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<document id="C21F9ECD0097918922B56413141C9DF5" ID-CLB-Dataset="46570" ID-DOI="10.11646/zootaxa.3755.5.9" ID-GBIF-Dataset="4c3ba205-d1d6-4a9e-87c7-eaeb0d829ca7" ID-ISSN="1175-5326" ID-Zenodo-Dep="226656" ID-ZooBank="96AA169A-0FB4-4475-BB99-559FC2EBA860" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460375474015" checkinUser="plazi" docAuthor="Gill, Anthony C., Bogorodsky, Sergey V. &amp; Mal, Ahmad O." docDate="2014" docId="482B87B8FFB9FFE011CFFF2A7BBFFDDA" docLanguage="en" docName="zt03755p495.pdf" docOrigin="Zootaxa 3755 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gymnoxenisthmus Gill, Bogorodsky &amp; Mal, 2014, new genus" docType="treatment" docVersion="9" lastPageNumber="495" masterDocId="B412FFC0FFB8FFE41158FFBD7829FFE6" masterDocTitle="Gymnoxenisthmus tigrellus, new genus and species of gobioid fish from the Red Sea (Gobioidei: Xenisthmidae)" masterLastPageNumber="495" masterPageNumber="491" pageNumber="492" updateTime="1698296230469" updateUser="ExternalLinkService">
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<mods:title id="16D75CE3CF5527FB2884C53C2241B9AE">Gymnoxenisthmus tigrellus, new genus and species of gobioid fish from the Red Sea (Gobioidei: Xenisthmidae)</mods:title>
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<mods:namePart id="0739E7A6FCBFF13058CAFCEC7EDD5AE1">Gill, Anthony C.</mods:namePart>
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<mods:namePart id="3DE5E4D5AD5822271A058F6FE611C1C1">Bogorodsky, Sergey V.</mods:namePart>
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<mods:namePart id="73B1CC3C01800DD6B658C10B9165852A">Mal, Ahmad O.</mods:namePart>
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<mods:title id="DA805CFEBD8506679CAA1106E027E756">Zootaxa</mods:title>
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<mods:part id="D24ED9F49120D8AD27313DEFDE4E9E4C">
<mods:date id="7374AFDC1AD7C19C1C42C1F9DE869B68">2014</mods:date>
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<mods:identifier id="6D7441B54CAEDFF171F093B29FAF6268" type="DOI">10.11646/zootaxa.3755.5.9</mods:identifier>
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<treatment id="482B87B8FFB9FFE011CFFF2A7BBFFDDA" ID-DOI="http://doi.org/10.5281/zenodo.5671203" ID-GBIF-Taxon="119413029" ID-Zenodo-Dep="5671203" LSID="urn:lsid:plazi:treatment:482B87B8FFB9FFE011CFFF2A7BBFFDDA" httpUri="http://treatment.plazi.org/id/482B87B8FFB9FFE011CFFF2A7BBFFDDA" lastPageId="4" lastPageNumber="495" pageId="1" pageNumber="492">
<subSubSection id="88986525FFB9FFE511CFFF2A7A20FF54" box="[151,521,151,178]" pageId="1" pageNumber="492" type="nomenclature">
<paragraph id="C03D36AEFFB9FFE511CFFF2A7A20FF54" blockId="1.[151,521,151,178]" box="[151,521,151,178]" pageId="1" pageNumber="492">
<heading id="9B7581C2FFB9FFE511CFFF2A7A20FF54" bold="true" box="[151,521,151,178]" fontSize="11" level="1" pageId="1" pageNumber="492" reason="1">
<emphasis id="F2F6EABCFFB9FFE511CFFF2A7A20FF54" bold="true" box="[151,521,151,178]" pageId="1" pageNumber="492">
<taxonomicName id="07824D2DFFB9FFE511CFFF2A7950FF57" ID-CoL="4RYL" box="[151,377,151,177]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="F2F6EABCFFB9FFE511CFFF2A7950FF57" bold="true" box="[151,377,151,177]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E9C557C7FFB9FFE510D0FF257A20FF54" box="[392,521,152,178]" pageId="1" pageNumber="492" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
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<subSubSection id="88986525FFB9FFE511CFFF637AE3FF12" box="[151,714,222,244]" pageId="1" pageNumber="492" type="materials_examined">
<paragraph id="C03D36AEFFB9FFE511CFFF637AE3FF12" blockId="1.[151,714,222,244]" box="[151,714,222,244]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE511CFFF63790BFF12" bold="true" box="[151,290,222,244]" pageId="1" pageNumber="492">
<typeStatus id="1F39880CFFB9FFE511CFFF6378E5FF12" box="[151,204,222,244]" pageId="1" pageNumber="492">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="07824D2DFFB9FFE51071FF637A15FF12" box="[297,572,222,244]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="species" species="tigrellus" status="sp. nov.">
<emphasis id="F2F6EABCFFB9FFE51071FF637A15FF12" box="[297,572,222,244]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus tigrellus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="E9C557C7FFB9FFE51311FF627AEEFF12" box="[585,711,222,244]" pageId="1" pageNumber="492" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="88986525FFB9FFE511CFFE9D7A37FC78" pageId="1" pageNumber="492" type="diagnosis">
<paragraph id="C03D36AEFFB9FFE511CFFE9D7C40FE64" blockId="1.[151,1437,288,1646]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE511CFFE9D7938FEDF" bold="true" box="[151,273,288,313]" pageId="1" pageNumber="492">Diagnosis.</emphasis>
The following combination of characters distinguishes
<taxonomicName id="07824D2DFFB9FFE512EEFE9C7CAAFEDC" box="[950,1155,289,314]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE512EEFE9C7CAAFEDC" box="[950,1155,289,314]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
from other xenisthmid genera: head pores absent; scales lacking; first dorsal fin with five spines; dorsal- and anal-fin rays branched; at least some pectoral-fin rays branched; pelvic fin with a spine and five unbranched rays.
</paragraph>
<paragraph id="C03D36AEFFB9FFE5119FFE317A37FC78" blockId="1.[151,1437,288,1646]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE5119FFE317944FE43" bold="true" box="[199,365,396,421]" pageId="1" pageNumber="492">Comparisons.</emphasis>
Characters distinguishing xenisthmid genera are summarised in
<tableCitation id="8D000315FFB9FFE51506FE307C9CFE40" box="[1118,1205,397,422]" captionStart="TABLE 1" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1405,459,564]" captionTargetPageId="2" captionText="TABLE 1. Comparison of selected characters of xenisthmid genera. * indicates where data are included from undescribed species." httpUri="http://table.plazi.org/id/94FD6626FFBAFFE611CFFEF1795AFE60" pageId="1" pageNumber="492" tableUuid="94FD6626FFBAFFE611CFFEF1795AFE60">Table 1</tableCitation>
. The genus is most similar to
<taxonomicName id="07824D2DFFB9FFE51050FE0C7995FE2C" box="[264,444,433,458]" class="Actinopterygii" family="Xenisthmidae" genus="Paraxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51050FE0C7995FE2C" box="[264,444,433,458]" italics="true" pageId="1" pageNumber="492">Paraxenisthmus</emphasis>
</taxonomicName>
and
<taxonomicName id="07824D2DFFB9FFE510ABFE0C7A5DFE2C" box="[499,628,433,458]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE510ABFE0C7A5DFE2C" box="[499,628,433,458]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
in general body form, in most meristic details, and in having dorsal-and anal-fin rays and at least some pectoral-fin rays branched. It differs from
<taxonomicName id="07824D2DFFB9FFE5128BFE687CA1FE08" box="[979,1160,469,494]" class="Actinopterygii" family="Xenisthmidae" genus="Paraxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5128BFE687CA1FE08" box="[979,1160,469,494]" italics="true" pageId="1" pageNumber="492">Paraxenisthmus</emphasis>
</taxonomicName>
in lacking laterosensory pores on the head, vomerine and palatine teeth, and scales, and in having only five (versus six) spines in the first dorsal fin and a relatively broad (versus narrow) proximal head on the third branchiostegal ray. It differs from
<taxonomicName id="07824D2DFFB9FFE511CFFDFC7931FDBC" box="[151,280,577,602]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE511CFFDFC7931FDBC" box="[151,280,577,602]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
in lacking scales, laterosensory pores on the head, and developed gill rakers (though tiny rudiments are present), and in having only five (versus six) spines in the first dorsal fin, a narrower proximal head on the third branchiostegal ray (see Remarks below), and in having only unbranched rays in the pelvic fins (versus anterior four segmented rays branched in
<taxonomicName id="07824D2DFFB9FFE51086FD107A76FD20" box="[478,607,685,710]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51086FD107A76FD20" box="[478,607,685,710]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
).
<taxonomicName id="07824D2DFFB9FFE51321FD107B6FFD20" box="[633,838,685,710]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51321FD107B6FFD20" box="[633,838,685,710]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
might also be confused with
<taxonomicName id="07824D2DFFB9FFE515C4FD137CDAFD23" box="[1180,1267,686,709]" class="Actinopterygii" family="Xenisthmidae" genus="Rotuma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE515C4FD137CDAFD23" box="[1180,1267,686,709]" italics="true" pageId="1" pageNumber="492">Rotuma</emphasis>
</taxonomicName>
, with which it shares the following characters: first dorsal fin with five spines; pelvic fins with a spine and five unbranched segmented rays, the inner of which is vestigial; no head pores; and no scales. It is readily distinguished from
<taxonomicName id="07824D2DFFB9FFE511CFFCA778C6FCD7" box="[151,239,794,817]" class="Actinopterygii" family="Xenisthmidae" genus="Rotuma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE511CFFCA778C6FCD7" box="[151,239,794,817]" italics="true" pageId="1" pageNumber="492">Rotuma</emphasis>
</taxonomicName>
in having branched (versus unbranched) dorsal-, anal- and pectoral-fin rays, a relatively broad (versus narrow) proximal head on the third branchiostegal ray; autogenous anterior and posterior ceratohyals (versus a single ceratohyal ossification present); and more segmented rays in second dorsal and anal fins (13 and 12, respectively, versus
<quantity id="077A9B4BFFB9FFE51021FC3B798DFC78" box="[377,420,901,926]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="1" pageNumber="492" unit="in" value="9.0">9 in</quantity>
both fins).
</paragraph>
</subSubSection>
<subSubSection id="88986525FFB9FFE5119FFC14799FF9C0" pageId="1" pageNumber="492" type="discussion">
<paragraph id="C03D36AEFFB9FFE5119FFC14799FF9C0" blockId="1.[151,1437,288,1646]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE5119FFC147912FC27" bold="true" box="[199,315,937,961]" pageId="1" pageNumber="492">Remarks.</emphasis>
<bibRefCitation id="A4134B5FFFB9FFE51013FC147A76FC24" author="Gill" box="[331,607,937,962]" pageId="1" pageNumber="492" refString="Gill, A. C. &amp; Hoese, D. F. (1993) Paraxenisthmus springeri, new genus and species of gobioid fish from the West Pacific, and its phylogenetic position within the Xenisthmidae. Copeia, 1993, 1049 - 1057. http: // dx. doi. org / 10.2307 / 1447083" type="journal article" year="1993">Gill and Hoese (1993)</bibRefCitation>
hypothesised the following relationships among xenisthmid genera: (
<taxonomicName id="07824D2DFFB9FFE511F8FC70797CFC00" box="[160,341,973,998]" class="Actinopterygii" family="Xenisthmidae" genus="Paraxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE511F8FC70797CFC00" box="[160,341,973,998]" italics="true" pageId="1" pageNumber="492">Paraxenisthmus</emphasis>
</taxonomicName>
(
<taxonomicName id="07824D2DFFB9FFE5103EFC7079CEFC00" box="[358,487,973,998]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5103EFC7079CEFC00" box="[358,487,973,998]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
(
<taxonomicName id="07824D2DFFB9FFE510A1FC737A79FC03" box="[505,592,974,997]" class="Actinopterygii" family="Xenisthmidae" genus="Rotuma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE510A1FC737A79FC03" box="[505,592,974,997]" italics="true" pageId="1" pageNumber="492">Rotuma</emphasis>
</taxonomicName>
+
<taxonomicName id="07824D2DFFB9FFE51329FC737A9BFC03" box="[625,690,974,997]" class="Actinopterygii" family="Xenisthmidae" genus="Tyson" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51329FC737A9BFC03" box="[625,690,974,997]" italics="true" pageId="1" pageNumber="492">Tyson</emphasis>
</taxonomicName>
+
<taxonomicName id="07824D2DFFB9FFE5138BFC707BBDFC00" box="[723,916,973,998]" class="Actinopterygii" family="Xenisthmidae" genus="Allomicrodesmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5138BFC707BBDFC00" box="[723,916,973,998]" italics="true" pageId="1" pageNumber="492">Allomicrodesmus</emphasis>
</taxonomicName>
))). Character evidence for these relationships was in part from osteological characters, which have not yet been verified in
<taxonomicName id="07824D2DFFB9FFE5155CFC4C7CECFBEC" box="[1028,1221,1009,1034]" class="Actinopterygii" family="Xenisthmidae" genus="Allomicrodesmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5155CFC4C7CECFBEC" box="[1028,1221,1009,1034]" italics="true" pageId="1" pageNumber="492">Allomicrodesmus</emphasis>
</taxonomicName>
. We similarly lack detailed osteological information for
<taxonomicName id="07824D2DFFB9FFE51365FBA87B23FBC8" box="[573,778,1045,1070]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51365FBA87B23FBC8" box="[573,778,1045,1070]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
. Considering those characters that can be observed in the two genera, evidence supports placement of
<taxonomicName id="07824D2DFFB9FFE513C3FB847B41FBB4" box="[667,872,1081,1106]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE513C3FB847B41FBB4" box="[667,872,1081,1106]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
in a clade that also includes
<taxonomicName id="07824D2DFFB9FFE51598FB877D3EFBB7" box="[1216,1303,1082,1105]" class="Actinopterygii" family="Xenisthmidae" genus="Rotuma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51598FB877D3EFBB7" box="[1216,1303,1082,1105]" italics="true" pageId="1" pageNumber="492">Rotuma</emphasis>
</taxonomicName>
,
<taxonomicName id="07824D2DFFB9FFE51471FB877D41FBB7" box="[1321,1384,1082,1105]" class="Actinopterygii" family="Xenisthmidae" genus="Tyson" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51471FB877D41FBB7" box="[1321,1384,1082,1105]" italics="true" pageId="1" pageNumber="492">Tyson</emphasis>
</taxonomicName>
and
<taxonomicName id="07824D2DFFB9FFE511CFFBE07971FB90" box="[151,344,1117,1142]" class="Actinopterygii" family="Xenisthmidae" genus="Allomicrodesmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE511CFFBE07971FB90" box="[151,344,1117,1142]" italics="true" pageId="1" pageNumber="492">Allomicrodesmus</emphasis>
</taxonomicName>
(synapomorphies: five or fewer spines in first dorsal fin; sensory pores absent; scales absent; fifth segmented pelvic-fin ray vestigial or absent). Within this clade,
<taxonomicName id="07824D2DFFB9FFE51207FB3F7B9FFB7F" box="[863,950,1154,1177]" class="Actinopterygii" family="Xenisthmidae" genus="Rotuma" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51207FB3F7B9FFB7F" box="[863,950,1154,1177]" italics="true" pageId="1" pageNumber="492">Rotuma</emphasis>
</taxonomicName>
,
<taxonomicName id="07824D2DFFB9FFE5129CFB3F7C2CFB7F" box="[964,1029,1154,1177]" class="Actinopterygii" family="Xenisthmidae" genus="Tyson" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5129CFB3F7C2CFB7F" box="[964,1029,1154,1177]" italics="true" pageId="1" pageNumber="492">Tyson</emphasis>
</taxonomicName>
and
<taxonomicName id="07824D2DFFB9FFE51563FB3C7CD5FB7C" box="[1083,1276,1153,1178]" class="Actinopterygii" family="Xenisthmidae" genus="Allomicrodesmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51563FB3C7CD5FB7C" box="[1083,1276,1153,1178]" italics="true" pageId="1" pageNumber="492">Allomicrodesmus</emphasis>
</taxonomicName>
in turn form a clade that excludes
<taxonomicName id="07824D2DFFB9FFE510DAFB187A66FB58" box="[386,591,1189,1214]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE510DAFB187A66FB58" box="[386,591,1189,1214]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
(synapomorphies: all segmented second dorsal-fin rays unbranched; all segmented anal-fin rays unbranched; all pectoral-fin rays unbranched; ceratohyals represented by a single ossification). Contrary to these relationships,
<taxonomicName id="07824D2DFFB9FFE513CDFB507B4BFAE0" box="[661,866,1261,1286]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE513CDFB507B4BFAE0" box="[661,866,1261,1286]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
shares a possible synapomorphy with
<taxonomicName id="07824D2DFFB9FFE5144BFB507DBDFAE0" box="[1299,1428,1261,1286]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5144BFB507DBDFAE0" box="[1299,1428,1261,1286]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
: enlarged proximal head on third branchiostegal ray.
<bibRefCitation id="A4134B5FFFB9FFE513AAFAAC7BDAFACC" author="Gill" box="[754,1011,1297,1322]" pageId="1" pageNumber="492" refString="Gill, A. C. &amp; Hoese, D. F. (1993) Paraxenisthmus springeri, new genus and species of gobioid fish from the West Pacific, and its phylogenetic position within the Xenisthmidae. Copeia, 1993, 1049 - 1057. http: // dx. doi. org / 10.2307 / 1447083" type="journal article" year="1993">Gill and Hoese (1993)</bibRefCitation>
noted that
<taxonomicName id="07824D2DFFB9FFE51521FAAC7CD3FACC" box="[1145,1274,1297,1322]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51521FAAC7CD3FACC" box="[1145,1274,1297,1322]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
species differ from other xenisthmids in having a very broad proximal head on the third branchiostegal ray (see
<bibRefCitation id="A4134B5FFFB9FFE515ABFA887DB1FAA8" author="Springer" box="[1267,1432,1333,1358]" pageId="1" pageNumber="492" refString="Springer, V. G. (1983) Tyson belos, new genus and species of western Pacific fish (Gobiidae, Xenisthminae), with discussions of gobioid osteology and classification. Smithsonian Contributions to Zoology, 390, 1 - 40. http: // dx. doi. org / 10.5479 / si. 00810282.390" type="journal article" year="1983">Springer 1983</bibRefCitation>
: fig. 9), which they interpreted as an autapomorphy of the genus. The proximal head of the third branchiostegal ray is also enlarged in
<taxonomicName id="07824D2DFFB9FFE5103FFAC07A1DFA70" box="[359,564,1405,1430]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE5103FFAC07A1DFA70" box="[359,564,1405,1430]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
, but not to the same extent as in
<taxonomicName id="07824D2DFFB9FFE512FBFAC07C0DFA70" box="[931,1060,1405,1430]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE512FBFAC07C0DFA70" box="[931,1060,1405,1430]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
species (slightly broader than the proximal head on the fourth ray in
<taxonomicName id="07824D2DFFB9FFE51346FA1C7AC2FA5C" box="[542,747,1441,1466]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51346FA1C7AC2FA5C" box="[542,747,1441,1466]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
, versus about twice the width of the fourth ray in
<taxonomicName id="07824D2DFFB9FFE51443FA1C7DB5FA5C" box="[1307,1436,1441,1466]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51443FA1C7DB5FA5C" box="[1307,1436,1441,1466]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
species). More detailed phylogenetic analysis must await more complete osteological studies of
<taxonomicName id="07824D2DFFB9FFE51596FA787DB2FA38" box="[1230,1435,1477,1502]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51596FA787DB2FA38" box="[1230,1435,1477,1502]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus</emphasis>
</taxonomicName>
and
<taxonomicName id="07824D2DFFB9FFE51191FA5479A3F9E4" box="[201,394,1513,1538]" class="Actinopterygii" family="Xenisthmidae" genus="Allomicrodesmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51191FA5479A3F9E4" box="[201,394,1513,1538]" italics="true" pageId="1" pageNumber="492">Allomicrodesmus</emphasis>
</taxonomicName>
(which in turn are dependent on the discovery of additional specimens of both genera for osteological preparation).
</paragraph>
</subSubSection>
<subSubSection id="88986525FFB9FFE5119FF98F797FF918" pageId="1" pageNumber="492" type="etymology">
<paragraph id="C03D36AEFFB9FFE5119FF98F7C57F988" blockId="1.[151,1437,288,1646]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE5119FF98F7967F9AC" bold="true" box="[199,334,1586,1610]" pageId="1" pageNumber="492">Etymology.</emphasis>
The generic name is a combination of the Greek
<emphasis id="F2F6EABCFFB9FFE51222F98F7BE6F9AF" box="[890,975,1586,1609]" italics="true" pageId="1" pageNumber="492">gymnos</emphasis>
, meaning bare or naked, and the gobioid genus
<taxonomicName id="07824D2DFFB9FFE51187F9E87949F988" box="[223,352,1621,1646]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="genus">
<emphasis id="F2F6EABCFFB9FFE51187F9E87949F988" box="[223,352,1621,1646]" italics="true" pageId="1" pageNumber="492">Xenisthmus</emphasis>
</taxonomicName>
, and alludes to the absence of scales on the body. Gender is masculine.
</paragraph>
<paragraph id="C03D36AEFFB9FFE511CFF97C797FF918" blockId="1.[151,621,1729,1790]" pageId="1" pageNumber="492">
<taxonomicName id="07824D2DFFB9FFE511CFF97C79FCF93C" box="[151,469,1729,1754]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="492" phylum="Chordata" rank="species" species="tigrellus" status="sp. nov.">
<emphasis id="F2F6EABCFFB9FFE511CFF97C79FCF93C" bold="true" box="[151,469,1729,1754]" italics="true" pageId="1" pageNumber="492">Gymnoxenisthmus tigrellus</emphasis>
</taxonomicName>
͵
<taxonomicNameLabel id="E9C557C7FFB9FFE510BBF97C7A44F93C" box="[483,621,1729,1754]" pageId="1" pageNumber="492" rank="species">
<emphasis id="F2F6EABCFFB9FFE510BBF97C7A44F93C" bold="true" box="[483,621,1729,1754]" pageId="1" pageNumber="492">new species</emphasis>
</taxonomicNameLabel>
<figureCitation id="58B92A2BFFB9FFE511CFF95878DCF918" box="[151,245,1765,1790]" captionStart="FIGURE 1" captionStartId="3.[151,250,950,972]" captionTargetBox="[170,1416,595,926]" captionTargetId="figure@3.[170,1417,595,926]" captionTargetPageId="3" captionText="FIGURE 1. Gymnoxenisthmus tigrellus, holotype, SMF 34903, 15.2 mm SL, gravid female, Farasan Archipelago, Red Sea. Photo by Sven Traenkner." httpUri="https://zenodo.org/record/226657/files/figure.png" pageId="1" pageNumber="492">Figure 1</figureCitation>
,
<tableCitation id="8D000315FFB9FFE5105BF958797FF918" box="[259,342,1765,1790]" captionStart="TABLE 1" captionStartId="2.[151,239,332,355]" captionTargetBox="[159,1405,459,564]" captionTargetPageId="2" captionText="TABLE 1. Comparison of selected characters of xenisthmid genera. * indicates where data are included from undescribed species." httpUri="http://table.plazi.org/id/94FD6626FFBAFFE611CFFEF1795AFE60" pageId="1" pageNumber="492" tableUuid="94FD6626FFBAFFE611CFFEF1795AFE60">Table 1</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="88986525FFB9FFE511CFF8907C42F88C" pageId="1" pageNumber="492" type="materials_examined">
<paragraph id="C03D36AEFFB9FFE511CFF8907C42F88C" blockId="1.[151,1436,1837,2006]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE511CFF8907922F8A3" bold="true" box="[151,267,1837,1861]" pageId="1" pageNumber="492">
<typeStatus id="1F39880CFFB9FFE511CFF890792FF8A3" box="[151,262,1837,1861]" pageId="1" pageNumber="492" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="A693AE6BFFB9FFE51049F8937960F8A0" LSID="urn:lsid:biocol.org:col:34838" box="[273,329,1838,1862]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="492">SMF</collectionCode>
34903,
<quantity id="077A9B4BFFB9FFE510FCF8907A21F8A3" box="[420,520,1837,1862]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="1" pageNumber="492" unit="mm" value="15.2">15.2 mm</quantity>
SL, gravid female, Red Sea,
<collectingCountry id="B895763EFFB9FFE51216F8907BCDF8A0" box="[846,996,1837,1862]" name="Saudi Arabia" pageId="1" pageNumber="492">Saudi Arabia</collectingCountry>
, Farasan Archipelago, unnamed island,
<geoCoordinate id="A5B65069FFB9FFE511CFF8EC7900F88C" box="[151,297,1873,1898]" direction="north" orientation="latitude" pageId="1" pageNumber="492" precision="1" value="16.79085">16°47.451’N</geoCoordinate>
<geoCoordinate id="A5B65069FFB9FFE51077F8EF79E4F88C" box="[303,461,1874,1898]" direction="east" orientation="longitude" pageId="1" pageNumber="492" precision="1" value="42.1973">042°11.838’E</geoCoordinate>
, coll. S.V. Bogorodsky &amp; T. Alpermann,
<date id="B43C106EFFB9FFE512C4F8EF7C4EF88C" box="[924,1127,1873,1898]" pageId="1" pageNumber="492" value="2012-02-22">22 February 2012</date>
.
</paragraph>
</subSubSection>
<subSubSection id="88986525FFB9FFE5119FF8C979F6F868" box="[199,479,1908,1934]" pageId="1" pageNumber="492" type="diagnosis">
<paragraph id="C03D36AEFFB9FFE5119FF8C979F6F868" blockId="1.[151,1436,1837,2006]" box="[199,479,1908,1934]" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE5119FF8C97968F86B" bold="true" box="[199,321,1908,1933]" pageId="1" pageNumber="492">Diagnosis.</emphasis>
<collectingCountry id="B895763EFFB9FFE51010F8CB794EF868" box="[328,359,1910,1934]" name="American Samoa" pageId="1" pageNumber="492">As</collectingCountry>
for genus.
</paragraph>
</subSubSection>
<subSubSection id="88986525FFB9FFE0119FF8247973FE6E" lastPageId="4" lastPageNumber="495" pageId="1" pageNumber="492" type="description">
<paragraph id="C03D36AEFFB9FFE6119FF82479FEFEFA" blockId="1.[151,1436,1837,2006]" lastBlockId="2.[151,1436,151,285]" lastPageId="2" lastPageNumber="493" pageId="1" pageNumber="492">
<emphasis id="F2F6EABCFFB9FFE5119FF8247970F854" bold="true" box="[199,345,1945,1970]" pageId="1" pageNumber="492">Description.</emphasis>
Dorsal-fin rays V + I,13, all segmented rays branched; first dorsal-fin pterygiophore formula 3- 2210; anal-fin rays I,12, all segmented rays branched; pectoral-fin rays 15/15, upper 2/2 and lower 1/1 rays unbranched; pelvic-fin rays I,5, all segmented rays unbranched, inner ray vestigial; segmented caudal-fin rays 9 + 8; branched caudal-fin rays 6 + 6; upper unsegmented caudal-fin rays 6; lower unsegmented caudal-fin rays 6; total caudal-fin rays 29; no developed gill rakers (about 6 tiny rudiments present on upper part of ceratobranchial 1); vertebrae 10 + 16; epurals 2.
</paragraph>
<caption id="94FD6626FFBAFFE611CFFEF1795AFE60" ID-Table-UUID="94FD6626FFBAFFE611CFFEF1795AFE60" httpUri="http://table.plazi.org/id/94FD6626FFBAFFE611CFFEF1795AFE60" pageId="2" pageNumber="493" targetBox="[159,1405,459,564]" targetIsTable="true" targetPageId="2">
<paragraph id="C03D36AEFFBAFFE611CFFEF1795AFE60" blockId="2.[151,1436,331,390]" pageId="2" pageNumber="493">
<emphasis id="F2F6EABCFFBAFFE611CFFEF17938FE85" bold="true" box="[151,273,332,355]" pageId="2" pageNumber="493">TABLE 1.</emphasis>
Comparison of selected characters of xenisthmid genera. * indicates where data are included from undescribed species.
</paragraph>
</caption>
<paragraph id="C03D36AEFFBAFFE61028FE1F7BC5FC41" blockId="2.[368,1405,418,440]" lastBlockId="2.[159,1329,851,1007]" pageId="2" pageNumber="493">
<emphasis id="F2F6EABCFFBAFFE61028FE1F7D54FE5E" box="[368,1405,418,440]" italics="true" pageId="2" pageNumber="493">
<taxonomicName id="07824D2DFFBAFFE61028FE1F7A3AFE5E" box="[368,531,418,440]" class="Actinopterygii" family="Xenisthmidae" genus="Paraxenisthmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="493" phylum="Chordata" rank="genus">Paraxenisthmus</taxonomicName>
<taxonomicName id="07824D2DFFBAFFE6137BFE1F7AB3FE5E" box="[547,666,418,440]" class="Actinopterygii" family="Xenisthmidae" genus="Xenisthmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="493" phylum="Chordata" rank="genus">Xenisthmus</taxonomicName>
*
<taxonomicName id="07824D2DFFBAFFE6139EFE1F7BA9FE5E" box="[710,896,418,440]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="493" phylum="Chordata" rank="genus">Gymnoxenisthmus</taxonomicName>
<taxonomicName id="07824D2DFFBAFFE612CEFE1E7BCCFE5E" box="[918,997,419,440]" class="Actinopterygii" family="Xenisthmidae" genus="Rotuma" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="493" phylum="Chordata" rank="genus">Rotuma</taxonomicName>
<taxonomicName id="07824D2DFFBAFFE6156AFE1E7C44FE5E" box="[1074,1133,419,440]" class="Actinopterygii" family="Xenisthmidae" genus="Tyson" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="493" phylum="Chordata" rank="genus">Tyson</taxonomicName>
<taxonomicName id="07824D2DFFBAFFE61599FE1F7D5AFE5E" authority="Pelvic" authorityName="Pelvic" box="[1217,1395,418,440]" class="Actinopterygii" family="Xenisthmidae" genus="Allomicrodesmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="493" phylum="Chordata" rank="genus">Allomicrodesmus</taxonomicName>
*
</emphasis>
Pelvic I,
<date id="B43C106EFFBAFFE61026FCEE7A04FC8F" box="[382,557,851,873]" pageId="2" pageNumber="493">5 I</date>
,
<date id="B43C106EFFBAFFE6136AFCEE7AF9FC8F" box="[562,720,851,873]" pageId="2" pageNumber="493">5 I</date>
,5, inner ray I,5, spine and 1 3 or absent vestigial inner ray vestigial
</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FE767D29FDD2" pageId="2" pageNumber="493">
<table id="B282C40EFFBA001B11C7FE767D54FDD2" box="[159,1405,459,564]" gridcols="7" gridrows="3" pageId="2" pageNumber="493">
<tr id="7EB234ECFFBA001B11C7FE767D54FE07" box="[159,1405,459,481]" gridrow="0" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FE7678C9FE07" box="[159,224,459,481]" gridcol="0" gridrow="0" pageId="2" pageNumber="493">Scales</th>
<th id="3D635D90FFBA001B1028FE767994FE07" box="[368,445,459,481]" gridcol="1" gridrow="0" pageId="2" pageNumber="493">present</th>
<th id="3D635D90FFBA001B137BFE767A59FE07" box="[547,624,459,481]" gridcol="2" gridrow="0" pageId="2" pageNumber="493">present</th>
<th id="3D635D90FFBA001B139EFE767B2EFE07" box="[710,775,459,481]" gridcol="3" gridrow="0" pageId="2" pageNumber="493">absent</th>
<th id="3D635D90FFBA001B12CEFE767BFEFE07" box="[918,983,459,481]" gridcol="4" gridrow="0" pageId="2" pageNumber="493">absent</th>
<th id="3D635D90FFBA001B156AFE767C5AFE07" box="[1074,1139,459,481]" gridcol="5" gridrow="0" pageId="2" pageNumber="493">absent</th>
<th id="3D635D90FFBA001B1599FE767D54FE07" box="[1217,1405,459,481]" gridcol="6" gridrow="0" pageId="2" pageNumber="493">absent</th>
</tr>
<tr id="7EB234ECFFBA001B11C7FE497D54FDEC" box="[159,1405,500,522]" gridrow="1" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FE4978C9FDEC" box="[159,224,500,522]" gridcol="0" gridrow="1" pageId="2" pageNumber="493">D1</th>
<td id="3D635D90FFBA001B1028FE497994FDEC" box="[368,445,500,522]" gridcol="1" gridrow="1" pageId="2" pageNumber="493">VI</td>
<td id="3D635D90FFBA001B137BFE497A59FDEC" box="[547,624,500,522]" gridcol="2" gridrow="1" pageId="2" pageNumber="493">VI</td>
<td id="3D635D90FFBA001B139EFE497B2EFDEC" box="[710,775,500,522]" gridcol="3" gridrow="1" pageId="2" pageNumber="493">V</td>
<td id="3D635D90FFBA001B12CEFE497BFEFDEC" box="[918,983,500,522]" gridcol="4" gridrow="1" pageId="2" pageNumber="493">V</td>
<td id="3D635D90FFBA001B156AFE497C5AFDEC" box="[1074,1139,500,522]" gridcol="5" gridrow="1" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B1599FE497D54FDEC" box="[1217,1405,500,522]" gridcol="6" gridrow="1" pageId="2" pageNumber="493">II</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FDA37D54FDD2" box="[159,1405,542,564]" gridrow="2" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FDA378C9FDD2" box="[159,224,542,564]" gridcol="0" gridrow="2" pageId="2" pageNumber="493">D2</th>
<td id="3D635D90FFBA001B1028FDA37994FDD2" box="[368,445,542,564]" gridcol="1" gridrow="2" pageId="2" pageNumber="493">I,11–12</td>
<td id="3D635D90FFBA001B137BFDA37A59FDD2" box="[547,624,542,564]" gridcol="2" gridrow="2" pageId="2" pageNumber="493">I,11–15</td>
<td id="3D635D90FFBA001B139EFDA37B2EFDD2" box="[710,775,542,564]" gridcol="3" gridrow="2" pageId="2" pageNumber="493">I,13</td>
<td id="3D635D90FFBA001B12CEFDA37BFEFDD2" box="[918,983,542,564]" gridcol="4" gridrow="2" pageId="2" pageNumber="493">I,9</td>
<td id="3D635D90FFBA001B156AFDA37C5AFDD2" box="[1074,1139,542,564]" gridcol="5" gridrow="2" pageId="2" pageNumber="493">I,8–9</td>
<td id="3D635D90FFBA001B1599FDA37D54FDD2" box="[1217,1405,542,564]" gridcol="6" gridrow="2" pageId="2" pageNumber="493">29–33</td>
</tr>
</table>
</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FDFA7CF2FCC7" pageId="2" pageNumber="493">
<table id="B282C40EFFBA001B11C7FDFA7D54FCA6" box="[159,1405,583,832]" gridcols="6" gridrows="5" pageId="2" pageNumber="493">
<tr id="7EB234ECFFBA001B11C7FDFA7D54FD9A" box="[159,1405,583,636]" gridrow="0" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FDFA7987FD9A" box="[159,430,583,636]" gridcol="0" gridrow="0" pageId="2" pageNumber="493">Segmented D2 yes rays branched</th>
<th id="3D635D90FFBA001B137BFDFA7A58FD9A" box="[547,625,583,636]" gridcol="1" gridrow="0" pageId="2" pageNumber="493">yes</th>
<th id="3D635D90FFBA001B139EFDFA7AC7FD9A" box="[710,750,583,636]" gridcol="2" gridrow="0" pageId="2" pageNumber="493">yes</th>
<th id="3D635D90FFBA001B12CEFDFA7B98FD9A" box="[918,945,583,636]" gridcol="3" gridrow="0" pageId="2" pageNumber="493">no</th>
<th id="3D635D90FFBA001B156AFDFA7C58FD9A" box="[1074,1137,583,636]" gridcol="4" gridrow="0" pageId="2" pageNumber="493">no</th>
<th id="3D635D90FFBA001B1599FDFA7D54FD9A" box="[1217,1405,583,636]" gridcol="5" gridrow="0" pageId="2" pageNumber="493">no</th>
</tr>
<tr id="7EB234ECFFBA001B11C7FD327D54FD43" box="[159,1405,655,677]" gridrow="1" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FD327987FD43" box="[159,430,655,677]" gridcol="0" gridrow="1" pageId="2" pageNumber="493">A I,10</th>
<td id="3D635D90FFBA001B137BFD327A58FD43" box="[547,625,655,677]" gridcol="1" gridrow="1" pageId="2" pageNumber="493">I,10–14</td>
<td id="3D635D90FFBA001B139EFD327AC7FD43" box="[710,750,655,677]" gridcol="2" gridrow="1" pageId="2" pageNumber="493">I,12</td>
<td id="3D635D90FFBA001B12CEFD327B98FD43" box="[918,945,655,677]" gridcol="3" gridrow="1" pageId="2" pageNumber="493">I,9</td>
<td id="3D635D90FFBA001B156AFD327C58FD43" box="[1074,1137,655,677]" gridcol="4" gridrow="1" pageId="2" pageNumber="493">I,8–9</td>
<td id="3D635D90FFBA001B1599FD327D54FD43" box="[1217,1405,655,677]" gridcol="5" gridrow="1" pageId="2" pageNumber="493">23–26</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FD047D54FD29" box="[159,1405,697,719]" gridrow="2" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FD047987FD29" box="[159,430,697,719]" gridcol="0" gridrow="2" pageId="2" pageNumber="493">A rays branched yes</th>
<td id="3D635D90FFBA001B137BFD047A58FD29" box="[547,625,697,719]" gridcol="1" gridrow="2" pageId="2" pageNumber="493">yes</td>
<td id="3D635D90FFBA001B139EFD047AC7FD29" box="[710,750,697,719]" gridcol="2" gridrow="2" pageId="2" pageNumber="493">yes</td>
<td id="3D635D90FFBA001B12CEFD047B98FD29" box="[918,945,697,719]" gridcol="3" gridrow="2" pageId="2" pageNumber="493">no</td>
<td id="3D635D90FFBA001B156AFD047C58FD29" box="[1074,1137,697,719]" gridcol="4" gridrow="2" pageId="2" pageNumber="493">no</td>
<td id="3D635D90FFBA001B1599FD047D54FD29" box="[1217,1405,697,719]" gridcol="5" gridrow="2" pageId="2" pageNumber="493">no</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FD5F7D54FD1E" box="[159,1405,738,760]" gridrow="3" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FD5F7987FD1E" box="[159,430,738,760]" gridcol="0" gridrow="3" pageId="2" pageNumber="493">Pectoral 15–18</th>
<td id="3D635D90FFBA001B137BFD5F7A58FD1E" box="[547,625,738,760]" gridcol="1" gridrow="3" pageId="2" pageNumber="493">15–18</td>
<td id="3D635D90FFBA001B139EFD5F7AC7FD1E" box="[710,750,738,760]" gridcol="2" gridrow="3" pageId="2" pageNumber="493">15</td>
<td id="3D635D90FFBA001B12CEFD5F7B98FD1E" box="[918,945,738,760]" gridcol="3" gridrow="3" pageId="2" pageNumber="493">16</td>
<td id="3D635D90FFBA001B156AFD5F7C58FD1E" box="[1074,1137,738,760]" gridcol="4" gridrow="3" pageId="2" pageNumber="493">17–21</td>
<td id="3D635D90FFBA001B1599FD5F7D54FD1E" box="[1217,1405,738,760]" gridcol="5" gridrow="3" pageId="2" pageNumber="493">10–12</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FCB67D54FCA6" box="[159,1405,779,832]" gridrow="4" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FCB67987FCA6" box="[159,430,779,832]" gridcol="0" gridrow="4" pageId="2" pageNumber="493">Pectoral rays yes branched</th>
<td id="3D635D90FFBA001B137BFCB67A58FCA6" box="[547,625,779,832]" gridcol="1" gridrow="4" pageId="2" pageNumber="493">yes</td>
<td id="3D635D90FFBA001B139EFCB67AC7FCA6" box="[710,750,779,832]" gridcol="2" gridrow="4" pageId="2" pageNumber="493">yes</td>
<td id="3D635D90FFBA001B12CEFCB67B98FCA6" box="[918,945,779,832]" gridcol="3" gridrow="4" pageId="2" pageNumber="493">no</td>
<td id="3D635D90FFBA001B156AFCB67C58FCA6" box="[1074,1137,779,832]" gridcol="4" gridrow="4" pageId="2" pageNumber="493">no</td>
<td id="3D635D90FFBA001B1599FCB67D54FCA6" box="[1217,1405,779,832]" gridcol="5" gridrow="4" pageId="2" pageNumber="493">no</td>
</tr>
</table>
</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FC077CF3FC36" blockId="2.[159,1329,851,1007]" box="[159,1242,954,976]" pageId="2" pageNumber="493">Pelvic rays no yes, outer 4 no no no no</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FC6478D5FC09" blockId="2.[159,1329,851,1007]" box="[159,252,985,1007]" pageId="2" pageNumber="493">branched</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FBBF7CCEFB4E" pageId="2" pageNumber="493">
<table id="B282C40EFFBA001B11C7FBBF7D54FB03" box="[159,1405,1026,1253]" gridcols="6" gridrows="3" pageId="2" pageNumber="493">
<tr id="7EB234ECFFBA001B11C7FBBF7D54FBFE" box="[159,1405,1026,1048]" gridrow="0" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FBBF798AFBFE" box="[159,419,1026,1048]" gridcol="0" gridrow="0" pageId="2" pageNumber="493">Segmented C rays 9 + 8</th>
<th id="3D635D90FFBA001B137BFBBF7A7EFBFE" box="[547,599,1026,1048]" gridcol="1" gridrow="0" pageId="2" pageNumber="493">9 + 8</th>
<th id="3D635D90FFBA001B139EFBBF7AD0FBFE" box="[710,761,1026,1048]" gridcol="2" gridrow="0" pageId="2" pageNumber="493">9 + 8</th>
<th id="3D635D90FFBA001B12CEFBBF7BE3FBFE" box="[918,970,1026,1048]" gridcol="3" gridrow="0" pageId="2" pageNumber="493">9 + 8</th>
<th id="3D635D90FFBA001B156AFBBF7C4CFBFE" box="[1074,1125,1026,1048]" gridcol="4" gridrow="0" pageId="2" pageNumber="493">8 + 7</th>
<th id="3D635D90FFBA001B1599FBBF7D54FBFE" box="[1217,1405,1026,1048]" gridcol="5" gridrow="0" pageId="2" pageNumber="493">8 + 7</th>
</tr>
<tr id="7EB234ECFFBA001B11C7FB967D54FB99" box="[159,1405,1067,1151]" gridrow="1" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FB96798AFB99" box="[159,419,1067,1151]" gridcol="0" gridrow="1" pageId="2" pageNumber="493">Upper 7–8 unsegmented C rays</th>
<td id="3D635D90FFBA001B137BFB967A7EFB99" box="[547,599,1067,1151]" gridcol="1" gridrow="1" pageId="2" pageNumber="493">6–9</td>
<td id="3D635D90FFBA001B139EFB967AD0FB99" box="[710,761,1067,1151]" gridcol="2" gridrow="1" pageId="2" pageNumber="493">6</td>
<td id="3D635D90FFBA001B12CEFB967BE3FB99" box="[918,970,1067,1151]" gridcol="3" gridrow="1" pageId="2" pageNumber="493">6</td>
<td id="3D635D90FFBA001B156AFB967C4CFB99" box="[1074,1125,1067,1151]" gridcol="4" gridrow="1" pageId="2" pageNumber="493">8–9</td>
<td id="3D635D90FFBA001B1599FB967D54FB99" box="[1217,1405,1067,1151]" gridcol="5" gridrow="1" pageId="2" pageNumber="493">7</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FB2F7D54FB03" box="[159,1405,1170,1253]" gridrow="2" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FB2F798AFB03" box="[159,419,1170,1253]" gridcol="0" gridrow="2" pageId="2" pageNumber="493">Lower 7–8 unsegmented C rays</th>
<td id="3D635D90FFBA001B137BFB2F7A7EFB03" box="[547,599,1170,1253]" gridcol="1" gridrow="2" pageId="2" pageNumber="493">5–9</td>
<td id="3D635D90FFBA001B139EFB2F7AD0FB03" box="[710,761,1170,1253]" gridcol="2" gridrow="2" pageId="2" pageNumber="493">6</td>
<td id="3D635D90FFBA001B12CEFB2F7BE3FB03" box="[918,970,1170,1253]" gridcol="3" gridrow="2" pageId="2" pageNumber="493">6</td>
<td id="3D635D90FFBA001B156AFB2F7C4CFB03" box="[1074,1125,1170,1253]" gridcol="4" gridrow="2" pageId="2" pageNumber="493">9</td>
<td id="3D635D90FFBA001B1599FB2F7D54FB03" box="[1217,1405,1170,1253]" gridcol="5" gridrow="2" pageId="2" pageNumber="493">6–7</td>
</tr>
</table>
</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FB447D2BFA52" pageId="2" pageNumber="493">
<table id="B282C40EFFBA001B11C7FB447D54FA52" box="[159,1405,1273,1460]" gridcols="7" gridrows="5" pageId="2" pageNumber="493">
<tr id="7EB234ECFFBA001B11C7FB447D54FAE9" box="[159,1405,1273,1295]" gridrow="0" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FB447911FAE9" box="[159,312,1273,1295]" gridcol="0" gridrow="0" pageId="2" pageNumber="493">Vertebrae</th>
<th id="3D635D90FFBA001B1028FB447997FAE9" box="[368,446,1273,1295]" gridcol="1" gridrow="0" pageId="2" pageNumber="493">10 + 16</th>
<th id="3D635D90FFBA001B137BFB447ABEFAE9" box="[547,663,1273,1295]" gridcol="2" gridrow="0" pageId="2" pageNumber="493">10 + 16–17</th>
<th id="3D635D90FFBA001B139EFB447B3AFAE9" box="[710,787,1273,1295]" gridcol="3" gridrow="0" pageId="2" pageNumber="493">10 + 16</th>
<th id="3D635D90FFBA001B12CEFB447BCBFAE9" box="[918,994,1273,1295]" gridcol="4" gridrow="0" pageId="2" pageNumber="493">11 + 15</th>
<th id="3D635D90FFBA001B156AFB447CA9FAE9" box="[1074,1152,1273,1295]" gridcol="5" gridrow="0" pageId="2" pageNumber="493">13 + 13</th>
<th id="3D635D90FFBA001B1599FB447D54FAE9" box="[1217,1405,1273,1295]" gridcol="6" gridrow="0" pageId="2" pageNumber="493">17 + 26–28</th>
</tr>
<tr id="7EB234ECFFBA001B11C7FA9F7D54FADE" box="[159,1405,1314,1336]" gridrow="1" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FA9F7911FADE" box="[159,312,1314,1336]" gridcol="0" gridrow="1" pageId="2" pageNumber="493">Epurals</th>
<td id="3D635D90FFBA001B1028FA9F7997FADE" box="[368,446,1314,1336]" gridcol="1" gridrow="1" pageId="2" pageNumber="493">2</td>
<td id="3D635D90FFBA001B137BFA9F7ABEFADE" box="[547,663,1314,1336]" gridcol="2" gridrow="1" pageId="2" pageNumber="493">2</td>
<td id="3D635D90FFBA001B139EFA9F7B3AFADE" box="[710,787,1314,1336]" gridcol="3" gridrow="1" pageId="2" pageNumber="493">2</td>
<td id="3D635D90FFBA001B12CEFA9F7BCBFADE" box="[918,994,1314,1336]" gridcol="4" gridrow="1" pageId="2" pageNumber="493">1</td>
<td id="3D635D90FFBA001B156AFA9F7CA9FADE" box="[1074,1152,1314,1336]" gridcol="5" gridrow="1" pageId="2" pageNumber="493">1</td>
<td id="3D635D90FFBA001B1599FA9F7D54FADE" box="[1217,1405,1314,1336]" gridcol="6" gridrow="1" pageId="2" pageNumber="493">?</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FAF67D54FA87" box="[159,1405,1355,1377]" gridrow="2" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FAF67911FA87" box="[159,312,1355,1377]" gridcol="0" gridrow="2" pageId="2" pageNumber="493">Head pores</th>
<td id="3D635D90FFBA001B1028FAF67997FA87" box="[368,446,1355,1377]" gridcol="1" gridrow="2" pageId="2" pageNumber="493">present</td>
<td id="3D635D90FFBA001B137BFAF67ABEFA87" box="[547,663,1355,1377]" gridcol="2" gridrow="2" pageId="2" pageNumber="493">present</td>
<td id="3D635D90FFBA001B139EFAF67B3AFA87" box="[710,787,1355,1377]" gridcol="3" gridrow="2" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B12CEFAF67BCBFA87" box="[918,994,1355,1377]" gridcol="4" gridrow="2" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B156AFAF67CA9FA87" box="[1074,1152,1355,1377]" gridcol="5" gridrow="2" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B1599FAF67D54FA87" box="[1217,1405,1355,1377]" gridcol="6" gridrow="2" pageId="2" pageNumber="493">absent</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FAC97D54FA6C" box="[159,1405,1396,1418]" gridrow="3" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FAC97911FA6C" box="[159,312,1396,1418]" gridcol="0" gridrow="3" pageId="2" pageNumber="493">Vomerine teeth</th>
<td id="3D635D90FFBA001B1028FAC97997FA6C" box="[368,446,1396,1418]" gridcol="1" gridrow="3" pageId="2" pageNumber="493">present</td>
<td id="3D635D90FFBA001B137BFAC97ABEFA6C" box="[547,663,1396,1418]" gridcol="2" gridrow="3" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B139EFAC97B3AFA6C" box="[710,787,1396,1418]" gridcol="3" gridrow="3" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B12CEFAC97BCBFA6C" box="[918,994,1396,1418]" gridcol="4" gridrow="3" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B156AFAC97CA9FA6C" box="[1074,1152,1396,1418]" gridcol="5" gridrow="3" pageId="2" pageNumber="493">present</td>
<td id="3D635D90FFBA001B1599FAC97D54FA6C" box="[1217,1405,1396,1418]" gridcol="6" gridrow="3" pageId="2" pageNumber="493">absent</td>
</tr>
<tr id="7EB234ECFFBA001B11C7FA237D54FA52" box="[159,1405,1438,1460]" gridrow="4" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FA237911FA52" box="[159,312,1438,1460]" gridcol="0" gridrow="4" pageId="2" pageNumber="493">Palatine teeth</th>
<td id="3D635D90FFBA001B1028FA237997FA52" box="[368,446,1438,1460]" gridcol="1" gridrow="4" pageId="2" pageNumber="493">present</td>
<td id="3D635D90FFBA001B137BFA237ABEFA52" box="[547,663,1438,1460]" gridcol="2" gridrow="4" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B139EFA237B3AFA52" box="[710,787,1438,1460]" gridcol="3" gridrow="4" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B12CEFA237BCBFA52" box="[918,994,1438,1460]" gridcol="4" gridrow="4" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B156AFA237CA9FA52" box="[1074,1152,1438,1460]" gridcol="5" gridrow="4" pageId="2" pageNumber="493">absent</td>
<td id="3D635D90FFBA001B1599FA237D54FA52" box="[1217,1405,1438,1460]" gridcol="6" gridrow="4" pageId="2" pageNumber="493">absent</td>
</tr>
</table>
</paragraph>
<paragraph id="C03D36AEFFBAFFE611C7FA7A7D21F90E" pageId="2" pageNumber="493">
<table id="B282C40EFFBA001B11C7FA7A7D54F8EF" box="[159,1405,1479,1801]" gridcols="6" gridrows="3" pageId="2" pageNumber="493">
<tr id="7EB234ECFFBA001B11C7FA7A7D54F9BE" box="[159,1405,1479,1624]" gridrow="0" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7FA7A7978F9BE" box="[159,337,1479,1624]" gridcol="0" gridrow="0" pageId="2" pageNumber="493">Gill rakers</th>
<th id="3D635D90FFBA001B1028FA7A79CDF9BE" box="[368,484,1479,1624]" gridcol="1" gridrow="0" pageId="2" pageNumber="493">absent</th>
<th id="3D635D90FFBA001B137CFA7A7B51F9BE" box="[548,888,1479,1624]" gridcol="2" gridrow="0" pageId="2" pageNumber="493">at least some no developed developed, 2– rakers, about 6 4 + 8–14 = tiny rudiments on 10–17 upper part of ceratobranchial 1</th>
<th id="3D635D90FFBA001B12CEFA7A7C2EF9BE" box="[918,1031,1479,1624]" gridcol="3" gridrow="0" pageId="2" pageNumber="493">absent, but tiny teeth present</th>
<th id="3D635D90FFBA001B156AFA7A7C8AF9BE" box="[1074,1187,1479,1624]" gridcol="4" gridrow="0" pageId="2" pageNumber="493">absent</th>
<th id="3D635D90FFBA001B1599FA7A7D54F9BE" box="[1217,1405,1479,1624]" gridcol="5" gridrow="0" pageId="2" pageNumber="493">absent</th>
</tr>
<tr id="7EB234ECFFBA001B11C7F9D67D54F959" box="[159,1405,1643,1727]" gridrow="1" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7F9D67978F959" box="[159,337,1643,1727]" gridcol="0" gridrow="1" pageId="2" pageNumber="493">Anterior and posterior ceratohyals</th>
<td id="3D635D90FFBA001B1028F9D679CDF959" box="[368,484,1643,1727]" gridcol="1" gridrow="1" pageId="2" pageNumber="493">autogenous</td>
<td id="3D635D90FFBA001B137CF9D67B51F959" box="[548,888,1643,1727]" gridcol="2" gridrow="1" pageId="2" pageNumber="493">autogenous autogenous</td>
<td id="3D635D90FFBA001B12CEF9D67C2EF959" box="[918,1031,1643,1727]" gridcol="3" gridrow="1" pageId="2" pageNumber="493">undifferen- tiated</td>
<td id="3D635D90FFBA001B156AF9D67C8AF959" box="[1074,1187,1643,1727]" gridcol="4" gridrow="1" pageId="2" pageNumber="493">undifferen- tiated</td>
<td id="3D635D90FFBA001B1599F9D67D54F959" box="[1217,1405,1643,1727]" gridcol="5" gridrow="1" pageId="2" pageNumber="493">undifferentiated</td>
</tr>
<tr id="7EB234ECFFBA001B11C7F96F7D54F8EF" box="[159,1405,1746,1801]" gridrow="2" pageId="2" pageNumber="493">
<th id="3D635D90FFBA001B11C7F96F7978F8EF" box="[159,337,1746,1801]" gridcol="0" gridrow="2" pageId="2" pageNumber="493">Proximal head on 3rd branchiostegal</th>
<td id="3D635D90FFBA001B1028F96F79CDF8EF" box="[368,484,1746,1801]" gridcol="1" gridrow="2" pageId="2" pageNumber="493">narrow</td>
<td id="3D635D90FFBA001B137CF96F7B51F8EF" box="[548,888,1746,1801]" gridcol="2" gridrow="2" pageId="2" pageNumber="493">very broad broad</td>
<td id="3D635D90FFBA001B12CEF96F7C2EF8EF" box="[918,1031,1746,1801]" gridcol="3" gridrow="2" pageId="2" pageNumber="493">narrow</td>
<td id="3D635D90FFBA001B156AF96F7C8AF8EF" box="[1074,1187,1746,1801]" gridcol="4" gridrow="2" pageId="2" pageNumber="493">narrow</td>
<td id="3D635D90FFBA001B1599F96F7D54F8EF" box="[1217,1405,1746,1801]" gridcol="5" gridrow="2" pageId="2" pageNumber="493">narrow</td>
</tr>
</table>
</paragraph>
<paragraph id="C03D36AEFFBAFFE7119FF8F37955FE82" blockId="2.[151,1436,1869,2003]" lastBlockId="3.[151,1436,151,501]" lastPageId="3" lastPageNumber="494" pageId="2" pageNumber="493">
<collectingCountry id="B895763EFFBAFFE6119FF8F378CFF880" box="[199,230,1870,1894]" name="American Samoa" pageId="2" pageNumber="493">As</collectingCountry>
percentage of SL: head length 28.3; predorsal length 39.5; prepelvic length 29.6; preanal length 60.5; first dorsal-fin origin to second dorsal-fin origin 17.1; second dorsal-fin base length 30.9; anal-fin base length 25.7; pectoral-fin base depth 6.6; first dorsal-fin origin to pelvic-fin origin 17.1; second dorsal-fin origin to anal-fin origin 13.2; snout length 5.9; orbit diameter 7.2; head width 14.5; body width 13.2; bony interorbital width 1.3; snout tip to retroarticular tip 13.2; caudal-peduncle length 15.1; caudal-peduncle depth 9.2; length of first spine of first dorsal fin 12.5; length of third spine of first dorsal fin 14.5; length of spine of second dorsal fin 9.2; length of first segmented ray of second dorsal fin 10.5; length of last segmented ray of second dorsal fin 11.8; anal-fin spine length 7.2; length of first segmented anal-fin ray 8.6; length of last segmented anal-fin ray 13.2; pectoral-fin length 21.7; fourth segmented pelvic-fin ray length 23.0; caudal-fin length 18.4, ray possibly regrown (longest mid-ray on lower hypural 23.7).
</paragraph>
<paragraph id="C03D36AEFFBBFFE7119FFED27A3BFE13" blockId="3.[151,1436,151,501]" pageId="3" pageNumber="494">Scales absent; laterosensory head pores absent; lower lip fleshy and protruding, with uninterrupted, free ventral margin; anterior naris in short tube (abnormally branched on left side); posterior naris without raised rim or membranous flap; tongue tip rounded; gill opening extending anteriorly to vertical through about midpoint between preopercle edge and eye.</paragraph>
<caption id="94FD6626FFBBFFE711CFFC0B79B7FC0C" httpUri="https://zenodo.org/record/226657/files/figure.png" pageId="3" pageNumber="494" targetBox="[170,1416,595,926]" targetPageId="3">
<paragraph id="C03D36AEFFBBFFE711CFFC0B79B7FC0C" blockId="3.[151,1436,950,1002]" pageId="3" pageNumber="494">
<emphasis id="F2F6EABCFFBBFFE711CFFC0B793CFC2A" bold="true" box="[151,277,950,972]" pageId="3" pageNumber="494">FIGURE 1.</emphasis>
<taxonomicName id="07824D2DFFBBFFE71045FC0B7A1BFC2A" box="[285,562,950,972]" class="Actinopterygii" family="Xenisthmidae" genus="Gymnoxenisthmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="494" phylum="Chordata" rank="species" species="tigrellus">
<emphasis id="F2F6EABCFFBBFFE71045FC0B7A1BFC2A" box="[285,562,950,972]" italics="true" pageId="3" pageNumber="494">Gymnoxenisthmus tigrellus</emphasis>
</taxonomicName>
, holotype, SMF 34903, 15.2 mm SL, gravid female, Farasan Archipelago, Red Sea. Photo by Sven Traenkner.
</paragraph>
</caption>
<paragraph id="C03D36AEFFBBFFE7119FFBA57CC8FBB2" blockId="3.[151,1436,1047,2008]" pageId="3" pageNumber="494">Upper jaw with 2 or 3 (anteriorly) to 2 (posteriorly) rows of slightly curved conical teeth; lower jaw with 2 or 3 (anteriorly) to 2 (posteriorly) rows of slightly curved conical teeth; vomer and palatine edentate.</paragraph>
<paragraph id="C03D36AEFFBBFFE0119FFBE27963FF33" blockId="3.[151,1436,1047,2008]" lastBlockId="4.[151,1437,151,573]" lastPageId="4" lastPageNumber="495" pageId="3" pageNumber="494">
Live coloration (based on colour photograph of
<typeStatus id="1F39880CFFBBFFE71253FBE27B44FB9E" box="[779,877,1119,1144]" pageId="3" pageNumber="494" type="holotype">holotype</typeStatus>
when freshly dead;
<figureCitation id="58B92A2BFFBBFFE71532FBE27CE7FB9E" box="[1130,1230,1119,1144]" captionStart="FIGURE 1" captionStartId="3.[151,250,950,972]" captionTargetBox="[170,1416,595,926]" captionTargetId="figure@3.[170,1417,595,926]" captionTargetPageId="3" captionText="FIGURE 1. Gymnoxenisthmus tigrellus, holotype, SMF 34903, 15.2 mm SL, gravid female, Farasan Archipelago, Red Sea. Photo by Sven Traenkner." httpUri="https://zenodo.org/record/226657/files/figure.png" pageId="3" pageNumber="494">Figure 1</figureCitation>
): head and body translucent bluish grey; body with fourteen equally spaced, mid-lateral orange markings (first a spot just anterior to pectoral-fin base; second a short bar just behind pectoral-fin base; third a chevron below anterior part of first dorsal fin; fourth a chevron below posterior edge of first dorsal fin; fifth a chevron through space between dorsal fins; sixth a chevron through first segmented ray of second dorsal fin; seventh a bar through third segmented ray of second dorsal fin; eighth a bar through fifth segmented ray of second dorsal fin; ninth a bar through seventh and eighth segmented rays of second dorsal; tenth a bar through tenth segmented ray of second dorsal fin; eleventh through base of last second dorsal-fin ray; twelfth and thirteenth bars slightly oblique and less distinct, through caudal peduncle; fourteenth a narrow, short bar along posterior edge of hypurals); small, indistinct pale orange or yellow spots present between most midlateral orange bars and spots; scattered melanophores present on head and body, mostly confined to orange areas; head with orange stripe extending from mid-upper part of upper lip to upper half of eye, then from behind eye to point above upper edge of preopercle, with large (almost pupil-sized) orange spot near edge of opercle; second, oblique orange stripe extending from just behind and below eye to middle of operculum; isolated orange spot on anterior part of operculum, between two orange stripes; lower lip orange; orange “L”-shaped marking on cheek extending from just below anterior margin of eye, with bottom of “L” extending along lower cheek edge; iris mostly orange on dorsal two-thirds, remainder pale yellow to pale gold; pectoral-fin base orange anteriorly and dorsally, with two pale orange spots, one on mid-upper and the other on mid-lower part of fin base; first dorsal fin with first spine base orange; remainder of fin translucent on outer third, followed proximally with silvery white stripe, orange stripe, and silvery white basal stripe; scattered melanophores on first dorsal, these densest over orange stripe; second dorsal fin orange (basally) to dusky orange (distally), with narrow (anteriorly) to broad (posteriorly) distal margin translucent; orange bars from body extending on to fin base; silvery white stripe through middle of fin, with a basal series of short, silvery white oblique bars that extend anterodorsally from between orange bars from body; a few scattered silvery white spots present on translucent part of fin; anal fin translucent with narrow indistinct dusky orange stripe through basal third to half of fin, indistinctly bordered basally with silvery white spots (anteriorly) and stripe (posteriorly); caudal fin mostly translucent, with two large indistinct pale grey-orange spots on basal part of fin, one dorsal and the other ventral; pectoral and pelvic fins translucent.
</paragraph>
<paragraph id="C03D36AEFFBCFFE0119FFF627973FE6E" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="495">Preserved coloration: head and body generally pale beige, greyish brown on lower abdomen; melanophores within orange bars on body remain, though indistinct and confined to midside and upper half of body; melanophores within orange areas on lips, upper stripe on head and upper half of pectoral-fin base remain; orange markings on first dorsal fin become dark grey; orange markings on second dorsal and anal fins remain, becoming dark grey-brown.</paragraph>
</subSubSection>
<subSubSection id="88986525FFBCFFE0119FFE297BBFFDDA" pageId="4" pageNumber="495" type="etymology">
<paragraph id="C03D36AEFFBCFFE0119FFE297D7AFE36" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="495">
<emphasis id="F2F6EABCFFBCFFE0119FFE297967FE4A" bold="true" box="[199,334,404,428]" pageId="4" pageNumber="495">Etymology.</emphasis>
The specific epithet is from the Latin, meaning a little tiger, alludes to the orange bars on the body. The name was selected by school children at the Australian Museum Science Festival Expo in
<date id="B43C106EFFBCFFE015E4FE057D66FE36" box="[1212,1359,440,464]" pageId="4" pageNumber="495" value="2013-08">August 2013</date>
.
</paragraph>
<paragraph id="C03D36AEFFBCFFE0119FFE667BBFFDDA" blockId="4.[151,1437,151,573]" pageId="4" pageNumber="495">
<emphasis id="F2F6EABCFFBCFFE0119FFE667903FE12" bold="true" box="[199,298,475,500]" pageId="4" pageNumber="495">Habitat.</emphasis>
The
<typeStatus id="1F39880CFFBCFFE0103DFE6179EEFE13" box="[357,455,476,501]" pageId="4" pageNumber="495" type="holotype">holotype</typeStatus>
was collected from an unnamed rocky island with a narrow reef flat, and a slope with patches of corals and a rocky wall of about
<quantity id="077A9B4BFFBCFFE01320FDBD7AB5FDFE" box="[632,668,512,536]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="495" unit="m" value="3.0">3m</quantity>
with small caves and shelters. The sandy slope began at depths of
<quantity id="077A9B4BFFBCFFE014D8FDBD78FAFDDA" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="4" pageNumber="495" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8– 10 m</quantity>
; the
<typeStatus id="1F39880CFFBCFFE01050FD997943FDDB" box="[264,362,548,573]" pageId="4" pageNumber="495" type="holotype">holotype</typeStatus>
was collected on sand at the base of coral in
<specimenCount id="D684FD27FFBCFFE0123CFD997BBFFDDA" box="[868,918,548,572]" pageId="4" pageNumber="495" type="generic">
<quantity id="077A9B4BFFBCFFE0123CFD997BBBFDDA" box="[868,914,548,572]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="495" unit="m" value="8.0">8 m</quantity>
.
</specimenCount>
</paragraph>
</subSubSection>
</treatment>
</document>
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