legs (Figs
<figureCitation id="E84CC4504EF715BD7E7B40E28C95648C" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4E, F</figureCitation>
,
<figureCitation id="6ACCBB644B132A1A6F13D62E13126EA0" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6A</figureCitation>
): all claws of legs with small dorsal accessory tooth (Figs
<figureCitation id="F22B896A259769E8B4963C7AE51D7DD7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4E</figureCitation>
,
<figureCitation id="F9F698755018FDD23EDABDCCD5BBE83F" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Neobisium (Neobisium) adjaricum sp. nov., holotype male, light microscope images A distal part of left pedal tarsus IV, dorsal view B basal part of pedipalpal patella, dorsal view C epistome, dorsal view D surface of chelal hand, dorsal view E basal part of right chelal fixed finger, lateral view F part of left chelicera (part), dorsal view G part of chelal fingers, lateral view H surface of carapace, dorsal view I pedicel of chela, lateral view J right eyes, dorsal view. Abbreviations: trichobothria on fixed chelal finger: eb - exterior basal, esb - exterior sub-basal, isb - interior sub-basal; trichobothria on movable chelal finger: st - sub-terminal; dat - dorsal accessory tooth; fa, fb - lyrifissures; gs, ls, is, sbs - cheliceral setae; pc - coupled sensilla; tis - tooth close to seta is on chelicera; arrows show glandular pores. Scale bar: 0.1 mm." figureDoi="10.3897/zookeys.1100.81910.figure6" httpUri="https://binary.pensoft.net/fig/684540" pageId="0" pageNumber="165">6A</figureCitation>
), arolia simple and shorter than claws. Leg I femur 4.00
<normalizedToken id="6B316FB51E48022132E693BD3B127C0D" originalValue="×">x</normalizedToken>
(4.44-5.50
<normalizedToken id="C0A299AEF81AD14A242727B3A7F22B51" originalValue="×">x</normalizedToken>
) and patella 3.30
<normalizedToken id="4A114D00621058C40C8A774BB4CFB9F0" originalValue="×">x</normalizedToken>
(3.30-3.40
<normalizedToken id="A01F600A478B72B12A3566143C5B66E9" originalValue="×">x</normalizedToken>
) as long as deep, femur 1.21
<normalizedToken id="4D90AE5748080F276A7356ABB01B33B9" originalValue="×">x</normalizedToken>
(1.48-1.66
<normalizedToken id="C026A51419BBF31FAEC413F1736AA8B1" originalValue="×">x</normalizedToken>
) as long as patella, tibia 4.12
<normalizedToken id="FED8550B5F22B6775A30528EA4EF6CFA" originalValue="×">x</normalizedToken>
(4.28-4.37
<normalizedToken id="638E09A11F333ACD0A820F32B487861E" originalValue="×">x</normalizedToken>
), metatarsus 3.00
<normalizedToken id="9B6E207FB6D28B2E7A384E88FF36ED4C" originalValue="×">x</normalizedToken>
(3.12-3.14
<normalizedToken id="9E57E1E669B752D8894FCBD1ADA90AA6" originalValue="×">x</normalizedToken>
) and tarsus 4.28
<normalizedToken id="233365CEAFEEDFA15437F3863D449146" originalValue="×">x</normalizedToken>
(4.0-4.28
<normalizedToken id="5417264BFC9FF2B411981F926EE528D3" originalValue="×">x</normalizedToken>
) as long as deep, tarsus 1.25
<normalizedToken id="2A7CBBF501C178AA28B36AF7D1BB4BD9" originalValue="×">x</normalizedToken>
(1.20-1.27
<normalizedToken id="E8F8652C63E5A04F93859AEEC3EEC42D" originalValue="×">x</normalizedToken>
) as long as metatarsus; leg IV femur 2.00
<normalizedToken id="9D3C6582684EAD7829028BCC8DE83915" originalValue="×">x</normalizedToken>
(1.76-2.00
<normalizedToken id="724EFEAF66E453D0DCFA48533089550D" originalValue="×">x</normalizedToken>
), patella 2.22
<normalizedToken id="E03DC0C3B0D05B805C95722B99902349" originalValue="×">x</normalizedToken>
(2.05-2.63
<normalizedToken id="73BD33932BCBD001AF6AEA9E34F96758" originalValue="×">x</normalizedToken>
), femur+patella 4.22
<normalizedToken id="FD77677E1963FF9CAE98BAF23D35DB90" originalValue="×">x</normalizedToken>
(3.94-4.50
<normalizedToken id="75F5D00F60C92054348013DCBF5B7517" originalValue="×">x</normalizedToken>
), tibia 6.0
<normalizedToken id="C874E72738EA8540E0E2DBF903CF06BD" originalValue="×">x</normalizedToken>
(5.80-6.36
<normalizedToken id="04057BA33FE5ED952E9CAD4BEEF6DDF6" originalValue="×">x</normalizedToken>
), metatarsus 3.75
<normalizedToken id="27CA3AA3E4219CDF613E5217F8F33FAA" originalValue="×">x</normalizedToken>
(3.75-4.12
<normalizedToken id="DC39F70082919A398228C9AB8E82FCA9" originalValue="×">x</normalizedToken>
) and tarsus 5.71
<normalizedToken id="F0A86E84C4ECAA6D3E049DEE0876AFF2" originalValue="×">x</normalizedToken>
(5.42-5.75
<normalizedToken id="6198D0EA99ED5759F4B4DABFEA629B30" originalValue="×">x</normalizedToken>
) as long as deep, tarsus 1.33
<normalizedToken id="017918D298D49D06B70B4879C47E8003" originalValue="×">x</normalizedToken>
(1.26-1.39
<normalizedToken id="9AE95B1BC0C3A3453EF8A32AE7CE4706" originalValue="×">x</normalizedToken>
) as long as metatarsus; tibia IV with long tactile seta situated slightly proximal to middle (T = 0.40), metatarsus IV with long tactile seta situated basally (T = 0.125), tarsus with tactile seta situated proximal to middle (T = 0.38); sub-terminal setae branched, basal ramus short and smooth (Fig.
<figureCitation id="4E871A626D72C2E1DDA3225B66A38D2B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Neobisium (Neobisium) adjaricum sp. nov., holotype male A carapace, dorsal view B coxae of pedipalp and legs I-IV, ventral view C sternites II-IV, ventral view D male genitalia, ventral view E right leg IV (without trochanter), dorsal view F sub-terminal seta on pedal tarsus IV. Abbreviation: dat - dorsal accessory tooth. Scale bars: 1.0 mm (A-C, E); 0.1 mm (D, F)." figureDoi="10.3897/zookeys.1100.81910.figure4" httpUri="https://binary.pensoft.net/fig/684538" pageId="0" pageNumber="165">4F</figureCitation>
).
</paragraph>
<paragraph id="7231B877FF9D0392A8663B914BCDAE72" pageId="0" pageNumber="165">Sexual dimorphism not pronounced. Measurements: body length 2.95 (2.87-2.90); carapace 0.65/0.82 (0.64-0.65/0.72-0.77); chelicera 0.45/0.24 (0.40-0.50/0.22-0.26), movable finger of chelicera 0.28 (0.26-0.30); pedipalp: trochanter 0.40/0.20 (0.37-0.40/0.20-0.25), femur 0.90/0.20 (0.77-1.00/0.15-0.20), patella 0.63/0.24 (0.50-0.60/0.20-0.23), chela+ 1.52 (1.40-1.55), chela- 1.48 (1.34-1.50), hand+ 0.65/0.40 (0.62-0.71/0.38-0.40), hand- 0.61 (0.60-0.65), movable finger 0.86 (0.79-0.95); leg I: trochanter 0.18/0.15 (0.18-0.22/0.15), femur 0.40/0.10 (0.40-0.55/0.09-0.10), patella 0.33/0.10 (0.27-0.33/0.08-0.10), tibia 0.33/0.08 (0.30-0.35/0.07-0.08), metatarsus 0.24/0.08 (0.22-0.25/0.07-0.08), tarsus 0.30/0.07 (0.28-0.30/0.07); leg IV: trochanter 0.40/0.15 (0.40/0.18), femur 0.36/0.18 (0.30-0.40/0.17-0.20), patella 0.40/0.18 (0.37-0.50/0.18-0.19), femur+patella 0.76 (0.67-0.90), tibia 0.60/0.10 (0.58-0.70/0.10-0.11), metatarsus 0.30/0.08 (0.30-0.33/0.08), tarsus 0.40/0.07 (0.38-0.46/0.07-0.08).</paragraph>
</subSubSection>
<subSubSection id="F3EE519F116BC4660F1C862B09083D94" pageId="0" pageNumber="165" type="comparisons">
<paragraph id="EB024E4BBD4371544544A77C5145BC2B" pageId="0" pageNumber="165">Comparison.</paragraph>
<paragraph id="004F3D16C91A5F84016A3A375A899944" pageId="0" pageNumber="165">
The new species is most similar to
<taxonomicName id="0B56DFAF8F3DC521BCD3D7400C9FC78B" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) vilcekii" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="vilcekii" subGenus="N.">N. (N.) vilcekii</taxonomicName>
and
<taxonomicName id="5B3E7E9250DC7B6B41FE4DFEA56C9222" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) speleophilum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="speleophilum" subGenus="N.">N. (N.) speleophilum</taxonomicName>
to the following characters: tarsal claw IV with dorsal accessory tooth, palpal femur without granulate, trichobothrium
<emphasis id="0DF39964B1D9062E2F1623D22813ABE5" italics="true" pageId="0" pageNumber="165">ist</emphasis>
located distinctly proximal to
<emphasis id="F53048864BEF3CE6D93F4A7BFE273F62" italics="true" pageId="0" pageNumber="165">t.</emphasis>
The new species differs from
<taxonomicName id="3AF38909BE0FDCA10257EE34E23CF1B4" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) vilcekii" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="vilcekii" subGenus="N.">N. (N.) vilcekii</taxonomicName>
in the following characters: shorter pedipalpal femur (0.77-1.00 vs. 1.35-1.72 in
<taxonomicName id="33E5ADE8AEAF99B2B0F88B2EA404BAC3" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) vilcekii" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="vilcekii" subGenus="N.">N. (N.) vilcekii</taxonomicName>
), shorter hand+ (0.62-0.71 vs. 1.07-1.45) and movable finger (0.79-0.95 vs. 1.32-1.57), higher ratio of length of movable finger to hand+ (1.27-1.33
<normalizedToken id="88998D534BB740BD27E9685E170CBFA3" originalValue="×">x</normalizedToken>
vs. 1.06-1.23
<normalizedToken id="4C5BE834BC2BE2C8CE9A7D7881F8462C" originalValue="×">x</normalizedToken>
), lower ratio of length to width of chelal hand+ (1.62-1.77
<normalizedToken id="A41EF75A63AFDB88488A6105A3F4A7C9" originalValue="×">x</normalizedToken>
vs. 1.81-2.15
<normalizedToken id="F8B29BAA5FF60607D4D2D18012202E20" originalValue="×">x</normalizedToken>
), longer epistome (moderately long vs. short), smaller number of setae on sternite II (5 vs. 10-15), longer anterolateral process on coxa I (0.06/0.04 vs. 0.04/0.05), smaller number of setae on pedipalpal and leg coxae (see Table
<tableCitation id="2E0B137065125ADA38C4FBF6FAC4E7FB" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Numbers of setae on the coxae of some species of the subgenus Neobisium Neobisium from the Caucasus and adjacent territories." httpUri="http://table.plazi.org/id/3E8801DBE711556FFE6C11EDDF8B06CD" pageId="0" pageNumber="165" tableUuid="3E8801DBE711556FFE6C11EDDF8B06CD">1</tableCitation>
), smaller number of setae on manducatory process of pedipalp (4 vs. 5), and the presence/absence of large median tooth on movable finger of chelicera (absent vs. present) (
<bibRefCitation id="981D779836E83E0C97A45B93384C0DBB" author="Krumpal, M" journalOrPublisher="Biologia" pageId="0" pageNumber="165" pagination="607 - 612" refId="B19" refString="Krumpal, M, 1983. Neobisium (N.) vilcekii sp. n., ein neuer Pseudoscorpion aus der UdSSR (Neobisiidae, Pseudoscorpiones). Ueber Pseudoscorpioniden-Fauna der UdSSR IV. Biologia 38 (6): 607 - 612" title="Neobisium (N.) vilcekii sp. n., ein neuer Pseudoscorpion aus der UdSSR (Neobisiidae, Pseudoscorpiones). Ueber Pseudoscorpioniden-Fauna der UdSSR IV." volume="38" year="1983">
<normalizedToken id="C7B9C21773312659011CD475E05056E8" originalValue="Krumpál">Krumpal</normalizedToken>
1983
</bibRefCitation>
;
<bibRefCitation id="25416BC2F75549EB9B5DA762324584B4" DOI="https://doi.org/10.2476/asjaa.70.77" author="Nassirkhani, M" journalOrPublisher="Acta Arachnologica" pageId="0" pageNumber="165" pagination="77 - 82" refId="B25" refString="Nassirkhani, M, Snegovaya, N, 2021. A redescription of Neobisium (N.) vilcekii Krumpal 1983 (Pseudoscorpiones: Neobisiidae) from Caucasian Russia. Acta Arachnologica 70 (2): 77 - 82, DOI: https://doi.org/10.2476/asjaa.70.77" title="A redescription of Neobisium (N.) vilcekii Krumpal 1983 (Pseudoscorpiones: Neobisiidae) from Caucasian Russia." url="https://doi.org/10.2476/asjaa.70.77" volume="70" year="2021">Nassirkhani and Snegovaya 2021</bibRefCitation>
). The new species differs from
<taxonomicName id="4D5CE7809C493D9E99A60C6A8AADE1DC" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) speleophilum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="speleophilum" subGenus="N.">N. (N.) speleophilum</taxonomicName>
in the following characters: shorter chela+ (1.40-1.55vs. 1.72-1.80 in
<taxonomicName id="85DBF8B66CA4A3FA325A9F58AB96EDCE" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) speleophilum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="speleophilum" subGenus="N.">N. (N.) speleophilum</taxonomicName>
), hand+ (0.62-0.71 vs. 0.73-0.80), and movable finger (0.79-0.95 vs. 1.12-1.20), lower ratio movable finger/hand+ (1.27-1.33
<normalizedToken id="D71254BA2BBA6A097EE1D0FB079C5DB1" originalValue="×">x</normalizedToken>
vs. 1.45-1.53
<normalizedToken id="95A3F86E8654F3524E4D00ED245E6897" originalValue="×">x</normalizedToken>
), sternite II with 5 setae vs. 11 or 12, longer (as long as lateral sacs) median sac of male genitalia vs. short (lateral sac longer than median) (
<bibRefCitation id="7DCED1D282F95AB8FAAF668423F84969" author="Krumpal, M" journalOrPublisher="Biologia" pageId="0" pageNumber="165" pagination="163 - 172" refId="B20" refString="Krumpal, M, 1986. Pseudoscorpione (Arachnida) aus Hoehlen der UdSSR. Ueber Pseudoscorpioniden-Fauna der UdSSR V. Biologia 41 (2): 163 - 172" title="Pseudoscorpione (Arachnida) aus Hoehlen der UdSSR. Ueber Pseudoscorpioniden-Fauna der UdSSR V." volume="41" year="1986">
<normalizedToken id="0A597D61959C67E07554D464F4D7DABB" originalValue="Krumpál">Krumpal</normalizedToken>
1986
</bibRefCitation>
;
<bibRefCitation id="3FC914F5D664F125CAFE8A1ED6504DFC" DOI="https://doi.org/10.13156/arac.2020.18.5.449" author="Nassirkhani, M" journalOrPublisher="Arachnology" pageId="0" pageNumber="165" pagination="449 - 461" refId="B28" refString="Nassirkhani, M, Zaragoza, JA, Snegovaya, N, Chumachenko, YA, 2020. Description of two new Neobisium (Neobisium) species and redescription of Neobisium (N.) speleophilum from Caucasian Russia, with a key to the Neobisium (Neobisium) species (Arachnida: Pseudoscorpiones) recorded from Russia. Arachnology 18 (5): 449 - 461, DOI: https://doi.org/10.13156/arac.2020.18.5.449" title="Description of two new Neobisium (Neobisium) species and redescription of Neobisium (N.) speleophilum from Caucasian Russia, with a key to the Neobisium (Neobisium) species (Arachnida: Pseudoscorpiones) recorded from Russia." url="https://doi.org/10.13156/arac.2020.18.5.449" volume="18" year="2020">Nassirkhani et al. 2020</bibRefCitation>
).
</paragraph>
<caption id="3E8801DBE711556FFE6C11EDDF8B06CD" ID-Table-UUID="3E8801DBE711556FFE6C11EDDF8B06CD" httpUri="http://table.plazi.org/id/3E8801DBE711556FFE6C11EDDF8B06CD" pageId="0" pageNumber="165" start="Table 1" startId="T1">
<paragraph id="52A805AC0FDAC24705EC62A61B18526E" pageId="0" pageNumber="165">
<emphasis id="63D897F5BB31A5367B867F419D88C1CA" bold="true" pageId="0" pageNumber="165">Table 1.</emphasis>
Numbers of setae on the coxae of some species of the subgenus
<taxonomicName id="AE509B16E18714EE385CB28E79AB3972" authority="Neobisium" class="Arachnida" family="Neobisiidae" genus="Neobisium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neobisium (Neobisium)" order="Pseudoscorpiones" pageId="0" pageNumber="165" phylum="Arthropoda" rank="subGenus" subGenus="Neobisium">Neobisium Neobisium</taxonomicName>
from the Caucasus and adjacent territories.
</paragraph>
</caption>
<paragraph id="FFCCDEEA38EC855D57B71AC2B8C085DF" pageId="0" pageNumber="165">
<table id="25DAAB08E6697832B6A473961DCD3FCB" pageId="0" pageNumber="165">
<tr id="10A8C7511AC14727A78539BA29E61577" pageId="0" pageNumber="165">
<th id="742BA621868CEEE16B30182AD2850BD0" colspan="1" pageId="0" pageNumber="165" rowspan="1">Species</th>
<th id="EF5348B649EC1EDAA0CDB4D962F09672" colspan="1" pageId="0" pageNumber="165" rowspan="1">Pedipalpal coxa (including manducatory process)</th>
<th id="51F22B688F5D4DD29F3260385D4B94D8" colspan="1" pageId="0" pageNumber="165" rowspan="1">Manducatory process of pedipalpal coxae</th>
<th id="A7A5944955706FFB3DE95831EE117A70" colspan="1" pageId="0" pageNumber="165" rowspan="1">Coxa I</th>
<th id="4499B69D0141C3877F100FA1E6D74F5F" colspan="1" pageId="0" pageNumber="165" rowspan="1">Coxa II</th>
<th id="FC33BF428DC9F058064A7A675899FCEB" colspan="1" pageId="0" pageNumber="165" rowspan="1">Coxa III</th>
<th id="A6428953AC4BDA69D3199E4CDC967B33" colspan="1" pageId="0" pageNumber="165" rowspan="1">Coxa IV</th>
</tr>
<tr id="34C1AE5BA97B34AE6634A155CDC6C80F" pageId="0" pageNumber="165">
<td id="215CE1E31210186512BC4E621072E946" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="E7390C39E0D9A323BD669114887F9E76" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) adjaricum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="N.">N. (N.) adjaricum</taxonomicName>
sp. nov.
</td>
<td id="DA23CBC29C619142AA9DCBC6BE776CB1" colspan="1" pageId="0" pageNumber="165" rowspan="1">11-12</td>
<td id="9604E1062E9649826B911C1A3D6ABCD2" colspan="1" pageId="0" pageNumber="165" rowspan="1">4</td>
<td id="69F43FF379EDE7577ADFBF4FE8A6A078" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-6</td>
<td id="C7F19A36C673363C9020DF92F1450F43" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-8</td>
<td id="D86A25D52E5AF5C8BCF851CA0D5E7337" colspan="1" pageId="0" pageNumber="165" rowspan="1">7</td>
<td id="1DF9481A9192B10E8EB6FCD4C11D385E" colspan="1" pageId="0" pageNumber="165" rowspan="1">10-11</td>
</tr>
<tr id="8AEBD0F2DE16731357A66892300573A5" pageId="0" pageNumber="165">
<td id="BEB8796DAE9997473FFC60888ABA8E29" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="EEFB3C66F0B913561F647E259103A449" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) anatolicum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="anatolicum" subGenus="N.">N. (N.) anatolicum</taxonomicName>
</td>
<td id="C8F39A9E6DBFC39F1F0FCFE66D119E69" colspan="1" pageId="0" pageNumber="165" rowspan="1">9</td>
<td id="32256C944252CD1392BDB71F8C61040C" colspan="1" pageId="0" pageNumber="165" rowspan="1">4</td>
<td id="1B629CFF39DD96B125AD2D279F5801A0" colspan="1" pageId="0" pageNumber="165" rowspan="1">10</td>
<td id="3762D33B069A267CE9ED8BA1D06A4C94" colspan="1" pageId="0" pageNumber="165" rowspan="1">7</td>
<td id="29562507913EEFEA46C06B7988F7EAFF" colspan="1" pageId="0" pageNumber="165" rowspan="1">6</td>
<td id="F3CEF7D5CD693D6B581904A102536448" colspan="1" pageId="0" pageNumber="165" rowspan="1">11</td>
</tr>
<tr id="AE978B7186B21781669083E34A671B26" pageId="0" pageNumber="165">
<td id="A8D623A3AF186392769CB5B3C758E923" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="C9C90AC346E372942F31ACF788D535FA" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) artaxerxesi" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="artaxerxesi" subGenus="N.">N. (N.) artaxerxesi</taxonomicName>
</td>
<td id="1E4F37BA0584E19084694B788366FE73" colspan="1" pageId="0" pageNumber="165" rowspan="1">10-13</td>
<td id="BF2A2127E9022852E53767BCBD5F2B92" colspan="1" pageId="0" pageNumber="165" rowspan="1">5</td>
<td id="0C73EEC898302E8A3EF1A1AA15180EC9" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-7</td>
<td id="EF63249BCAE5678B47B538EFE426BBEF" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-6</td>
<td id="7E01C7D0C766C1EBC01A8D67F4E54051" colspan="1" pageId="0" pageNumber="165" rowspan="1">4-6</td>
<td id="51D466A8153E603B572D497B9305E372" colspan="1" pageId="0" pageNumber="165" rowspan="1">9-13</td>
</tr>
<tr id="EEBAC67EE0B6F3FECBA374E04F2E6757" pageId="0" pageNumber="165">
<td id="4E3DF0A59E9E6FD92802C1B1F8220B54" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="30CAFB12EF88A44D18F30DD56EFF96E5" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) catherineae" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="catherineae" subGenus="N.">N. (N.) catherineae</taxonomicName>
</td>
<td id="E21B2AED9BEFAA7C270292C585B55891" colspan="1" pageId="0" pageNumber="165" rowspan="1">12</td>
<td id="0AC41717FDBA8D417278BF974AB6E256" colspan="1" pageId="0" pageNumber="165" rowspan="1">4</td>
<td id="53586CD6CA27BD97457859336250923E" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-7</td>
<td id="7E36B56AD85BF677626A6C4931028F78" colspan="1" pageId="0" pageNumber="165" rowspan="1">6</td>
<td id="F8AE405CE4ECF5DDBD0A7485794D2B6A" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-6</td>
<td id="A617F20C10CF71D4998FD393136CD1C7" colspan="1" pageId="0" pageNumber="165" rowspan="1">7-8</td>
</tr>
<tr id="784E993BDCC9586741B646FEEB9F9228" pageId="0" pageNumber="165">
<td id="58DAC74DB8428A568C8421A8C05F315F" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="53392B1E896D7EF676DB15FC1D3592D7" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) crassifemoratum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="crassifemoratum" subGenus="N.">N. (N.) crassifemoratum</taxonomicName>
</td>
<td id="EA29D9E3883CD5F17CB1EDFE7BF5F635" colspan="1" pageId="0" pageNumber="165" rowspan="1">8-9</td>
<td id="83CB5D2E90315B3BAC1DFA22EFB9DCB1" colspan="1" pageId="0" pageNumber="165" rowspan="1">3</td>
<td id="33F824D3711AFAA7512E17D80C88F1FF" colspan="1" pageId="0" pageNumber="165" rowspan="1">7-9</td>
<td id="BFB3FB9D23585AB83B622A43CD8B7BD8" colspan="1" pageId="0" pageNumber="165" rowspan="1">7-8</td>
<td id="D661338326564C1EFB8CA19BB810A07A" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-6</td>
<td id="4DE58AFFA923184764BB7113520DD518" colspan="1" pageId="0" pageNumber="165" rowspan="1">9-11</td>
</tr>
<tr id="DD5E5E400F9047572FBC3DDEC6A743FF" pageId="0" pageNumber="165">
<td id="C8C7E2E5F1990891DD157F39A925C377" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="3CEE912DE22A0673D289AF75A9F838D3" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) golovatchi" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="golovatchi" subGenus="N.">N. (N.) golovatchi</taxonomicName>
</td>
<td id="9FC2399AD3E87910F53B2C0FBAED344A" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-7</td>
<td id="E39122E561BB8B1F961CD42E37981CEE" colspan="1" pageId="0" pageNumber="165" rowspan="1">4</td>
<td id="6501F5817932CF8EDB081DC652FFC301" colspan="1" pageId="0" pageNumber="165" rowspan="1">4-7</td>
<td id="7616DFBE880F67D7B405E044E4B32E4E" colspan="1" pageId="0" pageNumber="165" rowspan="1">4-5</td>
<td id="13735D3DFACDCBC103080F207CF2C068" colspan="1" pageId="0" pageNumber="165" rowspan="1">3-5</td>
<td id="FE1B333DDC8ABE89153C10B7A19420AC" colspan="1" pageId="0" pageNumber="165" rowspan="1">9-10</td>
</tr>
<tr id="F915618CC7696C0DFFA8ACAB747341FC" pageId="0" pageNumber="165">
<td id="0C1E09DFCA37540659F5BC8BAAB7465D" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="D9EE137AF768A77F938291E57FAEA6D4" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) kamenskyi" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="kamenskyi" subGenus="N.">N. (N.) kamenskyi</taxonomicName>
</td>
<td id="E236D8D3B2E7ECF1B7BF1E74E25BA9F6" colspan="1" pageId="0" pageNumber="165" rowspan="1">9</td>
<td id="942A5B93E74BBC7283E86C81E143ECC9" colspan="1" pageId="0" pageNumber="165" rowspan="1">5</td>
<td id="B1F322B11D28A506B1F34494F04B2D79" colspan="1" pageId="0" pageNumber="165" rowspan="1">6</td>
<td id="DFCD99A04802E019F27A96A7CD2BB3AC" colspan="1" pageId="0" pageNumber="165" rowspan="1">6</td>
<td id="AD1C0E1B7361D3F130B7500C22AFDA9C" colspan="1" pageId="0" pageNumber="165" rowspan="1">6</td>
<td id="68797377979E2C970ADAA56583B5A9BE" colspan="1" pageId="0" pageNumber="165" rowspan="1">8</td>
</tr>
<tr id="B0CCA780C72F51532262766B5F146D3E" pageId="0" pageNumber="165">
<td id="93C2FD89602CD7CD9E5BE9694AE4BD6E" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="16F9F8A9DD1FF40DDA998469A8C56698" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) kobachidzei" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="kobachidzei" subGenus="N.">N. (N.) kobachidzei</taxonomicName>
</td>
<td id="8669C38C60E6B606CD2A7E1390086063" colspan="1" pageId="0" pageNumber="165" rowspan="1">10-12</td>
<td id="53BE5AA9506F6384E2A8AD6D983F07FF" colspan="1" pageId="0" pageNumber="165" rowspan="1">4</td>
<td id="7A3002358F00EF969CC6C598D0F9A21F" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-7</td>
<td id="CA44C616CC85781D1354EBC9B5C55061" colspan="1" pageId="0" pageNumber="165" rowspan="1">7</td>
<td id="000401B79743AE57B11A97B42D1EFFC2" colspan="1" pageId="0" pageNumber="165" rowspan="1">7</td>
<td id="9DE74A85133AC09F21C812BE29060D0A" colspan="1" pageId="0" pageNumber="165" rowspan="1">8-13</td>
</tr>
<tr id="12174CD225F4596040B3B7562418932B" pageId="0" pageNumber="165">
<td id="6534A2C24984CF3CC1258153D064FDA5" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="9F18313B08D260B2CA620867D0CEDB63" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) kovalevskayae" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="kovalevskayae" subGenus="N.">N. (N.) kovalevskayae</taxonomicName>
</td>
<td id="F363BF71E38AEC87379CB90C47EF9B83" colspan="1" pageId="0" pageNumber="165" rowspan="1">13</td>
<td id="94D8E8E5134D989C905FBF67828E77C9" colspan="1" pageId="0" pageNumber="165" rowspan="1">5</td>
<td id="8D0941DDC5CE7D7A2929293F3F266075" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-9</td>
<td id="A1B935AD2E3ABB88A843582166D2497F" colspan="1" pageId="0" pageNumber="165" rowspan="1">6</td>
<td id="C3CF0E331D072280E6602A98984FA590" colspan="1" pageId="0" pageNumber="165" rowspan="1">5-7</td>
<td id="ADCD15A23780C92338CFA2B845E2A54C" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-8</td>
</tr>
<tr id="4DD3FF87D3BDF6C21933172BE72A8EB4" pageId="0" pageNumber="165">
<td id="295A63B4B2C07591C28972950B38719E" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="B628FA0781D2830ABC98ED687A9C40C6" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) speleophilum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="speleophilum" subGenus="N.">N. (N.) speleophilum</taxonomicName>
</td>
<td id="5BDB615931ADA5D176DF29951154B3D8" colspan="1" pageId="0" pageNumber="165" rowspan="1">10-12</td>
<td id="1BB052887DA30818401F049A23879F6C" colspan="1" pageId="0" pageNumber="165" rowspan="1">4</td>
<td id="E2CF4665F06C0586278ADF6303224E0B" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-7</td>
<td id="FE03F45A018F52AA83C13B59987C1EC0" colspan="1" pageId="0" pageNumber="165" rowspan="1">4-5</td>
<td id="23AA2CA608DD8A45BBDAB8405CA6DB8E" colspan="1" pageId="0" pageNumber="165" rowspan="1">4-6</td>
<td id="0259F19A83D28020447F8434BC4252CB" colspan="1" pageId="0" pageNumber="165" rowspan="1">8-10</td>
</tr>
<tr id="103610E14347A6438B36448B19B13A9B" pageId="0" pageNumber="165">
<td id="38534A8642581693C6BF6E613205CEA4" colspan="1" pageId="0" pageNumber="165" rowspan="1">
<taxonomicName id="B06E57B17674511DE4934DECEB63BA65" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) vilcekii" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="vilcekii" subGenus="N.">N. (N.) vilcekii</taxonomicName>
</td>
<td id="14C77BB53F99DED3F9CE35CA48E68138" colspan="1" pageId="0" pageNumber="165" rowspan="1">12-15</td>
<td id="587AEE8DCAC483956B321D8013B5F9B7" colspan="1" pageId="0" pageNumber="165" rowspan="1">5</td>
<td id="B2DC23720D3E77105406009FF90B9ECC" colspan="1" pageId="0" pageNumber="165" rowspan="1">6-11</td>
<td id="2C640A37E83C292251BC32625E26B58D" colspan="1" pageId="0" pageNumber="165" rowspan="1">7-12</td>
<td id="73DE5A98AFD5E73066681E0EA7FD90DA" colspan="1" pageId="0" pageNumber="165" rowspan="1">7-11</td>
<td id="F9D11509FBCF27356F5FD2909AA173DD" colspan="1" pageId="0" pageNumber="165" rowspan="1">13-22</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection id="B5DAA05EB8BD29CD4EBA1947376CB4BD" pageId="0" pageNumber="165" type="distribution">
<paragraph id="223F5513413B803E9DAF01DE54743AAF" pageId="0" pageNumber="165">Distribution.</paragraph>
<paragraph id="871EEB1AB659E1C4DEB2753F4AD4F4BE" pageId="0" pageNumber="165">
Known only from two adjacent
<taxonomicName id="23F57AB1C8F305966F1BCC8F280D4D8D" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="A4092B69E56E219C2FDCD1F764D670EA" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
bogs, Ispani 1 and Ispani 2, situated near the town of Kobuleti in the Republic of Adjara (Georgia).
</paragraph>
</subSubSection>
<subSubSection id="88EA660A704D676FFCC6D88B27DDD694" pageId="0" pageNumber="165" type="habitat">
<paragraph id="FEB99EDD3E96B4154F4C76760DB5078E" pageId="0" pageNumber="165">Habitats.</paragraph>
<paragraph id="66A4AC8526ED8AD1AC59B8A69B6DCE38" pageId="0" pageNumber="165">
In Ispani 1 bog,
<taxonomicName id="3656753667C1F5847D4A368626CEF502" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) adjaricum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="N.">N. (N.) adjaricum</taxonomicName>
sp. nov. was collected in its northwestern part (site 1), with large relatively dry hummocks (20-40 cm high, 1-2 m wide) and moist flat spaces between them (Fig.
<figureCitation id="351281A28847556717A3F844E28E6370" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The habitat of Neobisium (Neobisium) adjaricum sp. nov.: Ispani 1 bog near the town of Kobuleti (Republic of Adjara, Georgia) A general view of site 1 (30 April 2019) B Sphagnum hummock, with exact location where two paratypes were collected (sample I- 18) indicated by an arrow (11 May 2019). Photographs by AP." figureDoi="10.3897/zookeys.1100.81910.figure2" httpUri="https://binary.pensoft.net/fig/684536" pageId="0" pageNumber="165">2A</figureCitation>
). The new species was collected only from hummocks. The hummocks consisted mainly of loose
<taxonomicName id="6EE3B7F7C6603C6F1EF8DB9BF71FD3FF" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="09F19EDE91A22E14ACCB2E52783BD6EB" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
(with predominant
<taxonomicName id="49629D66EBB24EE15516134980168EBF" lsidName="S. papillosum" pageId="0" pageNumber="165" rank="species" species="papillosum">
<emphasis id="AF5DAE9593160DDB2F7B45C14D965E88" italics="true" pageId="0" pageNumber="165">S. papillosum</emphasis>
</taxonomicName>
,
<taxonomicName id="903B2907EE225470FB913CF4E008AD71" lsidName="S. austinii" pageId="0" pageNumber="165" rank="species" species="austinii">
<emphasis id="E55ADBD984E6A203E3550587D26452F6" italics="true" pageId="0" pageNumber="165">S. austinii</emphasis>
</taxonomicName>
, and
<taxonomicName id="2DABADC49D3653BF9F97FB8E149FC56E" lsidName="S. palustre" pageId="0" pageNumber="165" rank="species" species="palustre">
<emphasis id="7F5F149AB2CCC15DA6F085F0C91E0652" italics="true" pageId="0" pageNumber="165">S. palustre</emphasis>
</taxonomicName>
; in particular, the latter species was predominant in sample I-37, where one paratype was collected; Fig.
<figureCitation id="2172F65A0B43F50B5466213E7F3B967B" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. The habitat of Neobisium (Neobisium) adjaricum sp. nov.: Ispani 1 bog near the town of Kobuleti (Republic of Adjara, Georgia) A general view of site 1 (30 April 2019) B Sphagnum hummock, with exact location where two paratypes were collected (sample I- 18) indicated by an arrow (11 May 2019). Photographs by AP." figureDoi="10.3897/zookeys.1100.81910.figure2" httpUri="https://binary.pensoft.net/fig/684536" pageId="0" pageNumber="165">2B</figureCitation>
), with common
<taxonomicName id="DCE14AFE99BD21095F7BA5FF54364DF5" class="Liliopsida" family="Juncaceae" genus="Juncus" kingdom="Plantae" lsidName="Juncus effusus" order="Poales" pageId="0" pageNumber="165" phylum="Tracheophyta" rank="species" species="effusus">
<emphasis id="BB343DD659E25128A969C8E22D31AA31" italics="true" pageId="0" pageNumber="165">Juncus effusus</emphasis>
</taxonomicName>
L.,
<taxonomicName id="ED3E73784AA1AAC08D5DD36F06055DC8" class="Liliopsida" family="Poaceae" genus="Molinia" kingdom="Plantae" lsidName="Molinia caerulea subsp. arundinacea" order="Poales" pageId="0" pageNumber="165" phylum="Tracheophyta" rank="subSpecies" species="caerulea" subSpecies="arundinacea">
<emphasis id="CA6AA5E567127CE0798C2D9F1BBFB2E0" italics="true" pageId="0" pageNumber="165">Molinia caerulea arundinacea</emphasis>
</taxonomicName>
(Schrank) K. Richt.,
<taxonomicName id="AF1789D238E6C733C8950EF844CDF61E" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" lsidName="Rhododendron" order="Ericales" pageId="0" pageNumber="165" phylum="Tracheophyta" rank="genus">
<emphasis id="E1C9BC46018736F477590650109D5EEE" italics="true" pageId="0" pageNumber="165">Rhododendron</emphasis>
</taxonomicName>
spp., and
<taxonomicName id="7401E1356CACF5D1829EC905DA1BB763" class="Polypodiopsida" family="Dennstaedtiaceae" genus="Pteridium" kingdom="Plantae" lsidName="Pteridium tauricum" order="Polypodiales" pageId="0" pageNumber="165" phylum="Tracheophyta" rank="species" species="tauricum">
<emphasis id="E1DDD4DB835DC56797368175BAB2FDD0" italics="true" pageId="0" pageNumber="165">Pteridium tauricum</emphasis>
</taxonomicName>
V.I. Krecz.
<taxonomicName id="7E9AA80B69414C2C53896286FF4D8809" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="871C9CEE27D3FC796F37CA43E1BA7676" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
was dry to slightly wet, with pH 2.9-3.0 and mineralization of 0.04-0.08 ppt.
</paragraph>
<paragraph id="33AC0F46FDA29255DE43C7C47FB745FB" pageId="0" pageNumber="165">
In Ispani 2 bog,
<taxonomicName id="8684EEB6CD166E68B42E1F8996D35414" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) adjaricum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="N.">N. (N.) adjaricum</taxonomicName>
sp. nov. was collected from a flat site in its (site 3; Fig.
<figureCitation id="C6DE0CE456518D4F452F9739BF1BF2B6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The habitat of Neobisium (Neobisium) adjaricum sp. nov.: Ispani 2 bog near the town of Kobuleti (Republic of Adjara, Georgia) A general view of site 3 (10 May 2019) B Sphagnum carpet, with exact location where the holotype was collected (sample I- 11) indicated by an arrow (10 May 2019). Photographs by AP." figureDoi="10.3897/zookeys.1100.81910.figure1" httpUri="https://binary.pensoft.net/fig/684535" pageId="0" pageNumber="165">1A</figureCitation>
) covered by a flat carpet of
<taxonomicName id="E0C233E55E932DC01F8B39E7D405CA94" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="0C9F8FF5967EFB92A4409F2F495C6A26" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
, with
<taxonomicName id="7B4423644A20776B2B17425A5AB2C0C5" lsidName="S. papillosum" pageId="0" pageNumber="165" rank="species" species="papillosum">
<emphasis id="026402BF19637F30AEA79B1582D7AEDC" italics="true" pageId="0" pageNumber="165">S. papillosum</emphasis>
</taxonomicName>
and
<taxonomicName id="4FB03B52873DDAE597D834802001C8A6" lsidName="S. palustre" pageId="0" pageNumber="165" rank="species" species="palustre">
<emphasis id="B7743F59DF2AB97A2CBDC4FFC8617727" italics="true" pageId="0" pageNumber="165">S. palustre</emphasis>
</taxonomicName>
predominant, and with the thinned stands of
<taxonomicName id="DF447E00C9DB038383CE971666F2B25D" lsidName="M. caerulea subsp. arundinacea" pageId="0" pageNumber="165" rank="subSpecies" species="caerulea" subSpecies="arundinacea">
<emphasis id="FE4CB1BDD0D08583E2AB4C516C0ED6F2" italics="true" pageId="0" pageNumber="165">M. caerulea arundinacea</emphasis>
</taxonomicName>
(Fig.
<figureCitation id="E5739552932F2301FE0CB097B1D975AC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. The habitat of Neobisium (Neobisium) adjaricum sp. nov.: Ispani 2 bog near the town of Kobuleti (Republic of Adjara, Georgia) A general view of site 3 (10 May 2019) B Sphagnum carpet, with exact location where the holotype was collected (sample I- 11) indicated by an arrow (10 May 2019). Photographs by AP." figureDoi="10.3897/zookeys.1100.81910.figure1" httpUri="https://binary.pensoft.net/fig/684535" pageId="0" pageNumber="165">1B</figureCitation>
).
<taxonomicName id="EDD842DBA6B0813172357D617E751FA7" authorityName="C.Linnaeus" authorityYear="1753" class="Arachnida" family="Sphagnaceae" genus="Sphagnum" kingdom="Plantae" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="165" phylum="Bryophyta" rank="genus">
<emphasis id="5FAF21263E9F5E3EBBA77CD2304470DA" italics="true" pageId="0" pageNumber="165">Sphagnum</emphasis>
</taxonomicName>
was wet to moist, with a pH of 4.0 and mineralization of 0.07 ppt.
</paragraph>
<paragraph id="20BAA14090262411BE07D413FCDC8069" pageId="0" pageNumber="165">
The new species seems to be rare, considering that only 1-3 specimens were collected at each site, in 10 quantitative samples (sample size of 0.05 m2). As distinct from other pseudoscorpion species, all specimens of
<taxonomicName id="E10F0C4B129CDEFFB835404C22927AEB" class="Arachnida" kingdom="Animalia" lsidName="N. (N.) adjaricum" order="Pseudoscorpionida" pageId="0" pageNumber="165" phylum="Arthropoda" rank="species" species="adjaricum" subGenus="N.">N. (N.) adjaricum</taxonomicName>
sp. nov. were collected only from quantitative samples and no specimens were collected by other techniques (sifting, pitfall trapping, and/or rearing from substrata in the lab).
</paragraph>
</subSubSection>
<subSubSection id="1D9381CF6F6FC90DF09B0777F144015D" pageId="0" pageNumber="165" type="etymology">
<paragraph id="740A2DB6E952574DBE0EB8C6B7A43090" pageId="0" pageNumber="165">Etymology.</paragraph>
<paragraph id="9EB540A25BC38BA0C0456746BFDD99B6" pageId="0" pageNumber="165">
The Latin adjective
<emphasis id="90967AE57591AE1B39A308830EF7BA28" italics="true" pageId="0" pageNumber="165">adjaricum</emphasis>
indicates that the new species is named after the Republic of Adjara (= Ajaria), Georgia, where it was discovered.
</paragraph>
</subSubSection>
</treatment>
</document>