184 lines
20 KiB
XML
184 lines
20 KiB
XML
<document id="3D47FF9BFC25B26A2B9268022952DF33" ID-DOI="http://dx.doi.org/10.3897/BDJ.1.e957" ID-PMC="PMC3964709" ID-Pensoft-Pub="1314-2828--957" ID-PubMed="24723759" ID-ZBK="CABF7796A0F44526917B5286F2C7CA0C" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1314-2828--e957" ModsDocOrigin="Biodiversity Data Journal " ModsDocTitle="The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp.n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida)" checkinTime="1553126941720" checkinUser="pensoft" docAuthor="Golovatch, Sergei & Stoev, Pavel" docDate="2013" docId="4D607CE9A8CE926F0928776D0BCCC1F5" docLanguage="en" docName="BiodivDatJour 1: e957" docOrigin="Biodiversity Data Journal 1" docSource="http://dx.doi.org/10.3897/BDJ.1.e957" docTitle="Eustrongylosoma penevi Golovatch & Stoev 2013, sp. n." docType="treatment" docUuid="3A600B5D-4791-44CE-AC7C-89BF768789D9" docUuidSource="ZooBank" docVersion="6" lastPageNumber="957" masterDocId="F979FFB6721F7E1F78569D4FFF85FFB0" masterDocTitle="The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp. n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida)" masterLastPageNumber="957" masterPageNumber="957" pageNumber="957" updateTime="1668122260419" updateUser="ExternalLinkService">
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<mods:mods id="7CA109DFFD7AAE52F147A101333C3A96" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="774C06B9559284B609E17867347EF28F">
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<mods:title id="B58329F6545A809970ED769F1FB8E079">The millipede family Paradoxosomatidae in the Philippines, with a description of Eustrongylosomapenevi sp. n., and notes on Anoplodesmusanthracinus Pocock, 1895, recorded in Malaysia and Sri Lanka for the first time (Diplopoda, Polydesmida)</mods:title>
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<mods:name id="3CC6D4CE1766D87C185FEE10A6B4E538" type="personal">
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<mods:roleTerm id="B7904AA7CD972038B58CFDAF15E883BB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0E113C873071459278C270826A4963D2">Golovatch, Sergei</mods:namePart>
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<mods:name id="42A96A5F2540FA2F5ABA265CAAFAC65E" type="personal">
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<mods:roleTerm id="0E4AA9FEA5D163DCF25688295F885777">Author</mods:roleTerm>
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<mods:namePart id="9E6C3758F8FAEF242F4CD02C1E7FD2C6">Stoev, Pavel</mods:namePart>
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<mods:typeOfResource id="AC66F43411206F54202A0921671C04C6">text</mods:typeOfResource>
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<mods:titleInfo id="EF36939E35B550CB68FD091579C96720">
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<mods:title id="92440166C17645B4C126FC0A3576648F">Biodiversity Data Journal</mods:title>
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</mods:titleInfo>
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<mods:part id="C6DAB18828491537B67BDD2E4542218F">
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<mods:date id="52854B44F0C9F115978B2E2483948D49">2013</mods:date>
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<mods:detail id="0B10513B19779322F47F8C39895F58CD" type="volume">
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<mods:number id="C02A3EA6F6B1EA9462F804FD27484222">1</mods:number>
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</mods:detail>
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<mods:extent id="5DFAF0C63CA36696C4A684C5A5194426" unit="page">
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<mods:start id="847B81FE92ACADD8D22D29AF33B97538">957</mods:start>
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<mods:end id="9502385375150933853A45F798F7F8B9">957</mods:end>
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<mods:location id="6449ED3F390CC584BDDAA4AFDFF409D5">
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<mods:url id="54C860FCBE4557662871342BC3F6D028">http://dx.doi.org/10.3897/BDJ.1.e957</mods:url>
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</mods:location>
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<mods:classification id="9A503FD9B8BC1963FAEC91B59EE6612F">journal article</mods:classification>
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<mods:identifier id="CD283C22846113C699011259DFDA0B9E" type="DOI">http://dx.doi.org/10.3897/BDJ.1.e957</mods:identifier>
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<mods:identifier id="C02083B323C45B3390A7CCDE714ECC01" type="Pensoft-Pub">1314-2828--957</mods:identifier>
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<mods:identifier id="C255044D61FF3994A905D93902029445" type="ZBK">CABF7796A0F44526917B5286F2C7CA0C</mods:identifier>
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<mods:identifier id="12A5662C0F71831CEF977212D6A1E213" type="ZooBank">CABF7796A0F44526917B5286F2C7CA0C</mods:identifier>
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</mods:mods>
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<treatment id="4D607CE9A8CE926F0928776D0BCCC1F5" LSID="urn:lsid:zoobank.org:act:3A600B5D-4791-44CE-AC7C-89BF768789D9" httpUri="http://treatment.plazi.org/id/4D607CE9A8CE926F0928776D0BCCC1F5" lastPageNumber="957" pageId="0" pageNumber="957">
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<subSubSection id="5343925D3958DD287AF00F0FB675C77A" pageId="0" pageNumber="957" type="nomenclature">
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<paragraph id="2BABE0133F4DED714DDD106325D6006A" pageId="0" pageNumber="957">
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<taxonomicName id="472AF49A255FCBBDFBD910DC5F16FE49" ID-CoL="3DBH2" LSID="3A600B5D-4791-44CE-AC7C-89BF768789D9" authority="Golovatch & Stoev, 2013" authorityName="Golovatch & Stoev" authorityYear="2013" class="Diplopoda" family="Paradoxosomatidae" genus="Eustrongylosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eustrongylosoma penevi" order="Polydesmida" pageId="0" pageNumber="957" phylum="Arthropoda" rank="species" species="penevi">Eustrongylosoma penevi Golovatch & Stoev, 2013</taxonomicName>
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<taxonomicNameLabel id="24B50E7E2F49F879FDD184E1B4E720BB" pageId="0" pageNumber="957">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="B6F3705D314E3DB38B5B71114EEBB2E4" pageId="0" pageNumber="957" type="materials_examined">
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<paragraph id="1754EE48D26DC4064DBD7E3227681DDD" pageId="0" pageNumber="957">Materials</paragraph>
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<paragraph id="D735412A85EB67E9C8D242F8627A6E06" pageId="0" pageNumber="957">
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<materialsCitation id="C17017BBCF28883785F83560D1D5A635" collectingDate="2012-07-06" collectionCode="NMNHS" collectorName="P. Stoev & L. Penev" country="Philippines" latitude="16.966112" location="Philippines" longLatPrecision="21" longitude="121.02695" pageId="0" pageNumber="957" specimenCount="1" specimenCount-male="1">
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Occurrence: recordedBy:
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<collectorName id="263CFDD678B9CB838D390962A3C8D80E" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="20E503C30E9A30AFBED15A1023DA3A0B" pageId="0" pageNumber="957">1</specimenCount>
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; sex:
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<specimenType id="F4530A7388749FA8C9F7223119FA1036" pageId="0" pageNumber="957">male</specimenType>
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; Location: island: Luzon Island; country:
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<collectingCountry id="DB66D9B050FA6F5EBE0DC94C95437C9C" pageId="0" pageNumber="957">Philippines</collectingCountry>
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; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation:
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<elevation id="EA296546175095F07313CA222F528C6F" pageId="0" pageNumber="957">1800-1900 m</elevation>
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; locationRemarks: under stones and logs; verbatimLatitude:
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<geoCoordinate id="6A0AD535ABE27BFA941D408895146160" direction="north" orientation="latitude" precision="15" value="16.966112">16°57'58"N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="B9658AC3C704F8242EBBA58FD12BBB1A" direction="east" orientation="longitude" precision="15" value="121.02695">121°1'37"E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="2AA0CED513C748C13C9D387D9E72C759" pageId="0" pageNumber="957" value="2012-07-06">6 July 2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="93DC1F3F9C6A0C9E7947E58E5F733F2E" pageId="0" pageNumber="957">NMNHS</collectionCode>
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</materialsCitation>
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<materialsCitation id="6C1303CE9AF5FF97202821F470811ED9" collectingDate="2012-07-06" collectionCode="NMNHS" collectorName="P. Stoev & L. Penev" country="Philippines" latitude="16.966112" location="Philippines" longLatPrecision="21" longitude="121.02695" pageId="0" pageNumber="957" specimenCount="4" specimenCount-male="4">
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Occurrence: recordedBy:
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<collectorName id="4BAC1FBD61B117B26F7E8BB9F6E327CB" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="A7BDECFE9F88BCFD5AB74E506DFA6A4C" pageId="0" pageNumber="957">4</specimenCount>
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; sex:
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<specimenType id="4DDBACFAD61CBF55418648B6833A2C3B" pageId="0" pageNumber="957">male</specimenType>
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; Location: island: Luzon Island; country:
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<collectingCountry id="B5C15553AEE39364E3573A041FB54987" pageId="0" pageNumber="957">Philippines</collectingCountry>
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; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation:
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<elevation id="45CA5D861C5D0BD74C1179762C39E35E" pageId="0" pageNumber="957">1800-1900 m</elevation>
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; locationRemarks: under stones and logs; verbatimLatitude:
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<geoCoordinate id="6268767D4E36A80AAA255B918CE9EA79" direction="north" orientation="latitude" precision="15" value="16.966112">16°57'58"N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="2D779DBC6CFD4BF9EEE04C41FDDD6984" direction="east" orientation="longitude" precision="15" value="121.02695">121°1'37"E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="627FEA5E4150871AA6F4AC54990124D6" pageId="0" pageNumber="957" value="2012-07-06">6 July 2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="35C3BAA92240052288CBBDCE5D4F73D7" pageId="0" pageNumber="957">NMNHS</collectionCode>
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</materialsCitation>
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<materialsCitation id="B4A2B3EB19E5A50CF9C4FC61D49FC225" collectingDate="2012-07-06" collectionCode="NMNHS" collectorName="P. Stoev & L. Penev" country="Philippines" latitude="16.966112" location="Philippines" longLatPrecision="21" longitude="121.02695" pageId="0" pageNumber="957" specimenCount="1" specimenCount-female="1">
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Occurrence: recordedBy:
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<collectorName id="DF722384D4CEE007CCFA601733303C5F" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="17A4B71FC73E17576486E5FF360C1E47" pageId="0" pageNumber="957">1</specimenCount>
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; sex:
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<specimenType id="0EDF860602FFA31DFCE057D88FC6CB91" pageId="0" pageNumber="957">female</specimenType>
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; Location: island: Luzon Island; country:
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<collectingCountry id="EDE8485F6E537A68D69B8C2844D6F45F" pageId="0" pageNumber="957">Philippines</collectingCountry>
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; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation:
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<elevation id="50692161C8EB296A7B6723CF04781A79" pageId="0" pageNumber="957">1800-1900 m</elevation>
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; locationRemarks: under stones and logs; verbatimLatitude:
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<geoCoordinate id="2763AF18729554AB415359CF6B5A96AC" direction="north" orientation="latitude" precision="15" value="16.966112">16°57'58"N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="C069EB826A7182A84E95B3DE8763639E" direction="east" orientation="longitude" precision="15" value="121.02695">121°1'37"E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="9E91F7B135D8F034513F9C255C1A8BF5" pageId="0" pageNumber="957" value="2012-07-06">6 July 2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="9AEDD7A04E574821A8CCD8C122175B67" pageId="0" pageNumber="957">NMNHS</collectionCode>
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</materialsCitation>
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<materialsCitation id="43A0CE65006094043457A6907EB6F70B" collectingDate="2012-07-06" collectionCode="NMNHS" collectorName="P. Stoev & L. Penev" country="Philippines" latitude="16.966112" location="Philippines" longLatPrecision="21" longitude="121.02695" pageId="0" pageNumber="957" specimenCount="1" specimenCount-juvenile="1">
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Occurrence: recordedBy:
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<collectorName id="60D0F366477E430EDEA0D8A41349B52D" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="5AFA073DE9CD86DA40061805FE6C59DB" pageId="0" pageNumber="957">1</specimenCount>
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; lifeStage:
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<specimenType id="F1540997419C2F4D523140DEED9595BD" pageId="0" pageNumber="957">juvenile</specimenType>
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; Location: island: Luzon Island; country:
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<collectingCountry id="8A042EF1EA19719CA40EAA1FE02B2296" pageId="0" pageNumber="957">Philippines</collectingCountry>
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|
; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation:
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<elevation id="39221B603C1BCF96CB27DC9529365287" pageId="0" pageNumber="957">1800-1900 m</elevation>
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|
; locationRemarks: under stones and logs; verbatimLatitude:
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<geoCoordinate id="F60FC85418CE59D44A928FE91A29AAD5" direction="north" orientation="latitude" precision="15" value="16.966112">16°57'58"N</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate id="E4CF6F4E8862DE381C7E0A20CD2240D2" direction="east" orientation="longitude" precision="15" value="121.02695">121°1'37"E</geoCoordinate>
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; Event: eventDate:
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<collectingDate id="AE694DBF52D950884F2D98DE3F865732" pageId="0" pageNumber="957" value="2012-07-06">6 July 2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="9013B803EBD1387313443B18D8CBC090" pageId="0" pageNumber="957">NMNHS</collectionCode>
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</materialsCitation>
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<materialsCitation id="D845BFF921D24430C493B6728BB09D7C" collectingDate="2012-07-06" collectionCode="ZMUM" collectorName="P. Stoev & L. Penev" country="Philippines" latitude="16.966112" location="Philippines" longLatPrecision="21" longitude="121.02695" pageId="0" pageNumber="957" specimenCount="1" specimenCount-male="1">
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Occurrence: recordedBy:
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<collectorName id="86F8440F2639DAE5DC689B35DAD37F90" pageId="0" pageNumber="957">P. Stoev & L. Penev</collectorName>
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; individualCount:
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<specimenCount id="7E865C2557A56CD12D2301DB44756B16" pageId="0" pageNumber="957">1</specimenCount>
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; sex:
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<specimenType id="555AEFE99359D02ED22071C50CC7EFE4" pageId="0" pageNumber="957">male</specimenType>
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; Location: island: Luzon Island; country:
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<collectingCountry id="2363DF27EBB8DB42420724697BCF71F3" pageId="0" pageNumber="957">Philippines</collectingCountry>
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; stateProvince: Mountain Province; verbatimLocality: Mt Polis Checkpoint on the road Banaue - Sagada; verbatimElevation:
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<elevation id="E870211130D3F0AFC05EA9EF9BDEF69A" pageId="0" pageNumber="957">1800-1900 m</elevation>
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|
; locationRemarks: under stones and logs; verbatimLatitude:
|
|
<geoCoordinate id="5E97818D96184EFDD6A233B322AC161B" direction="north" orientation="latitude" precision="15" value="16.966112">16°57'58"N</geoCoordinate>
|
|
; verbatimLongitude:
|
|
<geoCoordinate id="6320C27527DED496F67D939FE74DFEDA" direction="east" orientation="longitude" precision="15" value="121.02695">121°1'37"E</geoCoordinate>
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|
; Event: eventDate:
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|
<collectingDate id="0539E50C8C3A3674C97CFF97AFDACCB2" pageId="0" pageNumber="957" value="2012-07-06">6 July 2012</collectingDate>
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; Record Level: institutionCode:
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<collectionCode id="B0C35D4ADA46D42D8D1BAF5397E328A3" pageId="0" pageNumber="957">ZMUM</collectionCode>
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|
</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="751C5E42856CA110D13534B2DFD8C640" pageId="0" pageNumber="957" type="description">
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<paragraph id="DA2DB7919DCFDD53D8B3707C4606272E" pageId="0" pageNumber="957">Description</paragraph>
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<paragraph id="D723D1CBEB558D3FBFCBE73A547CF301" pageId="0" pageNumber="957">Length 18-22 (♂) or 23 mm (♀), width of midbody pro- and metazona 1.1-1.3 and 1.5-1.7 mm (♂), or 1.9 and 2.1 mm (♀), respectively. Holotype ca 22 mm long, width of pro- and metazona 1.3 and 1.6 mm, respectively. Coloration black to light grey-brown (Fig. 1a). Pattern mostly cingulate due to a large light grey band on prozona dorsally in front of stricture extending down until level of paraterga (Fig. 1a, b). Legs light grey-brown. Antennae increasingly infuscate distad, from light brown to blackish (Fig. 1a).</paragraph>
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<paragraph id="1358B9665ACB0B1A7C3A2571F826DE21" pageId="0" pageNumber="957">Body submoniliform. Antennomeres 2 to 6 subequal in length, antennae rather short, reaching behind segment 3 (♂) or 2 (♀) when stretched dorsally. Tegument generally smooth and shining. In width, segments 2 and 3 <collum = 4 <5-17, thereafter body gently tapering towards telson. Paraterga (Fig. 1b, c, d) modestly developed, keel-shaped, set low (at about 1/3rd of metazonal height), thinner in poreless, thicker in pore-bearing, segments, slightly reaching behind tergal margin only in segments 2 and 17-19, mostly slightly pointed, delimited by a complete and deep sulcus only dorsally, ventral sulcus being incomplete, developed only in posterior quarter to 1/5th extent. Head densely setose on clypeus and frons, bare on vertex. Collum semilunar, bearing two transverse rows of 2+2 setae, one row along front margin, second row in the middle; lateral edges broadly rounded. Metaterga faintly rugulose, a little more clearly so in postsulcus halves, surface below paraterga microgranular in segments 2-7. Tergal setae rather long, about 1/5th of metatergal length, arranged in two rows of 2+2 in each, one in front of, second behind sulcus. Sulcus starting from segment 5, deep, almost reaching the bases of paraterga. Stricture between pro- and metazona finely and densely ribbed. Ozopores lying close to caudal end of paraterga in a shallow ovoid groove, lateral, only partly visible from above. Pleurosternal carinae poorly developed ridges visible only in segments 2-4. Seta at about midway of each paratergum mostly broken off. Axial line wanting. Epiproct (Fig. 2a) subtruncate, pre-apical lateral papillae small. Hypoproct (Fig. 2b) semi-circular. Sternal lobe between coxae 4 subtrapeziform, densely setose (Fig. 2c). Legs very long and slender, about 2.0 (♂) or 1.3 (♀) times as long as midbody height; ♂ tarsal brushes traceable until about legs of segment 15, thereafter thinning out.</paragraph>
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<paragraph id="851EED90522F2844A1D8F2564D755267" pageId="0" pageNumber="957">Gonopods rather simple (Fig. 2d, e, f): coxite long, subcylindrical, bare; prefemoral portion small, about 1/3rd as long as femorite, the latter slender, ventral lobe somewhat better developed than dorsal one, apicolateral lobe (l) rounded, well developed, with a long transverse spine (s) at base. Solenophore subcircular, with a subterminal lobule.</paragraph>
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</subSubSection>
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<subSubSection id="8B55D7FBD9011289A874D5F72FA0AE63" pageId="0" pageNumber="957" type="diagnosis">
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<paragraph id="DED712A3C1B172A9B011B7D25778305F" pageId="0" pageNumber="957">Diagnosis</paragraph>
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<paragraph id="F8BCE07C41187915006B9AED16CFE216" pageId="0" pageNumber="957">
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Most similar to
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<taxonomicName id="CA6C7E24A10DCDD4C0DBBC4EFD781A16" lsidName="E. exiguum" pageId="0" pageNumber="957" rank="species" species="exiguum">E. exiguum</taxonomicName>
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Hoffman, 1978, from Papua New Guinea, and
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<taxonomicName id="69E8E54F51A270D758AC983C062DFB46" lsidName="E. kuekenthali" pageId="0" pageNumber="957" rank="species" species="kuekenthali">E. kuekenthali</taxonomicName>
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(Attems, 1897), from Borneo and Sulawesi, sharing the presence of a prominent distal spine on the gonopod femorite. Different from all congeners by the transverse orientation of the spine and noticeably long legs in the male (
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<bibRefCitation id="27B7B6A065B7C6C33DB577B4806A74DC" author="Hoffman, RL" journalOrPublisher="International Journal of Speleology" pageId="0" pageNumber="957" pagination="281 - 307" title="Diplopoda from Papuan Caves (Zoological Results of the British Speleological Expedition to Papua New Guinea, 1975, 4)." volume="9" year="1978">Hoffman 1978</bibRefCitation>
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,
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<bibRefCitation id="3B7CFF8DA5DA057E638EC4E857F2257F" author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="0" pageNumber="957" pagination="131 - 141" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda, Polydesmida), V." volume="5" year="1997">Golovatch 1997</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="FAB1C058DCFFF5912ABC108D33C97A0B" pageId="0" pageNumber="957" type="etymology">
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<paragraph id="BDE425762C698BE9AA579EB7015293F7" pageId="0" pageNumber="957">Etymology</paragraph>
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<paragraph id="C74C6D0B0156B08D4DD2518F788EE54E" pageId="0" pageNumber="957">Honours our good friend and colleague Lyubomir Penev, biologist and founder of the Biodiversity Data Journal and Pensoft Publishers. <br/></paragraph>
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</subSubSection>
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<subSubSection id="F90289100B6E9BC4B2734CD024780AD7" pageId="0" pageNumber="957" type="notes">
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<paragraph id="B8FDD569EA4412761929B7B970EBB7EC" pageId="0" pageNumber="957">Notes</paragraph>
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<paragraph id="1FF3C6F97EEBB6BD29A822784B288904" pageId="0" pageNumber="957">The species is hitherto known only from its type locality, Mt Polis Checkpoint on the road Banaue - Sagada (Fig. 3), where it was found close to a human settlement, under wooden plates and logs (Fig. 4).</paragraph>
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</subSubSection>
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</treatment>
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</document> |