122 lines
14 KiB
XML
122 lines
14 KiB
XML
<document id="01CAE8F74913A2A1B8812A6270613114" ID="10.11646/zootaxa.4178.3.5" ID-DOI="10.11646/zootaxa.4178.3.5" ID-GBIF-Dataset="fb113551-41a5-40e7-80fd-ce05a4dfcc00" ID-ISSN="1175-5326" ID-Zenodo-Dep="272503" ID-ZooBank="51B86192-BF51-444F-8C00-960C10E8DF5C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1477469536949" checkinUser="plazi" docAuthor="James, Samuel W. & Gamiette, Franciane" docDate="2016" docId="440C87ADFF9EC958FF19C399FCAEF8E9" docLanguage="en" docName="zootaxa.4178.3.5.pdf" docOrigin="Zootaxa 4178 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichogaster" docType="treatment" docVersion="8" lastPageNumber="399" masterDocId="B835FFD5FF96C950FF8EC12DFF91FFEA" masterDocTitle="New species of Dichogaster Beddard, 1888 (Clitellata: Benhamiidae) with additional records of earthworms from Guadeloupe (French West Indies)" masterLastPageNumber="408" masterPageNumber="391" pageNumber="399" updateTime="1698390840751" updateUser="plazi">
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<mods:mods id="066569821296A9E9A7290F248C36A0B1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D8FF57AEBE506A426B2412186E20D303">
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<mods:title id="F2273960908E94D3BCA58DC644D95B44">New species of Dichogaster Beddard, 1888 (Clitellata: Benhamiidae) with additional records of earthworms from Guadeloupe (French West Indies)</mods:title>
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</mods:titleInfo>
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<mods:name id="06F1EC5608E0FFB7301D9D44A877BFC9" type="personal">
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<mods:roleTerm id="6B0A3F117012D3B6A3511B262202BF37">Author</mods:roleTerm>
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<mods:namePart id="6EB16BB52E38C6D9BC8B20E30982C9B4">James, Samuel W.</mods:namePart>
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</mods:name>
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<mods:name id="10D1CC7262911926F87BAED8FE0E2788" type="personal">
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<mods:role id="50D16D407C5533347E680A52B4379E38">
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<mods:roleTerm id="2FE223BE360123FB6EF2F9D9C1401032">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6CB619FE3D276C0C16DE5FA60ACFD98F">Gamiette, Franciane</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="5DFF5FAA38C4803C84C7B54F53B81B1B">text</mods:typeOfResource>
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<mods:titleInfo id="A02C92A40AFC122B66337320099E3530">
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<mods:title id="4A0EFA68A55D0051DDDFFD32BC5D19FE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="47EA0F9873BBD0945655FFBE3732EF66">
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<mods:date id="8A5F3DDE39A99669A97EED50CCD8F912">2016</mods:date>
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<mods:detail id="C97FB8422768EA370E501DBD153F511A" type="volume">
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<mods:number id="8A0BC2F61489BD78934C4E8ABE7A9FF6">4178</mods:number>
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</mods:detail>
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<mods:detail id="F854B9FD345E67624FB19433FB1044D1" type="issue">
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<mods:number id="3721EAA908365E58176DFF4A48582496">3</mods:number>
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<mods:start id="22B92731EED69D41CF3D315ED194E65F">391</mods:start>
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<mods:end id="63CDF2B6544CE3AF9CC25ECAED6309A1">408</mods:end>
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<mods:classification id="ED5F5C8682744018E2EE6CA02AD3115A">journal article</mods:classification>
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<mods:identifier id="1B56A11E11F3BF50CDFF93751AB1B80B" type="DOI">10.11646/zootaxa.4178.3.5</mods:identifier>
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<mods:identifier id="EF1696DB6CB1845D5BFAC3DEC69D6732" type="GBIF-Dataset">fb113551-41a5-40e7-80fd-ce05a4dfcc00</mods:identifier>
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<mods:identifier id="092F26ABB2E1EF9DAFDE51BC2B665561" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="27F4A025E32C95C43566550415B98F1D" type="Zenodo-Dep">272503</mods:identifier>
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<mods:identifier id="A8A98D2B1B4C5333F4C4354601223D3E" type="ZooBank">51B86192-BF51-444F-8C00-960C10E8DF5C</mods:identifier>
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<treatment id="440C87ADFF9EC958FF19C399FCAEF8E9" ID-DOI="http://doi.org/10.5281/zenodo.6070208" ID-GBIF-Taxon="124558163" ID-Zenodo-Dep="6070208" LSID="urn:lsid:plazi:treatment:440C87ADFF9EC958FF19C399FCAEF8E9" httpUri="http://treatment.plazi.org/id/440C87ADFF9EC958FF19C399FCAEF8E9" lastPageNumber="399" pageId="8" pageNumber="399">
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<subSubSection id="84BF6530FF9EC958FF19C399FEFBFD24" box="[151,362,692,718]" pageId="8" pageNumber="399" type="nomenclature">
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<paragraph id="CC1A36BBFF9EC958FF19C399FEFBFD24" blockId="8.[151,362,692,718]" box="[151,362,692,718]" pageId="8" pageNumber="399">
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<heading id="975281D7FF9EC958FF19C399FEFBFD24" bold="true" box="[151,362,692,718]" fontSize="11" level="1" pageId="8" pageNumber="399" reason="1">
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<emphasis id="FED1EAA9FF9EC958FF19C399FEFBFD24" bold="true" box="[151,362,692,718]" pageId="8" pageNumber="399">
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<taxonomicName id="0BA54D38FF9EC958FF19C399FEBBFD24" box="[151,298,692,718]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="399" phylum="Annelida" rank="genus">
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<emphasis id="FED1EAA9FF9EC958FF19C399FEBBFD24" bold="true" box="[151,298,692,718]" italics="true" pageId="8" pageNumber="399">Dichogaster</emphasis>
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</taxonomicName>
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sp. 5
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="84BF6530FF9EC958FF19C3D7FCFEFCDD" pageId="8" pageNumber="399" type="materials_examined">
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<paragraph id="CC1A36BBFF9EC958FF19C3D7FCFEFCDD" blockId="8.[151,1437,762,1687]" pageId="8" pageNumber="399">
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<materialsCitation id="7CCD3CE6FF9EC958FF19C3D7FCFEFCDD" ID-GBIF-Occurrence="1324200685" country="Guadeloupe" county="Basse Terre" elevation="629" latitude="15.974934" location="Material" longLatPrecision="1" longitude="-61.687717" municipality="Vent Souffle" pageId="8" pageNumber="399" specimenCount="1">
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<emphasis id="FED1EAA9FF9EC958FF19C3D7FE13FCF9" bold="true" box="[151,386,762,787]" pageId="8" pageNumber="399">
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<location id="C97A6060FF9EC958FF19C3D7FF6EFCF9" LSID="urn:lsid:plazi:treatment:440C87ADFF9EC958FF19C399FCAEF8E9:C97A6060FF9EC958FF19C3D7FF6EFCF9" box="[151,255,762,787]" country="Guadeloupe" county="Basse Terre" latitude="15.974934" longLatPrecision="1" longitude="-61.687717" municipality="Vent Souffle" name="Material" pageId="8" pageNumber="399">Material</location>
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examined.
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</emphasis>
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3 late juveniles (one in alcohol),
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<location id="C97A6060FF9EC958FC88C3D6FC29FCF9" LSID="urn:lsid:plazi:treatment:440C87ADFF9EC958FF19C399FCAEF8E9:C97A6060FF9EC958FC88C3D6FC29FCF9" box="[774,952,762,787]" country="Guadeloupe" county="Basse Terre" latitude="15.974934" longLatPrecision="1" longitude="-61.687717" municipality="Vent Souffle" name="Monts Caraibes" pageId="8" pageNumber="399">Monts Caraibes</location>
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trail,
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<collectingMunicipality id="2C7EACC1FF9EC958FC70C3D6FB1DFCF9" box="[1022,1164,762,787]" pageId="8" pageNumber="399">Vent Soufflé</collectingMunicipality>
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in bromeliad leaf axils,
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<collectingCounty id="257B4E37FF9EC958FF19C232FE8CFCDC" box="[151,285,799,822]" pageId="8" pageNumber="399">Basse Terre</collectingCounty>
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,
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<collectingCountry id="B4B2762BFF9EC958FEA6C233FE23FCDD" box="[296,434,798,823]" name="Guadeloupe" pageId="8" pageNumber="399">Guadeloupe</collectingCountry>
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,
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<geoCoordinate id="A991507CFF9EC958FE33C233FDDEFCDD" box="[445,591,798,823]" direction="north" orientation="latitude" pageId="8" pageNumber="399" precision="1" value="15.974934">N15°58.496'</geoCoordinate>
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,
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<geoCoordinate id="A991507CFF9EC958FDD6C233FD61FCDD" box="[600,752,798,823]" direction="west" orientation="longitude" pageId="8" pageNumber="399" precision="1" value="-61.687717">W61°41.263'</geoCoordinate>
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,
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<elevation id="4788D188FF9EC958FD74C233FCD0FCDC" box="[762,833,798,823]" metricMagnitude="2" metricUnit="m" metricValue="6.29" pageId="8" pageNumber="399" unit="m" value="629.0">
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<quantity id="0B5D9B5EFF9EC958FD74C233FCD0FCDC" box="[762,833,798,823]" metricMagnitude="2" metricUnit="m" metricValue="6.29" pageId="8" pageNumber="399" unit="m" value="629.0">629 m</quantity>
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</elevation>
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asl.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="84BF6530FF9EC958FF49C26FFA15F9ED" pageId="8" pageNumber="399" type="description">
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<paragraph id="CC1A36BBFF9EC958FF49C26FFB99FBD9" blockId="8.[151,1437,762,1687]" pageId="8" pageNumber="399">
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<emphasis id="FED1EAA9FF9EC958FF49C26FFEC8FCB1" bold="true" box="[199,345,834,859]" pageId="8" pageNumber="399">Description.</emphasis>
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Dimensions
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<quantity id="0B5D9B5EFF9EC958FE63C26FFDF7FCB0" box="[493,614,834,859]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="8" pageNumber="399" unit="mm" value="43.5" valueMax="45.0" valueMin="42.0">42–45 mm</quantity>
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by
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<quantity id="0B5D9B5EFF9EC958FD1FC26EFC95FCB0" box="[657,772,835,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="8" pageNumber="399" unit="mm" value="2.1" valueMax="2.2" valueMin="2.0">2–2.2 mm</quantity>
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at segment x,
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<quantity id="0B5D9B5EFF9EC958FC26C26FFC6EFCB0" box="[936,1023,834,859]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="8" pageNumber="399" unit="mm" value="2.5">2.5 mm</quantity>
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at xxx; body cylindrical throughout, segments>102; one amputee at 102, one regenerate from 81 plus 44 regenerated segments. Setae all ventral, closely paired throughout; setal formula AA:AB:BC:CD= 4:1:4:1 at xxx; DD = 2/3 circumference. Prostomium schizolobous, furrow 1/2 obliterated; light olive brown dorsal pigmentation darker anteriorly; residual green color on first 3 or 4 segments in formalin preservation. First dorsal pore 10/11, 11/12; spermathecal pores not seen. Ovipores equatorial median to A in xiv; male pores not seen; prostatic pores at ends of straight grooves in A in xvii–xix; grooves straight. Clitellum not developed, no other genital markings.
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</paragraph>
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<paragraph id="CC1A36BBFF9EC958FF49C512FA98FA71" blockId="8.[151,1437,762,1687]" pageId="8" pageNumber="399">
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Septa
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<date id="B81B107BFF9EC958FE83C512FEEFFBBD" box="[269,382,1087,1111]" pageId="8" pageNumber="399" value="2014-08-09" valueMax="2014-08-13" valueMin="2014-08-09">8/9–13/14</date>
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slightly muscular, maximal at 11/12; septa 10/11, 11/12 fused at body wall. Alimentary canal with proventriculus in v, two long gizzards in vi, vii; esophagus valvular in xviii, intestinal origin xix; deeply sacculated xix–xxvi; typhlosole a simple thick fold originating in xxv, height one fifth lumen diameter, continuous very low lateral typhlosoles 25/26–xxx, typhlosole ends> xcii, or in cxi in regenerated segments. Calciferous glands reniform, paired in xv–xvii, composed of longitudinally-oriented lamellae opening to medial lumen; each gland with separate duct to esophagus; glands xv<xvi≤xvii. Micronephridia 10 per segment, pre-intestinal nephridia tubular; intestinal segment nephridia closed sac-shaped with oil droplets; median rows of posterior segments with nephrostomes; median row less mantled and with 3 or 4 lobes; no nephrostomes seen anteriorly; nephridia distributed AB-dorsum. Vascular system with ventral trunk, single dorsal trunk, these connected by commisural vessels in viii, ix, latero-esophageal hearts in x–xii. Commissural vessel of vii to gizzard.
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</paragraph>
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<paragraph id="CC1A36BBFF9EC958FF49C48BFA15F9ED" blockId="8.[151,1437,762,1687]" pageId="8" pageNumber="399">
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Ovaries in xiii; paired spermathecae in viii, ix, juvenile state showing spherical ampulla and small spherical sessile diverticulum. Male sexual system juvenile, testes of x in sub-esophageal sacs, formed by fusion of septa 10/ 11, 11/
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<quantity id="0B5D9B5EFF9EC958FF6DC4C2FE8AF9ED" box="[227,283,1518,1543]" metricMagnitude="-1" metricUnit="m" metricValue="3.048" pageId="8" pageNumber="399" unit="in" value="12.0">12 in</quantity>
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xi; very small tubular prostates in short zig-zags; penial setal follicles and penial setae undeveloped.
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</paragraph>
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</subSubSection>
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<subSubSection id="84BF6530FF9EC958FF49C73FFCAEF8E9" pageId="8" pageNumber="399" type="discussion">
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<paragraph id="CC1A36BBFF9EC958FF49C73FFC56F97D" blockId="8.[151,1437,762,1687]" pageId="8" pageNumber="399">
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<emphasis id="FED1EAA9FF9EC958FF49C73FFEA5F9C1" bold="true" box="[199,308,1554,1579]" pageId="8" pageNumber="399">Remarks</emphasis>
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. Although not quite adult, the combination of pigmentation, number of nephridia, shape of prostomium and extent of the lateral typhlosoles distinguishes this species from the other acaecate Guadeloupe
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<taxonomicName id="0BA54D38FF9EC958FF19C776FEB1F999" box="[151,288,1627,1651]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="399" phylum="Annelida" rank="genus">
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<emphasis id="FED1EAA9FF9EC958FF19C776FEB1F999" box="[151,288,1627,1651]" italics="true" pageId="8" pageNumber="399">Dichogaster</emphasis>
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</taxonomicName>
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.
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<taxonomicName id="0BA54D38FF9EC958FEA3C776FE29F999" box="[301,440,1627,1651]" class="Clitellata" family="Octochaetidae" genus="Dichogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Haplotaxida" pageId="8" pageNumber="399" phylum="Annelida" rank="genus">
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<emphasis id="FED1EAA9FF9EC958FEA3C776FE29F999" box="[301,440,1627,1651]" italics="true" pageId="8" pageNumber="399">Dichogaster</emphasis>
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</taxonomicName>
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sp. 4 has the same number of nephridia, but it is unpigmented, epilobous, the typhlosoles commence more anteriorly and it lives in soil as an endogeic earthworm.
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</paragraph>
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<paragraph id="CC1A36BBFF9EC958FF49C7EBFCAEF8E9" blockId="8.[151,1437,1734,1795]" pageId="8" pageNumber="399">The following new records all are based on adult material except where noted. The number in parentheses preceding the latitude is the number of individuals collected.</paragraph>
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</subSubSection>
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</treatment>
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</document> |