146 lines
12 KiB
XML
146 lines
12 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.732.21677" ID-GBIF-Dataset="05db50e8-b4e5-422a-b78f-9676d74ac319" ID-PMC="PMC5799741" ID-Pensoft-Pub="1313-2970-732-1" ID-PubMed="29416404" ID-ZBK="9995702B61464BA1BB5323DC9BA9650F" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-732-1" ModsDocOrigin="ZooKeys 732" ModsDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" checkinTime="1516831538731" checkinUser="pensoft" docAuthor="Mattia, Willy De, Neiber, Marco T. & Groh, Klaus" docDate="2018" docId="4DC7D21A08CD803FCFDBCEBB020520A6" docLanguage="en" docName="ZooKeys 732: 1-125" docOrigin="ZooKeys 732" docSource="http://dx.doi.org/10.3897/zookeys.732.21677" docTitle="Hystricella microcarinata De Mattia & Groh, sp. n." docType="treatment" docUuid="EE848F87-CEBC-4430-9D7F-4D00D2EE2FF9" docUuidSource="ZooBank" docVersion="6" lastPageNumber="18" masterDocId="FFA3FF812B32FFA8FFBAFFCDFF9AFF92" masterDocTitle="Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)" masterLastPageNumber="125" masterPageNumber="1" pageNumber="17" updateTime="1668165370779" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus-group Hystricella R. T. Lowe, 1855 from Porto Santo (Madeira Archipelago), with descriptions of new recent and fossil taxa (Gastropoda, Helicoidea, Geomitridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mattia, Willy De</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Neiber, Marco T.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Groh, Klaus</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>732</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>125</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.732.21677</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.732.21677</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-732-1</mods:identifier>
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<mods:identifier type="ZBK">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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<mods:identifier type="ZooBank">9995702B61464BA1BB5323DC9BA9650F</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="140417754" LSID="urn:lsid:zoobank.org:act:EE848F87-CEBC-4430-9D7F-4D00D2EE2FF9" httpUri="http://treatment.plazi.org/id/4DC7D21A08CD803FCFDBCEBB020520A6" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
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<subSubSection pageId="16" pageNumber="17" type="nomenclature">
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<paragraph pageId="16" pageNumber="17">
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†
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<taxonomicName LSID="http://zoobank.org/EE848F87-CEBC-4430-9D7F-4D00D2EE2FF9" authority="De Mattia & Groh" class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella microcarinata" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="microcarinata">Hystricella microcarinata De Mattia & Groh</taxonomicName>
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<taxonomicNameLabel pageId="16" pageNumber="17">sp. n.</taxonomicNameLabel>
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Figs 74, 76
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="type material">
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<paragraph pageId="16" pageNumber="17">Type material.</paragraph>
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<paragraph pageId="16" pageNumber="17">SMF 348924, holotype, from loc. typ. (locus typicus), leg. W. De Mattia & J. Macor, May 2016.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="locus typicus">
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<paragraph pageId="16" pageNumber="17">Locus typicus.</paragraph>
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<paragraph pageId="16" pageNumber="17">
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Porto Santo, E of Vila Baleira, S slope of the hill above Vale do Touro, 50 m W of the oil tanks, excavated Quaternary mixed gravel,
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="33.063053">33°03'47"N</geoCoordinate>
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/
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<geoCoordinate direction="west" orientation="longitude" precision="15" value="-16.32389">16°19'26"W</geoCoordinate>
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, 24 m.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="diagnosis">
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<paragraph pageId="16" pageNumber="17">Diagnosis.</paragraph>
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<paragraph pageId="16" pageNumber="17">
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Hystricella</taxonomicName>
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species with two keels on the body whorl, the upper less developed than the lower one; growth lines rather coarse, giving the impression of an irregular ribbing; tubercles on the base of the shell rather scattered and not as prominent as in
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<taxonomicName lsidName="H. bicarinata" pageId="16" pageNumber="17" rank="species" species="bicarinata">H. bicarinata</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="description">
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<paragraph pageId="16" pageNumber="17">Description of the shell of the holotype.</paragraph>
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<paragraph pageId="16" pageNumber="17">Shell very small for the genus, with 5⅓ regularly increasing whorls, the protoconch with 1.6 whorls. The form of the shell is rounded conical, the convex teleoconch whorls exhibit two flat keels that become more and more expressed; the upper keel is much less developed than the lower one; keels on the body whorl located in its upper half. The last whorl measures 69%, the penultimate whorl 15% of the total shell height. The lower half of the body whorl is, beneath the distinctly angled periphery, in frontal view nearly straight, only very slightly convex. The suture between the whorls is simple, not sunken. The aperture, which is inclined to the vertical axis of the shell in an angle of 50° and is descending in the last 5% of the last whorl in an angle of 36° to the horizontal axis, has an oblique-ovate form, its width is 43% of the shell width, its height 34% of the shell height. It has a slightly reflected lip, which is completely detached from the body whorl. The eccentric umbilicus, which is approximately 11% as wide as the shell, is shaped in the upper whorls like a pinhole. The protoconch is smooth. The teleoconch exhibits a number of oblique radial ribs, 16 in the penultimate quadrant of the body whorl and is additionally covered by numerous rough tubercles. The number of tubercles in the standard-quadrate of the base is 43. There are no traces of colouration (Fig. 74).</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="description">
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<paragraph pageId="16" pageNumber="17">Measurements.</paragraph>
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<paragraph pageId="16" pageNumber="17">D 4.1; H 4.2 mm; FW 2.7 mm; PA 49°; DU 0.3 mm; NT 12; NW 4.8 (n = 1). Ratio D/H 0.9; ratio FW/H 0.6.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="17" type="distribution">
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<paragraph pageId="16" pageNumber="17">Distribution.</paragraph>
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<paragraph pageId="16" pageNumber="17">
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The species is only known from the type locality where it appears to be extremely rare (Fig. 76). Its close resemblance to other (sub-) fossil
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="genus">Hystricella</taxonomicName>
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taxa may have led to misidentifications during previous collecting and sorting, thus the distribution area of the species may possibly become larger after further investigations.
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</paragraph>
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<caption pageId="16" pageNumber="17">
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<paragraph pageId="16" pageNumber="17">
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Figure 76. Distribution of
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Hystricella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hystricella microcarinata" order="Stylommatophora" pageId="16" pageNumber="17" phylum="Mollusca" rank="species" species="microcarinata">Hystricella microcarinata</taxonomicName>
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sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="etymology">
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<paragraph pageId="17" pageNumber="18">
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<pageBreakToken pageId="17" pageNumber="18" start="start">Etymology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="17" pageNumber="18">
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The name is a combination of the Greek name for small (μικρός =
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<normalizedToken originalValue="mikrós">mikros</normalizedToken>
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) and the Latin name for keeled (carinatus) and alludes to the small size of the shell with a keel at its periphery.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="comparisons">
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<paragraph pageId="17" pageNumber="18">Comparison and comments.</paragraph>
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<paragraph pageId="17" pageNumber="18">
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Being a (sub-) fossil species, only the shell features can be taken into account for comparisons. The globose shape is similar to that of some forms of
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<taxonomicName lsidName="H. echinulata" pageId="17" pageNumber="18" rank="species" species="echinulata">H. echinulata</taxonomicName>
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; nevertheless the overall dimensions and surface sculpture of the shell clearly distinguish
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<taxonomicName lsidName="H. microcarinata" pageId="17" pageNumber="18" rank="species" species="microcarinata">H. microcarinata</taxonomicName>
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sp. n. Because of its small size, the new species can also be confused with small specimens of
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<taxonomicName lsidName="H. aucta" pageId="17" pageNumber="18" rank="species" species="aucta">H. aucta</taxonomicName>
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Wollaston, 1878 which, however, possesses two well-developed keels, has a wider umbilicus, a different shape and differently positioned angles of the aperture, or with
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia beckmanni" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="beckmanni">Wollastonia beckmanni</taxonomicName>
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gen. et sp. n., which has a much flatter spire, has no keels on the whorls, a much wider umbilicus and also a different shape and differently positioned angles of the aperture.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="taxonomic remarks">
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<paragraph pageId="17" pageNumber="18">Taxonomic remarks.</paragraph>
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<paragraph pageId="17" pageNumber="18">
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The generic affiliation of this species is based exclusively on the shape of the shell and the presence of keels on the body whorl. A similar arrangement of the tubercles and keels can be found also in some
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<taxonomicName class="Gastropoda" family="Geomitridae" genus="Wollastonia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wollastonia" order="Stylommatophora" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">Wollastonia</taxonomicName>
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gen. n. species that will be described below.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="18" type="status and conservation">
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<paragraph pageId="17" pageNumber="18">Status and conservation.</paragraph>
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<paragraph pageId="17" pageNumber="18">
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Extinct before the
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<normalizedToken originalValue="islands’">islands'</normalizedToken>
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scientific exploration in the 19th century, possibly already before human settlement.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |