treatments-xml/data/4A/69/E5/4A69E52BFFE6FFE3ACA5FD33FC82D0A9.xml

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<mods:title id="47A3FCFA65E3D8D1FA2BFCCCD0231E32">A new Trimerocephalus species (Trilobita, Phacopidae) from the Late Devonian (Early Famennian) of Poland</mods:title>
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<mods:namePart id="A528B2982804FA56CD0A09B304AE75CA">Żejowski, Ej Bła</mods:namePart>
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<treatment id="4A69E52BFFE6FFE3ACA5FD33FC82D0A9" ID-DOI="http://doi.org/10.5281/zenodo.5671405" ID-GBIF-Taxon="119563756" ID-Zenodo-Dep="5671405" LSID="urn:lsid:plazi:treatment:4A69E52BFFE6FFE3ACA5FD33FC82D0A9" httpUri="http://treatment.plazi.org/id/4A69E52BFFE6FFE3ACA5FD33FC82D0A9" lastPageId="7" lastPageNumber="352" pageId="2" pageNumber="347">
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<paragraph id="C27F543DFFE6FFE6ACA5FD33FDACD127" blockId="2.[151,528,752,812]" box="[151,528,752,779]" pageId="2" pageNumber="347">
<heading id="9937E351FFE6FFE6ACA5FD33FDACD127" bold="true" box="[151,528,752,779]" fontSize="11" level="1" pageId="2" pageNumber="347" reason="1">
<emphasis id="F0B4882FFFE6FFE6ACA5FD33FDACD127" bold="true" box="[151,528,752,779]" pageId="2" pageNumber="347">
<taxonomicName id="05C02FBEFFE6FFE6ACA5FD33FE7BD126" box="[151,455,753,779]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="347" phylum="Arthropoda" rank="species" species="chopini" status="sp. nov.">
<emphasis id="F0B4882FFFE6FFE6ACA5FD33FE7BD126" bold="true" box="[151,455,753,779]" italics="true" pageId="2" pageNumber="347">Trimerocephalus chopini</emphasis>
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<taxonomicNameLabel id="EB873554FFE6FFE6ADFCFD32FDACD127" box="[462,528,752,778]" pageId="2" pageNumber="347" rank="species">n. sp.</taxonomicNameLabel>
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<paragraph id="C27F543DFFE6FFE6ACA5FCD6FEA0D101" blockId="2.[151,528,752,812]" box="[151,284,788,812]" pageId="2" pageNumber="347">
<figureCitation id="5AFB48B8FFE6FFE6ACA5FCD6FEA0D101" box="[151,284,788,812]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4. A" captionStartId-0="3.[151,250,1861,1884]" captionStartId-1="4.[151,250,1852,1875]" captionStartId-2="5.[151,250,1280,1303]" captionTargetBox-0="[151,1435,193,1839]" captionTargetBox-1="[176,1398,193,1815]" captionTargetBox-2="[151,1435,193,1258]" captionTargetId-0="figure@3.[151,1436,193,1840]" captionTargetId-1="figure@4.[176,1400,193,1816]" captionTargetId-2="figure@5.[151,1436,193,1259]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Partially complete meraspis (degree M 10; librigenae absent) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A and B: Dorsal and oblique lateral views, respectively. C: Enlargement of pygidium." captionText-1="FIGURE 3. Meraspides (degrees M 8 - M 10) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M 8 in dorsal view (pygidium is absent). B: Almost complete meraspis M 9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M 8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M 10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M 10 (holotype; ZPAL Tr. 8 / 12.02.79) in dorsal view (pygidium absent)." captionText-2="FIGURE 4. A lmost complete internal mould of a holaspid (?) cephalon of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A – C: Dorsal, anterior and lateral views, respectively. D: Oblique ventral view showing vincular furrow. E: Details of weakly crenulated genal part of vincular furrow, with white arrows indicating shallow pits." httpUri-0="https://zenodo.org/record/218662/files/figure.png" httpUri-1="https://zenodo.org/record/218663/files/figure.png" httpUri-2="https://zenodo.org/record/218664/files/figure.png" pageId="2" pageNumber="347">Figures 2–4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="8ADA07B6FFE6FFE6ACA5FC99FB97D191" pageId="2" pageNumber="347" type="materials_examined">
<paragraph id="C27F543DFFE6FFE6ACA5FC99FB97D191" blockId="2.[151,1437,859,2036]" pageId="2" pageNumber="347">
<bibRef id="1B60C0CBFFE6FFE6ACA5FC99FB97D191" pageId="2" pageNumber="347" type="journal article">
<title id="B0C4BF0BFFE6FFE6ACA5FC99FF43D159" box="[151,255,859,884]" jopScore="0.013404588" pageId="2" pageNumber="347">
<emphasis id="F0B4882FFFE6FFE6ACA5FC99FF43D159" bold="true" box="[151,255,859,884]" pageId="2" pageNumber="347">Material</emphasis>
</title>
. 
<typeStatus id="1D7BEA9FFFE6FFE6AD3DFC9EFEC6D159" box="[271,378,860,884]" pageId="2" pageNumber="347" type="holotype">Holotype</typeStatus>
, 
<collectingCountry id="BAD714ADFFE6FFE6ADB5FC9EFE1FD159" box="[391,419,860,884]" name="Chile" pageId="2" pageNumber="347">Tr</collectingCountry>
.8/12.02.79, near complete exoskeleton (pygidium absent) of a late meraspid (M10) (
<figureCitation id="5AFB48B8FFE6FFE6A95DFC9EFF19D1BA" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1875]" captionTargetBox="[176,1398,193,1815]" captionTargetId="figure@4.[176,1400,193,1816]" captionTargetPageId="4" captionText="FIGURE 3. Meraspides (degrees M 8 - M 10) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M 8 in dorsal view (pygidium is absent). B: Almost complete meraspis M 9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M 8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M 10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M 10 (holotype; ZPAL Tr. 8 / 12.02.79) in dorsal view (pygidium absent)." httpUri="https://zenodo.org/record/218663/files/figure.png" pageId="2" pageNumber="347">Fig. 3</figureCitation>
F), from 
<author id="1A7DBCD8FFE6FFE6AD3DFCBDFEE8D1BA" box="[271,340,895,919]" fns="I" npo="LF" pageId="2" pageNumber="347">Unit I</author>
(late Early Famennian, 
<emphasis id="F0B4882FFFE6FFE6AE5FFC42FC2CD1B5" box="[621,912,896,920]" italics="true" pageId="2" pageNumber="347">Palmatolepis marginifera</emphasis>
Zone) in Kowala Quarry, 
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km
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southwest of Kielce, Holy Cross Mountains, central 
<collectingCountry id="BAD714ADFFE6FFE6AE7EFC66FD21D191" box="[588,669,932,956]" name="Poland" pageId="2" pageNumber="347">Poland</collectingCountry>
, 
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<pagination id="FB09AAC9FFE6FFE6AE9AFC66FD7AD191" box="[680,710,932,956]" pageId="2" pageNumber="347" type="pagination">50</pagination>
° 47' 46.21&quot; N
</geoCoordinate>
<geoCoordinate id="A7F432FAFFE6FFE6AF59FC66FB9BD191" box="[875,1063,932,956]" direction="east" orientation="longitude" pageId="2" pageNumber="347" precision="1" value="20.565977">20° 33' 57.52&quot; E</geoCoordinate>
.
</bibRef>
</paragraph>
</subSubSection>
<subSubSection id="8ADA07B6FFE6FFE6ACF5FC0AFC84D629" pageId="2" pageNumber="347" type="etymology">
<paragraph id="C27F543DFFE6FFE6ACF5FC0AFC84D629" blockId="2.[151,1437,859,2036]" pageId="2" pageNumber="347">
<emphasis id="F0B4882FFFE6FFE6ACF5FC0AFEFAD1CD" bold="true" box="[199,326,968,992]" pageId="2" pageNumber="347">Etymology</emphasis>
. In honour of Fryderyk Franciszek Chopin, the most famous Polish composer during the Romantic period of classical music, also called the ‘poet of the piano’.
</paragraph>
</subSubSection>
<subSubSection id="8ADA07B6FFE6FFE6ACF5FBCDFBF1D6F1" pageId="2" pageNumber="347" type="diagnosis">
<paragraph id="C27F543DFFE6FFE6ACF5FBCDFBF1D6F1" blockId="2.[151,1437,859,2036]" pageId="2" pageNumber="347">
<emphasis id="F0B4882FFFE6FFE6ACF5FBCDFE86D605" bold="true" box="[199,314,1039,1064]" pageId="2" pageNumber="347">Diagnosis</emphasis>
. Facial suture running along marginal furrow and not cutting cheek; the latter showing a very slight recess in contact with suture. Border furrows generally indistinct, but very shallow and wide near truncated genal angle. Ocular protuberance absent. Preoccipital furrow continuous along its length. Preoccipital ring straight, with two small, median tubercles. Occipital ring without median node. Pygidium trapezoidal with posterior margin almost rectilinear, width to length ratio of 2.8:1. Glabella and cheeks covered by very large to massive granules, usually uniform in size. Carapace covered by fine and moderately dense granulation.
</paragraph>
</subSubSection>
<subSubSection id="8ADA07B6FFE6FFE3ACF5FB25FC82D0A9" lastPageId="7" lastPageNumber="352" pageId="2" pageNumber="347" type="description">
<paragraph id="C27F543DFFE6FFE6ACF5FB25FE2DD4D5" blockId="2.[151,1437,859,2036]" pageId="2" pageNumber="347">
<emphasis id="F0B4882FFFE6FFE6ACF5FB25FEEDD72D" bold="true" box="[199,337,1255,1280]" pageId="2" pageNumber="347">Description</emphasis>
. Cephalon semicircular in outline, length to width ratio about 0.5. Glabella subpentagonal in outline, wide, rounded anteriorly with antero-lateral angles truncated (
<figureCitation id="5AFB48B8FFE6FFE6AF8CFACEFBB9D709" box="[958,1029,1292,1316]" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1875]" captionTargetBox="[176,1398,193,1815]" captionTargetId="figure@4.[176,1400,193,1816]" captionTargetPageId="4" captionText="FIGURE 3. Meraspides (degrees M 8 - M 10) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M 8 in dorsal view (pygidium is absent). B: Almost complete meraspis M 9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M 8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M 10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M 10 (holotype; ZPAL Tr. 8 / 12.02.79) in dorsal view (pygidium absent)." httpUri="https://zenodo.org/record/218663/files/figure.png" pageId="2" pageNumber="347">Figs 3</figureCitation>
F and 4A–C), length to width ratio about 0.6. Glabellar furrows S2 and S3 usually poorly developed, only visible on partially exposed internal moulds of glabellar area on three individuals [specimens: Kow / TA 76 (Radwański 
<emphasis id="F0B4882FFFE6FFE6AFCBFA97FB8ED741" box="[1017,1074,1364,1388]" italics="true" pageId="2" pageNumber="347">et al.</emphasis>
, 2009, pl. 2, fig. 3b) and Kow / TA 89, 90]; S2 short, bent obliquely upward and positioned very close to preoccipital furrow; posterior ramus of S3 very short and bent obliquely downward (as in 
<taxonomicName id="05C02FBEFFE6FFE6AE96FA5FFCB7D799" box="[676,779,1437,1460]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="2" pageNumber="347" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE6FFE6AE96FA5FFCB7D799" box="[676,779,1437,1460]" italics="true" pageId="2" pageNumber="347">T. caecus</emphasis>
</taxonomicName>
). Preoccipital furrow straight, well developed and of equal depth. Preoccipital ring straight, slightly convex with two small median tubercles. Occipital furrow slightly bent medially. Occipital ring wide, convex and without median node. Facial suture runs within the antero-lateral border furrow and does not extend onto the cheeks; cheeks exhibit a small recess in contact with suture (e.g., 
<figureCitation id="5AFB48B8FFE6FFE6A972F9C5FA3BD432" box="[1344,1415,1543,1567]" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1875]" captionTargetBox="[176,1398,193,1815]" captionTargetId="figure@4.[176,1400,193,1816]" captionTargetPageId="4" captionText="FIGURE 3. Meraspides (degrees M 8 - M 10) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M 8 in dorsal view (pygidium is absent). B: Almost complete meraspis M 9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M 8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M 10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M 10 (holotype; ZPAL Tr. 8 / 12.02.79) in dorsal view (pygidium absent)." httpUri="https://zenodo.org/record/218663/files/figure.png" pageId="2" pageNumber="347">Figs 3</figureCitation>
F, 4A–C). In dorsal view, the margins of the glabella and cheeks are highly vaulted; cheek areas triangular in outline, with rounded to truncated posterolateral genal angles. Axial furrows deep and moderately wide, diverging forwards at an angle of about 55°. Lateral and posterior borders are widest at posterolateral corner of cephalon. Vincular furrow in holaspid moderately deep, straight medially, strongly directed transversely with smooth edges (
<figureCitation id="5AFB48B8FFE6FFE6A95DF955FF1AD4F9" captionStart="FIGURE 4. A" captionStartId="5.[151,250,1280,1303]" captionTargetBox="[151,1435,193,1258]" captionTargetId="figure@5.[151,1436,193,1259]" captionTargetPageId="5" captionText="FIGURE 4. A lmost complete internal mould of a holaspid (?) cephalon of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A – C: Dorsal, anterior and lateral views, respectively. D: Oblique ventral view showing vincular furrow. E: Details of weakly crenulated genal part of vincular furrow, with white arrows indicating shallow pits." httpUri="https://zenodo.org/record/218664/files/figure.png" pageId="2" pageNumber="347">Fig. 4</figureCitation>
D–E). In the genal part it is directed posteriorly (parallel to the cheek), and is very slightly crenulated with small, shallow pits (
<figureCitation id="5AFB48B8FFE6FFE6AD1FF922FECCD4D5" box="[301,368,1760,1784]" captionStart="FIGURE 4. A" captionStartId="5.[151,250,1280,1303]" captionTargetBox="[151,1435,193,1258]" captionTargetId="figure@5.[151,1436,193,1259]" captionTargetPageId="5" captionText="FIGURE 4. A lmost complete internal mould of a holaspid (?) cephalon of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A – C: Dorsal, anterior and lateral views, respectively. D: Oblique ventral view showing vincular furrow. E: Details of weakly crenulated genal part of vincular furrow, with white arrows indicating shallow pits." httpUri="https://zenodo.org/record/218664/files/figure.png" pageId="2" pageNumber="347">Fig. 4</figureCitation>
E).
</paragraph>
<paragraph id="C27F543DFFE6FFE6ACF5F8C6FF56D549" blockId="2.[151,1437,859,2036]" pageId="2" pageNumber="347">
Thorax of 11 segments. Axial rings straight (tr.). Pleurae with flat fulcrum (bent backward) that is covered in fine granules. Pleural furrows wide and distinct, subdividing two convex bands. The anterior band is narrower (
<figureCitation id="5AFB48B8FFE6FFE6A95CF8EAFF19D549" captionStart="FIGURE 2" captionStartId="3.[151,250,1861,1884]" captionTargetBox="[151,1435,193,1839]" captionTargetId="figure@3.[151,1436,193,1840]" captionTargetPageId="3" captionText="FIGURE 2. Partially complete meraspis (degree M 10; librigenae absent) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A and B: Dorsal and oblique lateral views, respectively. C: Enlargement of pygidium." httpUri="https://zenodo.org/record/218662/files/figure.png" pageId="2" pageNumber="347">Fig. 2</figureCitation>
A–B).
</paragraph>
<paragraph id="C27F543DFFE6FFE6ACF5F8B2FDD1D5D9" blockId="2.[151,1437,859,2036]" pageId="2" pageNumber="347">
Pygidium almost three times as wide as long. Axial furrows moderately deep. Pygidial axis long and wide (
<figureCitation id="5AFB48B8FFE6FFE6A95CF8B2FF1AD581" captionStart="FIGURE 2" captionStartId="3.[151,250,1861,1884]" captionTargetBox="[151,1435,193,1839]" captionTargetId="figure@3.[151,1436,193,1840]" captionTargetPageId="3" captionText="FIGURE 2. Partially complete meraspis (degree M 10; librigenae absent) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A and B: Dorsal and oblique lateral views, respectively. C: Enlargement of pygidium." httpUri="https://zenodo.org/record/218662/files/figure.png" pageId="2" pageNumber="347">Fig. 2</figureCitation>
C). Ring furrows (5 + 1) straight, with anteriormost furrows more distinct than weakly developed posterior furrows. Articulating half ring as wide as first axial ring. Interpleural and pleural furrows (4) rather indistinct, except first two pleural furrows (
<figureCitation id="5AFB48B8FFE6FFE6AE35F81EFDF7D5D9" box="[519,587,2012,2036]" captionStart="FIGURE 2" captionStartId="3.[151,250,1861,1884]" captionTargetBox="[151,1435,193,1839]" captionTargetId="figure@3.[151,1436,193,1840]" captionTargetPageId="3" captionText="FIGURE 2. Partially complete meraspis (degree M 10; librigenae absent) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A and B: Dorsal and oblique lateral views, respectively. C: Enlargement of pygidium." httpUri="https://zenodo.org/record/218662/files/figure.png" pageId="2" pageNumber="347">Fig. 2</figureCitation>
C).
</paragraph>
<caption id="96BF04B5FFE7FFE7ACA5F887FDF9D5B4" httpUri="https://zenodo.org/record/218662/files/figure.png" pageId="3" pageNumber="348" targetBox="[151,1435,193,1839]" targetPageId="3">
<paragraph id="C27F543DFFE7FFE7ACA5F887FDF9D5B4" blockId="3.[151,1436,1861,1945]" pageId="3" pageNumber="348">
<emphasis id="F0B4882FFFE7FFE7ACA5F887FEA4D576" bold="true" box="[151,280,1861,1884]" pageId="3" pageNumber="348">FIGURE 2.</emphasis>
Partially complete meraspis (degree M10; librigenae absent) of 
<taxonomicName id="05C02FBEFFE7FFE7AFE3F884FB6FD571" box="[977,1235,1862,1884]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="3" pageNumber="348" phylum="Arthropoda" rank="species" species="chopini" status="sp. nov.">
<emphasis id="F0B4882FFFE7FFE7AFE3F884FB6FD571" box="[977,1235,1862,1884]" italics="true" pageId="3" pageNumber="348">Trimerocephalus chopini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB873554FFE7FFE7A8EDF884FAA1D576" box="[1247,1309,1862,1883]" pageId="3" pageNumber="348" rank="species">
<emphasis id="F0B4882FFFE7FFE7A8EDF884FAA1D576" bold="true" box="[1247,1309,1862,1883]" pageId="3" pageNumber="348">n. sp.</emphasis>
</taxonomicNameLabel>
from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A and B: Dorsal and oblique lateral views, respectively. C: Enlargement of pygidium.
</paragraph>
</caption>
<caption id="96BF04B5FFE0FFE0ACA5F8FEFC89D5C6" httpUri="https://zenodo.org/record/218663/files/figure.png" pageId="4" pageNumber="349" targetBox="[176,1398,193,1815]" targetPageId="4">
<paragraph id="C27F543DFFE0FFE0ACA5F8FEFC89D5C6" blockId="4.[151,1436,1852,2028]" pageId="4" pageNumber="349">
<emphasis id="F0B4882FFFE0FFE0ACA5F8FEFEAFD57C" bold="true" box="[151,275,1852,1875]" pageId="4" pageNumber="349">FIGURE 3.</emphasis>
Meraspides (degrees M8-M10) of 
<taxonomicName id="05C02FBEFFE0FFE0AE48F8FEFCCBD57F" box="[634,887,1852,1874]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="4" pageNumber="349" phylum="Arthropoda" rank="species" species="chopini" status="sp. nov.">
<emphasis id="F0B4882FFFE0FFE0AE48F8FEFCCBD57F" box="[634,887,1852,1874]" italics="true" pageId="4" pageNumber="349">Trimerocephalus chopini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB873554FFE0FFE0AF4CF8FFFC04D57F" box="[894,952,1853,1874]" pageId="4" pageNumber="349" rank="species">
<emphasis id="F0B4882FFFE0FFE0AF4CF8FFFC04D57F" bold="true" box="[894,952,1853,1874]" pageId="4" pageNumber="349">n. sp.</emphasis>
</taxonomicNameLabel>
from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M8 in dorsal view (pygidium is absent). B: Almost complete meraspis M9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M10 (holotype; ZPAL Tr.8/12.02.79) in dorsal view (pygidium absent).
</paragraph>
</caption>
<caption id="96BF04B5FFE1FFE1ACA5FAC2FDFCD75F" httpUri="https://zenodo.org/record/218664/files/figure.png" pageId="5" pageNumber="350" targetBox="[151,1435,193,1258]" targetPageId="5">
<paragraph id="C27F543DFFE1FFE1ACA5FAC2FDFCD75F" blockId="5.[151,1436,1280,1394]" pageId="5" pageNumber="350">
<emphasis id="F0B4882FFFE1FFE1ACA5FAC2FE8BD738" bold="true" box="[151,311,1280,1303]" pageId="5" pageNumber="350">FIGURE 4. A</emphasis>
lmost complete internal mould of a holaspid(?) cephalon of 
<taxonomicName id="05C02FBEFFE1FFE1AFFEFAC2FB6CD73B" box="[972,1232,1280,1302]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="5" pageNumber="350" phylum="Arthropoda" rank="species" species="chopini" status="sp. nov.">
<emphasis id="F0B4882FFFE1FFE1AFFEFAC2FB6CD73B" box="[972,1232,1280,1302]" italics="true" pageId="5" pageNumber="350">Trimerocephalus chopini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB873554FFE1FFE1A8EEFAC3FAA7D73B" box="[1244,1307,1281,1302]" pageId="5" pageNumber="350" rank="species">
<emphasis id="F0B4882FFFE1FFE1A8EEFAC3FAA7D73B" bold="true" box="[1244,1307,1281,1302]" pageId="5" pageNumber="350">n. sp.</emphasis>
</taxonomicNameLabel>
from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A–C: Dorsal, anterior and lateral views, respectively. D: Oblique ventral view showing vincular furrow. E: Details of weakly crenulated genal part of vincular furrow, with white arrows indicating shallow pits.
</paragraph>
</caption>
<paragraph id="C27F543DFFE1FFE1ACF5FA5BFD20D7F8" blockId="5.[151,1436,1433,1493]" pageId="5" pageNumber="350">
<emphasis id="F0B4882FFFE1FFE1ACF5FA5BFE47D79F" bold="true" box="[199,507,1433,1458]" pageId="5" pageNumber="350">Morphological variation.</emphasis>
Amongst studied representatives of 
<taxonomicName id="05C02FBEFFE1FFE1AFF6FA5BFB59D79C" box="[964,1253,1433,1457]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="5" pageNumber="350" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE1FFE1AFF6FA5BFB59D79C" box="[964,1253,1433,1457]" italics="true" pageId="5" pageNumber="350">Trimerocephalus chopini</emphasis>
</taxonomicName>
, the following morphological variations have been observed:
</paragraph>
<paragraph id="C27F543DFFE1FFE1ACA5F9C7FE9BD448" blockId="5.[151,1436,1541,1998]" pageId="5" pageNumber="350">1) In most cephala examined, the genal angle is truncated, but in some it is rounded. Specimens that exhibit the latter condition are typically flattened, thus may represent taphonomic (as opposed to true morphological) variants.</paragraph>
<paragraph id="C27F543DFFE1FFE1ACA5F9B3FD47D4D8" blockId="5.[151,1436,1541,1998]" pageId="5" pageNumber="350">
2) Most studied specimens do not show clear lateral glabellar furrows S2 and S3, which are visible only on internal moulds of three specimens. However, in other internal moulds these furrows are completely absent. Similar variation in the development of the glabellar furrows has been documented in 
<taxonomicName id="05C02FBEFFE1FFE1A871F97BFA20D4FC" box="[1091,1436,1721,1745]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="5" pageNumber="350" phylum="Arthropoda" rank="species" species="dianopsoides">
<emphasis id="F0B4882FFFE1FFE1A871F97BFA20D4FC" box="[1091,1436,1721,1745]" italics="true" pageId="5" pageNumber="350">Trimerocephalus dianopsoides</emphasis>
</taxonomicName>
from the Holy Cross Mountains (Osmólska 1963).
</paragraph>
<paragraph id="C27F543DFFE1FFE1ACA5F8C3FD6DD5E0" blockId="5.[151,1436,1541,1998]" pageId="5" pageNumber="350">
3) Cephalic tuberculation varies throughout ontogenesis. Relatively small granules occur in earlier meraspides (e.g., 
<figureCitation id="5AFB48B8FFE1FFE1AD3AF8E7FEF0D510" box="[264,332,1829,1853]" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1875]" captionTargetBox="[176,1398,193,1815]" captionTargetId="figure@4.[176,1400,193,1816]" captionTargetPageId="4" captionText="FIGURE 3. Meraspides (degrees M 8 - M 10) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M 8 in dorsal view (pygidium is absent). B: Almost complete meraspis M 9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M 8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M 10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M 10 (holotype; ZPAL Tr. 8 / 12.02.79) in dorsal view (pygidium absent)." httpUri="https://zenodo.org/record/218663/files/figure.png" pageId="5" pageNumber="350">Fig. 3</figureCitation>
B, C) and become gradually larger on cephala assigned to subsequent meraspides (i.e., sub-adult representatives, e.g., 
<figureCitation id="5AFB48B8FFE1FFE1ADF2F88BFDB7D54C" box="[448,523,1865,1889]" captionStart="FIGURE 2" captionStartId="3.[151,250,1861,1884]" captionTargetBox="[151,1435,193,1839]" captionTargetId="figure@3.[151,1436,193,1840]" captionTargetPageId="3" captionText="FIGURE 2. Partially complete meraspis (degree M 10; librigenae absent) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A and B: Dorsal and oblique lateral views, respectively. C: Enlargement of pygidium." httpUri="https://zenodo.org/record/218662/files/figure.png" pageId="5" pageNumber="350">Figs 2</figureCitation>
A, B, 3E, F). Slight differences in cephalic tuberculation (i.e., dimension of granules) also occur between individuals representing the same ontogenetic stages (compare specimens in 
<figureCitation id="5AFB48B8FFE1FFE1A942F8AFFF68D584" captionStart="FIGURE 3" captionStartId="4.[151,250,1852,1875]" captionTargetBox="[176,1398,193,1815]" captionTargetId="figure@4.[176,1400,193,1816]" captionTargetPageId="4" captionText="FIGURE 3. Meraspides (degrees M 8 - M 10) of Trimerocephalus chopini n. sp. from Early Famennian marly shales at Kowala Quarry, Holy Cross Mountains, central Poland. A: Incomplete, exfoliated, dorso-laterally deformed meraspis M 8 in dorsal view (pygidium is absent). B: Almost complete meraspis M 9 in dorsal view (pygidium is not intact). C: Complete, slightly deformed meraspis M 8 with partially visible pygidium, in dorsal view. D and E: Nearly complete, slightly deformed meraspis M 10 (librigenae absent) in oblique posterior and dorsal views, respectively. F: Incomplete and dorsally deformed meraspis M 10 (holotype; ZPAL Tr. 8 / 12.02.79) in dorsal view (pygidium absent)." httpUri="https://zenodo.org/record/218663/files/figure.png" pageId="5" pageNumber="350">Fig. 3</figureCitation>
E, F). The cephalic ornamentation in 
<taxonomicName id="05C02FBEFFE1FFE1AE49F850FD57D584" box="[635,747,1937,1961]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="5" pageNumber="350" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE1FFE1AE49F850FD57D584" box="[635,747,1937,1961]" italics="true" pageId="5" pageNumber="350">T. chopini</emphasis>
</taxonomicName>
closely resembles that of 
<taxonomicName id="05C02FBEFFE1FFE1A820F853FA27D584" authority="Feist" authorityName="Feist" box="[1042,1435,1937,1961]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="5" pageNumber="350" phylum="Arthropoda" rank="species" species="shotoriensis">
<emphasis id="F0B4882FFFE1FFE1A820F853FAE1D584" box="[1042,1373,1937,1961]" italics="true" pageId="5" pageNumber="350">Trimerocephalus shotoriensis</emphasis>
Feist
</taxonomicName>
<emphasis id="F0B4882FFFE1FFE1ACF5F875FF61D5E3" box="[199,221,1975,1998]" italics="true" pageId="5" pageNumber="350">in</emphasis>
Feist, Yazdi and Becker, 2003 (see 
<tableCitation id="8F426186FFE1FFE1AE5CF877FD03D5E0" box="[622,703,1973,1997]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="5" pageNumber="350" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
).
</paragraph>
<caption id="96BF04B5FFE2FFE2ACA5FF5AFC9ED366" ID-Table-UUID="96BF04B5FFE2FFE2ACA5FF5AFC9ED366" httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" targetBox="[159,1423,361,672]" targetIsTable="true" targetPageId="6">
<paragraph id="C27F543DFFE2FFE2ACA5FF5AFC9ED366" blockId="6.[151,1436,152,332]" pageId="6" pageNumber="351">
<emphasis id="F0B4882FFFE2FFE2ACA5FF5AFEB8D280" bold="true" box="[151,260,152,173]" pageId="6" pageNumber="351">TABLE 1.</emphasis>
Comparison of nine morphological features between 
<taxonomicName id="05C02FBEFFE2FFE2AF16FF5AFB9DD283" box="[804,1057,152,174]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="chopini" status="sp. nov.">
<emphasis id="F0B4882FFFE2FFE2AF16FF5AFB9DD283" box="[804,1057,152,174]" italics="true" pageId="6" pageNumber="351">Trimerocephalus chopini</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EB873554FFE2FFE2A815FF5AFBDCD280" box="[1063,1120,152,173]" pageId="6" pageNumber="351" rank="species">
<emphasis id="F0B4882FFFE2FFE2A815FF5AFBDCD280" bold="true" box="[1063,1120,152,173]" pageId="6" pageNumber="351">n. sp.</emphasis>
</taxonomicNameLabel>
and selected representatives of the genus 
<taxonomicName id="05C02FBEFFE2FFE2AD38FF75FE09D2E0" box="[266,437,183,205]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="F0B4882FFFE2FFE2AD38FF75FE09D2E0" box="[266,437,183,205]" italics="true" pageId="6" pageNumber="351">Trimerocephalus</emphasis>
</taxonomicName>
. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present.
</paragraph>
</caption>
<paragraph id="C27F543DFFE2FFE2ACADFEA8FA84D08D" pageId="6" pageNumber="351">
<table id="B0C0A69DFFE2001BACADFEABFA33D08D" box="[159,1423,361,672]" gridcols="10" gridrows="8" pageId="6" pageNumber="351">
<tr id="7CF0567FFFE2001BACADFEABFA33D352" box="[159,1423,361,383]" gridrow="0" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFEABFEEDD352" box="[159,337,361,383]" gridcol="0" gridrow="0" pageId="6" pageNumber="351">Taxa</th>
<th id="3F213F03FFE2001BAD5FFEABFE70D352" box="[365,460,361,383]" gridcol="1" gridrow="0" pageId="6" pageNumber="351">Feature 1</th>
<th id="3F213F03FFE2001BADDEFEABFDF7D352" box="[492,587,361,383]" gridcol="2" gridrow="0" pageId="6" pageNumber="351">Feature 2</th>
<th id="3F213F03FFE2001BAE51FEABFD7ED352" box="[611,706,361,383]" gridcol="3" gridrow="0" pageId="6" pageNumber="351">Feature 3</th>
<th id="3F213F03FFE2001BAEE9FEABFC86D352" box="[731,826,361,383]" gridcol="4" gridrow="0" pageId="6" pageNumber="351">Feature 4</th>
<th id="3F213F03FFE2001BAF60FEABFC0DD352" box="[850,945,361,383]" gridcol="5" gridrow="0" pageId="6" pageNumber="351">Feature 5</th>
<th id="3F213F03FFE2001BAFFBFEABFB95D352" box="[969,1065,361,383]" gridcol="6" gridrow="0" pageId="6" pageNumber="351">Feature 6</th>
<th id="3F213F03FFE2001BA873FEABFB1CD352" box="[1089,1184,361,383]" gridcol="7" gridrow="0" pageId="6" pageNumber="351">Feature 7</th>
<th id="3F213F03FFE2001BA88AFEABFAA4D352" box="[1208,1304,361,383]" gridcol="8" gridrow="0" pageId="6" pageNumber="351">Feature 8</th>
<th id="3F213F03FFE2001BA902FEABFA33D352" box="[1328,1423,361,383]" gridcol="9" gridrow="0" pageId="6" pageNumber="351">Feature 9</th>
</tr>
<tr id="7CF0567FFFE2001BACADFE50FA33D385" box="[159,1423,402,424]" gridrow="1" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFE50FEEDD385" box="[159,337,402,424]" gridcol="0" gridrow="1" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFE51FEBED385" box="[159,258,402,424]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE2FFE2ACADFE51FEBED385" box="[159,258,402,424]" italics="true" pageId="6" pageNumber="351">T. chopini</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFE50FE70D385" box="[365,460,402,424]" gridcol="1" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BADDEFE50FDF7D385" box="[492,587,402,424]" gridcol="2" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAE51FE50FD7ED385" box="[611,706,402,424]" gridcol="3" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAEE9FE50FC86D385" box="[731,826,402,424]" gridcol="4" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAF60FE50FC0DD385" box="[850,945,402,424]" gridcol="5" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAFFBFE50FB95D385" box="[969,1065,402,424]" gridcol="6" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA873FE50FB1CD385" box="[1089,1184,402,424]" gridcol="7" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA88AFE50FAA4D385" box="[1208,1304,402,424]" gridcol="8" gridrow="1" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA902FE50FA33D385" box="[1328,1423,402,424]" gridcol="9" gridrow="1" pageId="6" pageNumber="351">+</td>
</tr>
<tr id="7CF0567FFFE2001BACADFE7FFA33D3FF" box="[159,1423,445,466]" gridrow="2" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFE7FFEEDD3FF" box="[159,337,445,466]" gridcol="0" gridrow="2" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFE7FFF41D3FF" box="[159,253,445,466]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE2FFE2ACADFE7FFF41D3FF" box="[159,253,445,466]" italics="true" pageId="6" pageNumber="351">T. caecus</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFE7FFE70D3FF" box="[365,460,445,466]" gridcol="1" gridrow="2" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BADDEFE7FFDF7D3FF" box="[492,587,445,466]" gridcol="2" gridrow="2" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAE51FE7FFD7ED3FF" box="[611,706,445,466]" gridcol="3" gridrow="2" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAEE9FE7FFC86D3FF" box="[731,826,445,466]" gridcol="4" gridrow="2" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAF60FE7FFC0DD3FF" box="[850,945,445,466]" gridcol="5" gridrow="2" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAFFBFE7FFB95D3FF" box="[969,1065,445,466]" gridcol="6" gridrow="2" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA873FE7FFB1CD3FF" box="[1089,1184,445,466]" gridcol="7" gridrow="2" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA88AFE7FFAA4D3FF" box="[1208,1304,445,466]" gridcol="8" gridrow="2" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA902FE7FFA33D3FF" box="[1328,1423,445,466]" gridcol="9" gridrow="2" pageId="6" pageNumber="351">-</td>
</tr>
<tr id="7CF0567FFFE2001BACADFE27FA33D3D6" box="[159,1423,485,507]" gridrow="3" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFE27FEEDD3D6" box="[159,337,485,507]" gridcol="0" gridrow="3" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFE24FEB6D3D6" box="[159,266,485,507]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="lelievrei">
<emphasis id="F0B4882FFFE2FFE2ACADFE24FEB6D3D6" box="[159,266,485,507]" italics="true" pageId="6" pageNumber="351">T. lelievrei</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFE27FE70D3D6" box="[365,460,485,507]" gridcol="1" gridrow="3" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BADDEFE27FDF7D3D6" box="[492,587,485,507]" gridcol="2" gridrow="3" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAE51FE27FD7ED3D6" box="[611,706,485,507]" gridcol="3" gridrow="3" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAEE9FE27FC86D3D6" box="[731,826,485,507]" gridcol="4" gridrow="3" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAF60FE27FC0DD3D6" box="[850,945,485,507]" gridcol="5" gridrow="3" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAFFBFE27FB95D3D6" box="[969,1065,485,507]" gridcol="6" gridrow="3" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA873FE27FB1CD3D6" box="[1089,1184,485,507]" gridcol="7" gridrow="3" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA88AFE27FAA4D3D6" box="[1208,1304,485,507]" gridcol="8" gridrow="3" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA902FE27FA33D3D6" box="[1328,1423,485,507]" gridcol="9" gridrow="3" pageId="6" pageNumber="351">-</td>
</tr>
<tr id="7CF0567FFFE2001BACADFDCDFA33D008" box="[159,1423,527,549]" gridrow="4" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFDCDFEEDD008" box="[159,337,527,549]" gridcol="0" gridrow="4" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFDD2FE82D008" box="[159,318,527,549]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="dianopsoides">
<emphasis id="F0B4882FFFE2FFE2ACADFDD2FE82D008" box="[159,318,527,549]" italics="true" pageId="6" pageNumber="351">T. dianopsoides</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFDCDFE70D008" box="[365,460,527,549]" gridcol="1" gridrow="4" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BADDEFDCDFDF7D008" box="[492,587,527,549]" gridcol="2" gridrow="4" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAE51FDCDFD7ED008" box="[611,706,527,549]" gridcol="3" gridrow="4" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAEE9FDCDFC86D008" box="[731,826,527,549]" gridcol="4" gridrow="4" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAF60FDCDFC0DD008" box="[850,945,527,549]" gridcol="5" gridrow="4" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAFFBFDCDFB95D008" box="[969,1065,527,549]" gridcol="6" gridrow="4" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA873FDCDFB1CD008" box="[1089,1184,527,549]" gridcol="7" gridrow="4" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA88AFDCDFAA4D008" box="[1208,1304,527,549]" gridcol="8" gridrow="4" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA902FDCDFA33D008" box="[1328,1423,527,549]" gridcol="9" gridrow="4" pageId="6" pageNumber="351">-</td>
</tr>
<tr id="7CF0567FFFE2001BACADFDFAFA33D063" box="[159,1423,568,590]" gridrow="5" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFDFAFEEDD063" box="[159,337,568,590]" gridcol="0" gridrow="5" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFDFBFEEDD063" box="[159,337,568,590]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mastophtalmus">
<emphasis id="F0B4882FFFE2FFE2ACADFDFBFEEDD063" box="[159,337,568,590]" italics="true" pageId="6" pageNumber="351">T. mastophtalmus</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFDFAFE70D063" box="[365,460,568,590]" gridcol="1" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BADDEFDFAFDF7D063" box="[492,587,568,590]" gridcol="2" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAE51FDFAFD7ED063" box="[611,706,568,590]" gridcol="3" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAEE9FDFAFC86D063" box="[731,826,568,590]" gridcol="4" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAF60FDFAFC0DD063" box="[850,945,568,590]" gridcol="5" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAFFBFDFAFB95D063" box="[969,1065,568,590]" gridcol="6" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA873FDFAFB1CD063" box="[1089,1184,568,590]" gridcol="7" gridrow="5" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA88AFDFAFAA4D063" box="[1208,1304,568,590]" gridcol="8" gridrow="5" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA902FDFAFA33D063" box="[1328,1423,568,590]" gridcol="9" gridrow="5" pageId="6" pageNumber="351">-</td>
</tr>
<tr id="7CF0567FFFE2001BACADFDA3FA33D05A" box="[159,1423,609,631]" gridrow="6" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFDA3FEEDD05A" box="[159,337,609,631]" gridcol="0" gridrow="6" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFDA0FF4BD05A" box="[159,247,609,631]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mimbi">
<emphasis id="F0B4882FFFE2FFE2ACADFDA0FF4BD05A" box="[159,247,609,631]" italics="true" pageId="6" pageNumber="351">T. mimbi</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFDA3FE70D05A" box="[365,460,609,631]" gridcol="1" gridrow="6" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BADDEFDA3FDF7D05A" box="[492,587,609,631]" gridcol="2" gridrow="6" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAE51FDA3FD7ED05A" box="[611,706,609,631]" gridcol="3" gridrow="6" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAEE9FDA3FC86D05A" box="[731,826,609,631]" gridcol="4" gridrow="6" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAF60FDA3FC0DD05A" box="[850,945,609,631]" gridcol="5" gridrow="6" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAFFBFDA3FB95D05A" box="[969,1065,609,631]" gridcol="6" gridrow="6" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA873FDA3FB1CD05A" box="[1089,1184,609,631]" gridcol="7" gridrow="6" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA88AFDA3FAA4D05A" box="[1208,1304,609,631]" gridcol="8" gridrow="6" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA902FDA3FA33D05A" box="[1328,1423,609,631]" gridcol="9" gridrow="6" pageId="6" pageNumber="351">-</td>
</tr>
<tr id="7CF0567FFFE2001BACADFD48FA33D08D" box="[159,1423,650,672]" gridrow="7" pageId="6" pageNumber="351">
<th id="3F213F03FFE2001BACADFD48FEEDD08D" box="[159,337,650,672]" gridcol="0" gridrow="7" pageId="6" pageNumber="351">
<taxonomicName id="05C02FBEFFE2FFE2ACADFD49FE8ED08D" box="[159,306,650,672]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="shotoriensis">
<emphasis id="F0B4882FFFE2FFE2ACADFD49FE8ED08D" box="[159,306,650,672]" italics="true" pageId="6" pageNumber="351">T. shotoriensis</emphasis>
</taxonomicName>
</th>
<td id="3F213F03FFE2001BAD5FFD48FE70D08D" box="[365,460,650,672]" gridcol="1" gridrow="7" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BADDEFD48FDF7D08D" box="[492,587,650,672]" gridcol="2" gridrow="7" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAE51FD48FD7ED08D" box="[611,706,650,672]" gridcol="3" gridrow="7" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAEE9FD48FC86D08D" box="[731,826,650,672]" gridcol="4" gridrow="7" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BAF60FD48FC0DD08D" box="[850,945,650,672]" gridcol="5" gridrow="7" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BAFFBFD48FB95D08D" box="[969,1065,650,672]" gridcol="6" gridrow="7" pageId="6" pageNumber="351">+</td>
<td id="3F213F03FFE2001BA873FD48FB1CD08D" box="[1089,1184,650,672]" gridcol="7" gridrow="7" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA88AFD48FAA4D08D" box="[1208,1304,650,672]" gridcol="8" gridrow="7" pageId="6" pageNumber="351">-</td>
<td id="3F213F03FFE2001BA902FD48FA33D08D" box="[1328,1423,650,672]" gridcol="9" gridrow="7" pageId="6" pageNumber="351">-</td>
</tr>
</table>
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACF5FD27FC7AD19C" blockId="6.[151,1436,741,1305]" pageId="6" pageNumber="351">
<emphasis id="F0B4882FFFE2FFE2ACF5FD27FCC8D0D3" bold="true" box="[199,884,741,766]" pageId="6" pageNumber="351">
Comparison of 
<taxonomicName id="05C02FBEFFE2FFE2ADB6FD27FD14D0D3" box="[388,680,741,766]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE2FFE2ADB6FD27FD14D0D3" bold="true" box="[388,680,741,766]" italics="true" pageId="6" pageNumber="351">Trimerocephalus chopini</emphasis>
</taxonomicName>
to other species.
</emphasis>
For the purposes of this comparison, we follow Crônier’s (2003) phylogenetic groupings for the better known representatives of 
<taxonomicName id="05C02FBEFFE2FFE2A82FFCCBFB65D10C" box="[1053,1241,777,801]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="F0B4882FFFE2FFE2A82FFCCBFB65D10C" box="[1053,1241,777,801]" italics="true" pageId="6" pageNumber="351">Trimerocephalus</emphasis>
</taxonomicName>
. Accordingly, on the basis of 23 morphological characters, three clades were distinguished among early Famennian representatives of this genus: 
<taxonomicName id="05C02FBEFFE2FFE2AD0AFC90FE1DD144" box="[312,417,850,873]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE2FFE2AD0AFC90FE1DD144" box="[312,417,850,873]" italics="true" pageId="6" pageNumber="351">T. caecus</emphasis>
</taxonomicName>
, 
<taxonomicName id="05C02FBEFFE2FFE2AD82FC90FD94D144" box="[432,552,849,873]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="lelievrei">
<emphasis id="F0B4882FFFE2FFE2AD82FC90FD94D144" box="[432,552,849,873]" italics="true" pageId="6" pageNumber="351">T. lelievrei</emphasis>
</taxonomicName>
, 
<taxonomicName id="05C02FBEFFE2FFE2AE05FC90FD60D144" box="[567,732,849,873]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="shotoriensis">
<emphasis id="F0B4882FFFE2FFE2AE05FC90FD60D144" box="[567,732,849,873]" italics="true" pageId="6" pageNumber="351">T. shotoriensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="05C02FBEFFE2FFE2AF26FC90FC7BD144" box="[788,967,849,873]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="tardispinosus">
<emphasis id="F0B4882FFFE2FFE2AF26FC90FC7BD144" box="[788,967,849,873]" italics="true" pageId="6" pageNumber="351">T. tardispinosus</emphasis>
</taxonomicName>
[clade 1]; 
<taxonomicName id="05C02FBEFFE2FFE2A871FC90FB73D144" box="[1091,1231,850,873]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="procurvus">
<emphasis id="F0B4882FFFE2FFE2A871FC90FB73D144" box="[1091,1231,850,873]" italics="true" pageId="6" pageNumber="351">T. procurvus</emphasis>
</taxonomicName>
, 
<taxonomicName id="05C02FBEFFE2FFE2A8EDFC90FAC4D144" box="[1247,1400,850,873]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="F0B4882FFFE2FFE2A8EDFC90FAC4D144" box="[1247,1400,850,873]" italics="true" pageId="6" pageNumber="351">T. interruptus</emphasis>
</taxonomicName>
, 
<taxonomicName id="05C02FBEFFE2FFE2A9B5FC90FF4CD1A0" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="sponsor">
<emphasis id="F0B4882FFFE2FFE2A9B5FC90FF4CD1A0" italics="true" pageId="6" pageNumber="351">T. sponsor</emphasis>
</taxonomicName>
and 
<taxonomicName id="05C02FBEFFE2FFE2AD00FCB4FE7CD1A0" box="[306,448,885,909]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="F0B4882FFFE2FFE2AD00FCB4FE7CD1A0" box="[306,448,885,909]" italics="true" pageId="6" pageNumber="351">T. polonicus</emphasis>
</taxonomicName>
[clade 2]; and 
<taxonomicName id="05C02FBEFFE2FFE2AE4DFCB4FC88D1A0" box="[639,820,885,909]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="dianopsoides">
<emphasis id="F0B4882FFFE2FFE2AE4DFCB4FC88D1A0" box="[639,820,885,909]" italics="true" pageId="6" pageNumber="351">T. dianopsoides</emphasis>
</taxonomicName>
, 
<taxonomicName id="05C02FBEFFE2FFE2AF7AFCB4FBF1D1A0" box="[840,1101,885,909]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="nigritus" subGenus="Trifoliops">
<emphasis id="F0B4882FFFE2FFE2AF7AFCB4FCE7D1A0" box="[840,859,886,909]" italics="true" pageId="6" pageNumber="351">T.</emphasis>
(
<emphasis id="F0B4882FFFE2FFE2AF41FCB7FC5CD1A0" box="[883,992,885,909]" italics="true" pageId="6" pageNumber="351">Trifoliops</emphasis>
) 
<emphasis id="F0B4882FFFE2FFE2AFC4FCB4FBF1D1A0" box="[1014,1101,886,909]" italics="true" pageId="6" pageNumber="351">nigritus</emphasis>
</taxonomicName>
, 
<taxonomicName id="05C02FBEFFE2FFE2A853FCB4FADBD1A0" box="[1121,1383,885,909]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="trifolius" subGenus="Trifoliops">
<emphasis id="F0B4882FFFE2FFE2A853FCB4FBD2D1A0" box="[1121,1134,886,909]" italics="true" pageId="6" pageNumber="351">T</emphasis>
. (
<emphasis id="F0B4882FFFE2FFE2A8B9FCB7FB44D1A0" box="[1163,1272,885,909]" italics="true" pageId="6" pageNumber="351">Trifoliops</emphasis>
) 
<emphasis id="F0B4882FFFE2FFE2A93CFCB7FADBD1A0" box="[1294,1383,885,909]" italics="true" pageId="6" pageNumber="351">trifolius</emphasis>
</taxonomicName>
, 
<emphasis id="F0B4882FFFE2FFE2A949FCB4FA34D1A0" box="[1403,1416,886,909]" italics="true" pageId="6" pageNumber="351">T</emphasis>
.? 
<emphasis id="F0B4882FFFE2FFE2ACA5FC5BFE90D19C" box="[151,300,921,945]" italics="true" pageId="6" pageNumber="351">steinachensis</emphasis>
and 
<taxonomicName id="05C02FBEFFE2FFE2AD51FC58FD89D19C" box="[355,565,921,945]" class="Trilobita" family="Phacopidae" genus="Phacops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mastophthalmus">
<emphasis id="F0B4882FFFE2FFE2AD51FC58FECCD19C" box="[355,368,922,945]" italics="true" pageId="6" pageNumber="351">T</emphasis>
. 
<emphasis id="F0B4882FFFE2FFE2AD4CFC5BFD89D19C" box="[382,565,921,945]" italics="true" pageId="6" pageNumber="351">mastophthalmus</emphasis>
</taxonomicName>
[clade 3] (cf. Crônier, 2003, fig. 8).
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACF5FC7CFD65D680" blockId="6.[151,1436,741,1305]" pageId="6" pageNumber="351">
Key morphological features of 
<taxonomicName id="05C02FBEFFE2FFE2AE16FC7FFC80D1F8" box="[548,828,957,981]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE2FFE2AE16FC7FFC80D1F8" box="[548,828,957,981]" italics="true" pageId="6" pageNumber="351">Trimerocephalus chopini</emphasis>
</taxonomicName>
that have phylogenetic significance include (
<tableCitation id="8F426186FFE2FFE2A904FC7FFA36D1F8" box="[1334,1418,957,981]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
): (1) facial sutures that do not cut the cheeks; (2) a well developed, straight preoccipital furrow, S1; (3) a lack of ocular protuberances; (4) coarse cephalic tuberculation; (5) a trapezoidal pygidium; (6) a preoccipital ring with two median tubercles; (7) the absence of an occipital median node; (8) a slight recess in the cheeks in association with the facial suture; and (9) a truncated genal angle. These features suggest a close relationship to members of clade 1 (mentioned above), in addition to the recently erected 
<taxonomicName id="05C02FBEFFE2FFE2AF39FBB0FCD3D6A4" box="[779,879,1137,1161]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mimbi">
<emphasis id="F0B4882FFFE2FFE2AF39FBB0FCD3D6A4" box="[779,879,1137,1161]" italics="true" pageId="6" pageNumber="351">T. mimbi</emphasis>
</taxonomicName>
from the Early Famennian (do II = 
<emphasis id="F0B4882FFFE2FFE2A927FBB3FA20D6A4" box="[1301,1436,1137,1161]" italics="true" pageId="6" pageNumber="351">rhomboidea</emphasis>
Zone) of 
<collectingCountry id="BAD714ADFFE2FFE2ACCDFB57FED9D680" box="[255,357,1173,1197]" name="Australia" pageId="6" pageNumber="351">Australia</collectingCountry>
(Feist 
<emphasis id="F0B4882FFFE2FFE2AD81FB54FE57D680" box="[435,491,1173,1197]" italics="true" pageId="6" pageNumber="351">et al.</emphasis>
, 2009) (see 
<tableCitation id="8F426186FFE2FFE2AE44FB57FD74D680" box="[630,712,1173,1197]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
).
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACF5FB7BFDBBD734" blockId="6.[151,1436,741,1305]" pageId="6" pageNumber="351">
The following list outlines the main morphological differences of Early Famennian species of 
<taxonomicName id="05C02FBEFFE2FFE2A8EDFB7BFA20D6FC" box="[1247,1436,1209,1233]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="genus">
<emphasis id="F0B4882FFFE2FFE2A8EDFB7BFA20D6FC" box="[1247,1436,1209,1233]" italics="true" pageId="6" pageNumber="351">Trimerocephalus</emphasis>
</taxonomicName>
from 
<collectingCountry id="BAD714ADFFE2FFE2ACE6FB1FFE9ED6D8" box="[212,290,1245,1269]" name="Poland" pageId="6" pageNumber="351">Poland</collectingCountry>
(mentioned above), 
<collectingCountry id="BAD714ADFFE2FFE2AE35FB1CFD88D6D8" box="[519,564,1246,1269]" name="Iran" pageId="6" pageNumber="351">Iran</collectingCountry>
(
<taxonomicName id="05C02FBEFFE2FFE2AE76FB1CFD5AD6D8" box="[580,742,1245,1269]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="shotoriensis">
<emphasis id="F0B4882FFFE2FFE2AE76FB1CFD5AD6D8" box="[580,742,1245,1269]" italics="true" pageId="6" pageNumber="351">T. shotoriensis</emphasis>
</taxonomicName>
; Feist 
<emphasis id="F0B4882FFFE2FFE2AF03FB1CFCD5D6D8" box="[817,873,1245,1269]" italics="true" pageId="6" pageNumber="351">et al.</emphasis>
, 2003) and 
<collectingCountry id="BAD714ADFFE2FFE2AFDEFB1FFBEFD6D8" box="[1004,1107,1245,1269]" name="Australia" pageId="6" pageNumber="351">Australia</collectingCountry>
(
<taxonomicName id="05C02FBEFFE2FFE2A851FB1CFB78D6D8" box="[1123,1220,1245,1269]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mimbi">
<emphasis id="F0B4882FFFE2FFE2A851FB1CFB78D6D8" box="[1123,1220,1245,1269]" italics="true" pageId="6" pageNumber="351">T. mimbi</emphasis>
</taxonomicName>
; Feist 
<emphasis id="F0B4882FFFE2FFE2A93DFB1CFAFAD6D8" box="[1295,1350,1245,1269]" italics="true" pageId="6" pageNumber="351">et al.</emphasis>
, 2009), and the new species (
<taxonomicName id="05C02FBEFFE2FFE2ADB4FAC0FE4AD734" box="[390,502,1281,1305]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE2FFE2ADB4FAC0FE4AD734" box="[390,502,1281,1305]" italics="true" pageId="6" pageNumber="351">T. chopini</emphasis>
</taxonomicName>
):
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5FA88FD65D7A8" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5FA8BFE6BD74C" box="[199,471,1353,1377]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE2FFE2ACF5FA8BFE6BD74C" box="[199,471,1353,1377]" italics="true" pageId="6" pageNumber="351">Trimerocephalus caecus</emphasis>
</taxonomicName>
differs in its anteriorly curved preoccipital furrow (S1), rounded genal angle, and the presence of an occipital median node (
<tableCitation id="8F426186FFE2FFE2AE41FAAFFD7BD7A8" box="[627,711,1389,1413]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
);
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5FA50FE18D7E0" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5FA53FDA2D784" box="[199,542,1425,1449]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="dianopsoides">
<emphasis id="F0B4882FFFE2FFE2ACF5FA53FDA2D784" box="[199,542,1425,1449]" italics="true" pageId="6" pageNumber="351">Trimerocephalus dianopsoides</emphasis>
</taxonomicName>
differs in the presence of ocular protuberances and the poor segmentation of the pygidium (
<tableCitation id="8F426186FFE2FFE2AD0CFA77FE2ED7E0" box="[318,402,1461,1485]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
);
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5FA18FC0DD438" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5FA1BFDB2D7DC" box="[199,526,1497,1521]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="F0B4882FFFE2FFE2ACF5FA1BFDB2D7DC" box="[199,526,1497,1521]" italics="true" pageId="6" pageNumber="351">Trimerocephalus interruptus</emphasis>
</taxonomicName>
differs in the outline of the facial suture that cuts the cheek, the fine and condensed granulation, and poorly segmented lenticular pygidium;
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5F9E0FBA8D470" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5F9E3FDFED414" box="[199,578,1569,1593]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mastophthalmus">
<emphasis id="F0B4882FFFE2FFE2ACF5F9E3FDFED414" box="[199,578,1569,1593]" italics="true" pageId="6" pageNumber="351">Trimerocephalus mastophthalmus</emphasis>
</taxonomicName>
differs in the outline of the facial suture that cuts the cheek, the presence of ocular protuberances, and much finer granulation of the cephalon (
<tableCitation id="8F426186FFE2FFE2AF9DF987FBBFD470" box="[943,1027,1605,1629]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
);
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5F9A8FC51D488" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
<emphasis id="F0B4882FFFE2FFE2ACA5F9A8FE6DD4AC" box="[151,465,1641,1665]" italics="true" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5F9ABFE6DD4AC" box="[199,465,1641,1665]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="mimbi">Trimerocephalus mimbi</taxonomicName>
</emphasis>
differs in having a medially ill-defined preoccipital furrow (S1), rounded genal angle, a deep lateral border furrow and much deeper vincular furrow (
<tableCitation id="8F426186FFE2FFE2AFBAF94FFC67D488" box="[904,987,1677,1701]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
);
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5F970FEDCD53C" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5F973FE40D4E4" box="[199,508,1713,1737]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="polonicus">
<emphasis id="F0B4882FFFE2FFE2ACF5F973FE40D4E4" box="[199,508,1713,1737]" italics="true" pageId="6" pageNumber="351">Trimerocephalus polonicus</emphasis>
</taxonomicName>
is represented only by cephala of early instars, but differences include a subsemicircular outline (but strongly pointed anteriorly), a facial suture that cuts the cheeks, and very sparse tuberculation;
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5F8DCFD4DD574" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5F8DFFDD6D518" box="[199,618,1821,1845]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="trifolius" subGenus="Trifoliops">
<emphasis id="F0B4882FFFE2FFE2ACF5F8DFFE38D518" box="[199,388,1821,1845]" italics="true" pageId="6" pageNumber="351">Trimerocephalus</emphasis>
(
<emphasis id="F0B4882FFFE2FFE2ADA6F8DFFDBDD518" box="[404,513,1821,1845]" italics="true" pageId="6" pageNumber="351">Trifoliops</emphasis>
) 
<emphasis id="F0B4882FFFE2FFE2AE23F8DFFDD6D518" box="[529,618,1821,1845]" italics="true" pageId="6" pageNumber="351">trifolius</emphasis>
</taxonomicName>
differs in the trilobed outline of the cephalon, a facial suture that cuts the cheeks, and the presence of ocular protuberances;
</paragraph>
<paragraph id="C27F543DFFE2FFE2ACA5F8A4FE1FD58C" blockId="6.[151,1436,1353,1953]" pageId="6" pageNumber="351">
- 
<taxonomicName id="05C02FBEFFE2FFE2ACF5F8A7FDA8D550" box="[199,532,1893,1917]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="6" pageNumber="351" phylum="Arthropoda" rank="species" species="shotoriensis">
<emphasis id="F0B4882FFFE2FFE2ACF5F8A7FDA8D550" box="[199,532,1893,1917]" italics="true" pageId="6" pageNumber="351">Trimerocephalus shotoriensis</emphasis>
</taxonomicName>
differs in exhibiting a rounded genal angle and a subtrapezoidal outline of the pygidium (
<tableCitation id="8F426186FFE2FFE2AD0CF84BFE2ED58C" box="[318,402,1929,1953]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="6" pageNumber="351" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
).
</paragraph>
<paragraph id="C27F543DFFE3FFE3ACF5FF55FC82D0A9" blockId="7.[151,1436,151,644]" pageId="7" pageNumber="352">
Crônier (2003) emphasised the acquisition of a marginal facial suture that does not cut the cheeks and the presence of an occipital median node as important morphological features in tracing the evolutionary trends in 
<taxonomicName id="05C02FBEFFE3FFE3ACA5FF22FEE8D2D5" box="[151,340,224,248]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="genus">
<emphasis id="F0B4882FFFE3FFE3ACA5FF22FEE8D2D5" box="[151,340,224,248]" italics="true" pageId="7" pageNumber="352">Trimerocephalus</emphasis>
</taxonomicName>
, despite being homoplastic. 
<taxonomicName id="05C02FBEFFE3FFE3AEABFF22FC0ED2D5" box="[665,946,224,248]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE3FFE3AEABFF22FC0ED2D5" box="[665,946,224,248]" italics="true" pageId="7" pageNumber="352">Trimerocephalus chopini</emphasis>
</taxonomicName>
conforms to most character acquisitions in the 
<taxonomicName id="05C02FBEFFE3FFE3ACF3FEC7FE95D331" box="[193,297,261,284]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE3FFE3ACF3FEC7FE95D331" box="[193,297,261,284]" italics="true" pageId="7" pageNumber="352">T. caecus</emphasis>
</taxonomicName>
plexus, but lacks the occipital median node, and appears to be most closely related to 
<taxonomicName id="05C02FBEFFE3FFE3A8CAFEC7FA27D331" box="[1272,1435,260,284]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="shotoriensis">
<emphasis id="F0B4882FFFE3FFE3A8CAFEC7FA27D331" box="[1272,1435,260,284]" italics="true" pageId="7" pageNumber="352">T. shotoriensis</emphasis>
</taxonomicName>
(see 
<tableCitation id="8F426186FFE3FFE3ACFEFEE5FE9DD312" box="[204,289,295,319]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="7" pageNumber="352" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
for comparison). Interestingly, specimens of 
<taxonomicName id="05C02FBEFFE3FFE3AF17FEEBFC32D36D" box="[805,910,297,320]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE3FFE3AF17FEEBFC32D36D" box="[805,910,297,320]" italics="true" pageId="7" pageNumber="352">T. caecus</emphasis>
</taxonomicName>
described by Osmólska (1958) from the Holy Cross Mountains also lack an occipital median node (as in 
<taxonomicName id="05C02FBEFFE3FFE3AF78FE8FFC07D349" box="[842,955,332,356]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE3FFE3AF78FE8FFC07D349" box="[842,955,332,356]" italics="true" pageId="7" pageNumber="352">T. chopini</emphasis>
</taxonomicName>
), but do show all the other characteristic features for this widely distributed species (for details, see Crônier, 2003). It also should be clearly emphasized here that all representatives of 
<taxonomicName id="05C02FBEFFE3FFE3AE3DFE57FDC7D381" box="[527,635,405,428]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE3FFE3AE3DFE57FDC7D381" box="[527,635,405,428]" italics="true" pageId="7" pageNumber="352">T. caecus</emphasis>
</taxonomicName>
known from outside of 
<collectingCountry id="BAD714ADFFE3FFE3AF9EFE56FC46D381" box="[940,1018,404,428]" name="Poland" pageId="7" pageNumber="352">Poland</collectingCountry>
possess the occipital median node (Osmólska, 1958; Crônier, 2003). It is therefore possible that specimens of 
<taxonomicName id="05C02FBEFFE3FFE3AFCCFE7BFBD4D3FD" box="[1022,1128,441,464]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="caecus">
<emphasis id="F0B4882FFFE3FFE3AFCCFE7BFBD4D3FD" box="[1022,1128,441,464]" italics="true" pageId="7" pageNumber="352">T. caecus</emphasis>
</taxonomicName>
from 
<collectingCountry id="BAD714ADFFE3FFE3A880FE75FABCD3E2" box="[1202,1280,439,463]" name="Poland" pageId="7" pageNumber="352">Poland</collectingCountry>
represent the most ancestral populations (lacking the occipital median node) or, more probably, specific ecophenotypic variants of that species. Consequently, the absence of the occipital median node in 
<taxonomicName id="05C02FBEFFE3FFE3AFD4FDC3FBE4D035" box="[998,1112,512,536]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="chopini">
<emphasis id="F0B4882FFFE3FFE3AFD4FDC3FBE4D035" box="[998,1112,512,536]" italics="true" pageId="7" pageNumber="352">T. chopini</emphasis>
</taxonomicName>
may be due to atavism. The only other species that seems to lack the occipital median node is 
<taxonomicName id="05C02FBEFFE3FFE3AFB7FDE7FC54D011" box="[901,1000,548,572]" class="Trilobita" family="Phacopidae" genus="Trimerocephalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="7" pageNumber="352" phylum="Arthropoda" rank="species" species="mimbi">
<emphasis id="F0B4882FFFE3FFE3AFB7FDE7FC54D011" box="[901,1000,548,572]" italics="true" pageId="7" pageNumber="352">T. mimbi</emphasis>
</taxonomicName>
(see 
<tableCitation id="8F426186FFE3FFE3A814FDE6FBC5D011" box="[1062,1145,548,572]" captionStart="TABLE 1" captionStartId="6.[151,235,152,173]" captionTargetBox="[159,1423,361,672]" captionTargetPageId="6" captionText="TABLE 1. Comparison of nine morphological features between Trimerocephalus chopini n. sp. and selected representatives of the genus Trimerocephalus. Feature 1: facial sutures that do not cut the cheeks; Feature 2: a well developed, straight preoccipital furrow, S 1; Feature 3: absence of ocular protuberances; Feature 4: coarse cephalic tuberculation; Feature 5: a trapezoidal pygidium; Feature 6: preoccipital ring (L 1) with two median tubercles; Feature 7: the absence of an occipital median node; Feature 8: cheeks form very slight recess in contact with facial suture; Feature 9: truncated genal angle. Plus symbol (+) denotes the above-mentioned conditions are present." httpUri="http://table.plazi.org/id/96BF04B5FFE2FFE2ACA5FF5AFC9ED366" pageId="7" pageNumber="352" tableUuid="96BF04B5FFE2FFE2ACA5FF5AFC9ED366">Table 1</tableCitation>
), but this taxon is known only from the 
<typeStatus id="1D7BEA9FFFE3FFE3AD09FD85FE21D072" box="[315,413,583,607]" pageId="7" pageNumber="352" type="holotype">holotype</typeStatus>
cephalon (specimen WAM. 07.284; Feist 
<emphasis id="F0B4882FFFE3FFE3AFB3FD8BFC07D04D" box="[897,955,584,608]" italics="true" pageId="7" pageNumber="352">et al.</emphasis>
, 2009, p. 23), hence additional specimens are needed to confirm if this is a true absence in this species.
</paragraph>
</subSubSection>
</treatment>
</document>