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<document id="10D6323ED99F05CAFFF4DCFFB9A31D56" ID-DOI="10.3853/j.2201-4349.74.2022.1821" ID-ISSN="2201-4349" ID-Zenodo-Dep="13228969" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722893937960" checkinUser="felipe" docAuthor="Reid, Chris A. M. &amp; Runagall-Mcnaull, Aidan" docDate="2022" docId="424B87F8EC55507DFF69B26FFB66FE01" docLanguage="en" docName="RecAustMus.74.5.131150.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 74 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.74.2022.1821" docStyle="DocumentStyle:997BA24B490466A2F99594A367FA147F.4:RecAustMus.2019-.journal_article.0cover" docStyleId="997BA24B490466A2F99594A367FA147F" docStyleName="RecAustMus.2019-.journal_article.0cover" docStyleVersion="4" docTitle="Onthophagus squalidus Lea 1923" docType="treatment" docVersion="2" lastPageNumber="147" masterDocId="BE72FF80EC5A506DFFD1B023FFBCFF9B" masterDocTitle="A Revision of the Onthophagus pexatus Species-group, with Description of a New Brachypterous Species (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="150" masterPageNumber="131" pageNumber="146" updateTime="1722901422291" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="2CB7BE92463BC4169FA80F24B3099993">A Revision of the Onthophagus pexatus Species-group, with Description of a New Brachypterous Species (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:namePart id="35E62487C53D55841476D0C2049C75A4">Reid, Chris A. M.</mods:namePart>
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<treatment id="424B87F8EC55507DFF69B26FFB66FE01" ID-DOI="http://doi.org/10.5281/zenodo.13229005" ID-Zenodo-Dep="13229005" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01" httpUri="http://treatment.plazi.org/id/424B87F8EC55507DFF69B26FFB66FE01" lastPageId="16" lastPageNumber="147" pageId="15" pageNumber="146">
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<paragraph id="CA5D36EEEC555062FF69B26FFD12FDF7" blockId="15.[184,686,588,620]" box="[184,686,588,620]" pageId="15" pageNumber="146">
<heading id="91158182EC555062FF69B26FFD12FDF7" bold="true" box="[184,686,588,620]" centered="true" fontSize="12" level="2" pageId="15" pageNumber="146" reason="2">
<taxonomicName id="0DE24D6DEC555062FF69B26FFD12FDF7" ID-CoL="7XP8L" authority="Lea, 1923" authorityName="Lea" authorityYear="1923" box="[184,686,588,620]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="146" phylum="Arthropoda" rank="species" species="squalidus">
<emphasis id="F896EAFCEC555062FF69B26FFD12FDF7" bold="true" box="[184,686,588,620]" pageId="15" pageNumber="146">
<emphasis id="F896EAFCEC555062FF69B26FFDAFFDF0" bold="true" box="[184,531,588,619]" italics="true" pageId="15" pageNumber="146">Onthophagus squalidus</emphasis>
<bibRefCitation id="AE734B1FEC555062FDCCB26EFD12FDF7" author="Lea, A. M." box="[541,686,589,620]" pageId="15" pageNumber="146" pagination="353 - 396" refId="ref13145" refString="Lea, A. M. 1923. Australian dung beetles of the subfamily Coprides. Records of the South Australian Museum 2: 353 - 396." type="journal article" year="1923">Lea, 1923</bibRefCitation>
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<subSubSection id="82F86565EC555062FF21B2A5FDCAFD3A" box="[240,630,646,673]" pageId="15" pageNumber="146" type="description">
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<figureCitation id="52D92A6BEC555062FF21B2A5FED6FD3B" box="[240,362,646,672]" captionStart="Figures 9, 10" captionStartId="6.[447,524,1048,1070]" captionTargetBox="[225,1362,170,1001]" captionTargetId="figure-496@6.[166,1466,148,1025]" captionTargetPageId="6" captionText="Figures 9, 10. Onthophagus squalidus Lea, male dorsal, lateral, and ventral." figureDoi="http://doi.org/10.5281/zenodo.13228975" httpUri="https://zenodo.org/record/13228975/files/figure.png" pageId="15" pageNumber="146">Figs 910</figureCitation>
,
<figureCitation id="52D92A6BEC555062FEA7B2A5FE78FD3B" box="[374,452,646,672]" captionStart="Figures 2124" captionStartId="10.[166,243,1231,1253]" captionTargetBox="[166,1466,149,1195]" captionTargetId="figure-11@10.[166,1466,149,1219]" captionTargetPageId="10" captionText="Figures 2124. Onthophagus species, head, male (left), and female (right), fresh specimens. (21, 22) O. pexatus Harold; (23, 24) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228985" httpUri="https://zenodo.org/record/13228985/files/figure.png" pageId="15" pageNumber="146">2324</figureCitation>
,
<figureCitation id="52D92A6BEC555062FE01B2A5FE4CFD3B" box="[464,496,646,672]" captionStart="Figures 2529" captionStartId="11.[121,198,1565,1587]" captionTargetBox="[121,748,151,1551]" captionTargetId="figure-106@11.[121,748,151,1553]" captionTargetPageId="11" captionText="Figures 2529. Onthophagus species, venter of right metafemur. (25) O. bulga sp. nov.; (26) O. longipes Paulian; (27) O. nammuldi Matthews; (28) O. pexatus Harold; (29) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228987" httpUri="https://zenodo.org/record/13228987/files/figure.png" pageId="15" pageNumber="146">29</figureCitation>
,
<figureCitation id="52D92A6BEC555062FE2FB2A5FDA2FD3B" box="[510,542,646,672]" captionStart="Figures 3034" captionStartId="12.[167,244,762,784]" captionTargetBox="[178,1466,144,748]" captionTargetId="figure-10@12.[166,1466,130,748]" captionTargetPageId="12" captionText="Figures 3034. Onthophagus species, left protibia. (30) O. bulga sp. nov.; (31) O. longipes Paulian; (32) O. nammuldi Matthews; (33) O. pexatus Harold; (34) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228991" httpUri="https://zenodo.org/record/13228991/files/figure.png" pageId="15" pageNumber="146">34</figureCitation>
,
<figureCitation id="52D92A6BEC555062FDFAB2A5FDF7FD3B" box="[555,587,646,672]" captionStart="Figures 3741" captionStartId="13.[819,896,1603,1625]" captionTargetBox="[121,1422,151,1679]" captionTargetId="figure-10@13.[121,1421,151,1679]" captionTargetPageId="13" captionText="Figures 3741. Onthophagus species, pygidium.(37) O. bulga sp. nov.; (38) O. longipes Paulian; (39) O. nammuldi Matthews; (40) O. pexatus Harold; (41) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228995" httpUri="https://zenodo.org/record/13228995/files/figure.png" pageId="15" pageNumber="146">41</figureCitation>
,
<figureCitation id="52D92A6BEC555062FD89B2A5FDCAFD3A" box="[600,630,646,673]" captionStart="Figures 4248" captionStartId="14.[167,244,1399,1421]" captionTargetBox="[167,1459,164,1379]" captionTargetId="figure-65@14.[166,1465,151,1381]" captionTargetPageId="14" captionText="Figures 4248. Onthophagus species, aedeagus, lateral. (42) O. bulga sp. nov.; (43) O. longipes Paulian; (44) O. nammuldi Matthews; (45) O. squalidus Lea; (46) O. pexatus, Dairy Park, NSW; (47) O. pexatus, Wombeyan Caves, NSW; (48) O. pexatus, Woolomin, NSW." figureDoi="http://doi.org/10.5281/zenodo.13228999" httpUri="https://zenodo.org/record/13228999/files/figure.png" pageId="15" pageNumber="146">45</figureCitation>
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<paragraph id="CA5D36EEEC555062FF46B297FD00FD65" blockId="15.[151,719,692,767]" pageId="15" pageNumber="146">
<treatmentCitationGroup id="EAF211C0EC555062FF46B297FD00FD65" pageId="15" pageNumber="146">
<treatmentCitation id="4B4310FFEC555062FF46B297FDB8FD52" author="Lea, A. M." box="[151,516,692,713]" page="390" pageId="15" pageNumber="146" year="1923">
<taxonomicName id="0DE24D6DEC555062FF46B297FDB8FD52" ID-CoL="7XP8L" authority="Lea, 1923: 390" authorityName="Lea" authorityPageNumber="390" authorityYear="1923" box="[151,516,692,713]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="146" phylum="Arthropoda" rank="species" species="squalidus">
<emphasis id="F896EAFCEC555062FF46B297FEC8FD52" box="[151,372,692,713]" italics="true" pageId="15" pageNumber="146">Onthophagus squalidus</emphasis>
<bibRefCitation id="AE734B1FEC555062FEA6B297FDB8FD52" author="Lea, A. M." box="[375,516,692,713]" pageId="15" pageNumber="146" pagination="353 - 396" refId="ref13145" refString="Lea, A. M. 1923. Australian dung beetles of the subfamily Coprides. Records of the South Australian Museum 2: 353 - 396." type="journal article" year="1923">Lea, 1923: 390</bibRefCitation>
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; Boucomont &amp; Gillet, 1927: 216;
<treatmentCitation id="4B4310FFEC555062FEF4B2ECFE48FD7F" author="Matthews, E." box="[293,500,719,741]" page="211" pageId="15" pageNumber="146" year="1972">
<bibRefCitation id="AE734B1FEC555062FEF4B2ECFE48FD7F" author="Matthews, E." box="[293,500,719,741]" pageId="15" pageNumber="146" pagination="3 - 330" refId="ref13173" refString="Matthews, E. 1972. A revision of the scarabaeine dung beetles of Australia. I. tribe Onthophagini. Australian Journal of Zoology. Supplementary Series 9: 3 - 330. https: // doi. org / 10.1071 / AJZS 009" type="book chapter" year="1972">Matthews, 1972: 211</bibRefCitation>
</treatmentCitation>
; Cassis &amp; Weir, 1992: 151; Monteith &amp; Kenyon, 2011: 57; Reid
<emphasis id="F896EAFCEC555062FD98B2C9FDCFFD65" box="[585,627,745,767]" italics="true" pageId="15" pageNumber="146">et al</emphasis>
., 2022.
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</paragraph>
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(
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,
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).
<emphasis id="F896EAFCEC555062FE58B302FD97FCAF" bold="true" box="[393,555,800,820]" pageId="15" pageNumber="146">
<collectingRegion id="0826F80CEC555062FE58B302FD97FCAF" box="[393,555,800,820]" country="Australia" name="New South Wales" pageId="15" pageNumber="146">New South Wales</collectingRegion>
</emphasis>
:
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<specimenCount id="DCE4FD67EC555062FDE6B302FDEDFCAF" box="[567,593,801,820]" count="2" pageId="15" pageNumber="146" type="male">2♂</specimenCount>
/
<collectorName id="67175338EC555062FD8EB302FF28FCD7" pageId="15" pageNumber="146">Boonoo Boonoo NP</collectorName>
,
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, burnt tall dry scler., macropod dung baited
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&amp; Runagall-McNaull (
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)
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;
<materialsCitation id="7A8A3CB3EC555062FF6EB34AFDD5FC0F" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="Boorook SF &amp; M Gray G Cassis" elevation="900" latitude="-28.816668" location="Survey" longLatPrecision="1230" longitude="152.18333" pageId="15" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC555062FF6EB34AFF64FCE7" box="[191,216,873,892]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
*/
<collectorName id="67175338EC555062FF3EB34BFEEAFCE7" box="[239,342,872,892]" pageId="15" pageNumber="146">Boorook SF</collectorName>
, 09AM
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E of main rd
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(NPWS
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<collectorName id="67175338EC555062FE47B3A2FD9AFC08" box="[406,550,896,916]" pageId="15" pageNumber="146">M Gray G Cassis</collectorName>
(
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)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FDA1B3A2FE54FC5F" collectingDate="1999-03-17" collectionCode="AMS" collectorName="Boyne SF &amp; Windywoppa &amp; Lookout Rds &amp; Tarnawski &amp; Lessau" latitude="-35.607777" longLatPrecision="19" longitude="150.20778" pageId="15" pageNumber="146" specimenCount="2" specimenCount-female="2" stateProvince="New South Wales">
<specimenCount id="DCE4FD67EC555062FDA1B3A2FD35FC0F" box="[624,649,897,916]" count="2" pageId="15" pageNumber="146" type="female">2♀</specimenCount>
/
<collectorName id="67175338EC555062FD44B3A2FD55FC0F" box="[661,745,897,916]" pageId="15" pageNumber="146">Boyne SF</collectorName>
, jn
<collectorName id="67175338EC555062FF5FB3BBFEBCFC30" box="[142,256,920,939]" pageId="15" pageNumber="146">Windywoppa</collectorName>
&amp;
<collectorName id="67175338EC555062FEC8B3BBFE35FC30" box="[281,393,920,939]" pageId="15" pageNumber="146">Lookout Rds</collectorName>
,
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, CBCR003-048
<date id="BE5C102EEC555062FFA8B393FF69FC58" box="[121,213,944,963]" pageId="15" pageNumber="146" value="1999-03-17">
<collectingDate id="AE18E9C6EC555062FFA8B393FF69FC58" box="[121,213,944,963]" pageId="15" pageNumber="146" value="1999-03-17">17.iii.1999</collectingDate>
</date>
J
<collectorName id="67175338EC555062FF36B393FEFCFC58" box="[231,320,944,963]" pageId="15" pageNumber="146">Tarnawski</collectorName>
&amp; S
<collectorName id="67175338EC555062FEB8B392FE18FC5F" box="[361,420,945,964]" pageId="15" pageNumber="146">Lessau</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FE7EB392FE5DFC5F" box="[431,481,945,964]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FE21B392FE7FFC6F" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="Carrai SF &amp; M Gray G Cassis" elevation="670" latitude="-30.916666" location="Survey" longLatPrecision="1219" longitude="152.38333" municipality="Fire Trail" pageId="15" pageNumber="146" specimenCount="1" specimenCount-female="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FE21B392FDBBFC5F" box="[496,519,945,964]" count="1" pageId="15" pageNumber="146" type="female">1♀</specimenCount>
/
<collectorName id="67175338EC555062FDC4B393FDDBFC5F" box="[533,615,944,964]" pageId="15" pageNumber="146">Carrai SF</collectorName>
38AR
<collectingRegion id="0826F80CEC555062FD72B393FD79FC58" box="[675,709,944,963]" country="United Kingdom" name="Fife" pageId="15" pageNumber="146">Fife</collectingRegion>
<collectingMunicipality id="2A39AC94EC555062FD1BB393FF18FC40" pageId="15" pageNumber="146">Fire Trail</collectingMunicipality>
,
<locationDeviation id="DBEF2FCCEC555062FF7AB3EBFE37FC40" box="[171,395,968,988]" location="Fifies Knob Rd" pageId="15" pageNumber="146">
<quantity id="0D1A9B0BEC555062FF7AB3EBFF59FC40" box="[171,229,968,987]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="15" pageNumber="146" unit="km" value="1.6">1.6 km</quantity>
NE Fifies Knob Rd
</locationDeviation>
<geoCoordinate id="AFD65029EC555062FE5EB3EBFE6EFC40" box="[399,466,968,987]" degrees="30" direction="south" minutes="55" orientation="latitude" pageId="15" pageNumber="146" precision="925" value="-30.916666">30°55'S</geoCoordinate>
<geoCoordinate id="AFD65029EC555062FE07B3EBFD99FC40" box="[470,549,968,987]" degrees="152" direction="east" minutes="23" orientation="longitude" pageId="15" pageNumber="146" precision="925" value="152.38333">152°23'E</geoCoordinate>
<quantity id="0D1A9B0BEC555062FDF8B3EBFDE2FC47" box="[553,606,968,988]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="15" pageNumber="146" unit="m" value="670.0">
<elevation id="41CFD1DDEC555062FDF8B3EBFDE2FC47" box="[553,606,968,988]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="15" pageNumber="146" unit="m" value="670.0">670 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC555062FD78B3EAFD5BFC47" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC555062FD78B3EAFD5BFC47" box="[681,743,969,988]" latitude="-30.916666" longLatPrecision="1219" longitude="152.38333" municipality="Fire Trail" name="Survey" pageId="15" pageNumber="146" stateProvince="Fife">Survey</location>
)
<date id="BE5C102EEC555062FFA8B3C3FF51FC68" box="[121,237,992,1011]" pageId="15" pageNumber="146" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC555062FFA8B3C3FF51FC68" box="[121,237,992,1011]" pageId="15" pageNumber="146" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC555062FF23B3C2FE3CFC68" box="[242,384,992,1012]" pageId="15" pageNumber="146">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FE5BB3C2FE00FC6F" box="[394,444,993,1012]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FE1BB3C2FEB7FBA7" collectingDate="2021-02-22" collectingDateMax="2021-02-23" collectingDateMin="2021-02-22" collectionCode="AMS" collectorName="Cathedral Rock NP &amp; Reid" elevation="1347" latitude="-30.4416" longLatPrecision="6" longitude="152.279" pageId="15" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FE1BB3C2FE5FFC6F" box="[458,483,993,1012]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
/
<collectorName id="67175338EC555062FE3FB3C3FD33FC6F" box="[494,655,992,1012]" pageId="15" pageNumber="146">Cathedral Rock NP</collectorName>
<geoCoordinate id="AFD65029EC555062FD42B3C3FD51FC6F" box="[659,749,991,1012]" degrees="30.4416" direction="south" orientation="latitude" pageId="15" pageNumber="146" precision="5" value="-30.4416">30.4416°S</geoCoordinate>
<geoCoordinate id="AFD65029EC555062FFA8B3DBFF63FB97" box="[121,223,1015,1036]" degrees="152.2790" direction="east" orientation="longitude" pageId="15" pageNumber="146" precision="5" value="152.279">152.2790°E</geoCoordinate>
<quantity id="0D1A9B0BEC555062FF34B3DAFE9AFB97" box="[229,294,1017,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.347" pageId="15" pageNumber="146" unit="m" value="1347.0">
<elevation id="41CFD1DDEC555062FF34B3DAFE9AFB97" box="[229,294,1017,1036]" metricMagnitude="3" metricUnit="m" metricValue="1.347" pageId="15" pageNumber="146" unit="m" value="1347.0">1347 m</elevation>
</quantity>
site 61 c2k from park entrance, burnt dry scler/open heath, macropod dung baited
<collectingMethod id="13A34EF9EC555062FE5BB433FE03FBB8" box="[394,447,1040,1059]" pageId="15" pageNumber="146">pitfall</collectingMethod>
<date id="BE5C102EEC555062FE16B433FDFDFBB8" box="[455,577,1040,1059]" pageId="15" pageNumber="146" value="2021-02-22" valueMax="2021-02-23" valueMin="2021-02-22">
<collectingDate id="AE18E9C6EC555062FE16B433FDFDFBB8" box="[455,577,1040,1059]" pageId="15" pageNumber="146" value="2021-02-22" valueMax="2021-02-23" valueMin="2021-02-22">2223.ii.2021</collectingDate>
</date>
<collectorName id="67175338EC555062FD9BB433FDCFFBB8" box="[586,627,1040,1059]" pageId="15" pageNumber="146">Reid</collectorName>
&amp; Runagall-McNaull (
<collectionCode id="ACF3AE2BEC555062FF03B40AFEB8FBA7" box="[210,260,1065,1084]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FEC7B40AFD20FBF7" collectingDate="2021-04-02" collectingDateMax="2021-04-03" collectingDateMin="2021-04-02" collectionCode="AMS" collectorName="Donaldson SF &amp; Reid" county="Site" elevation="524" latitude="-28.3418" location="Rabbit Fence" longLatPrecision="6" longitude="152.6693" municipality="Summerland Way" pageId="15" pageNumber="146" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FEC7B40AFE8DFBA7" box="[278,305,1065,1084]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
,
<specimenCount id="DCE4FD67EC555062FEECB40AFEE8FBA7" box="[317,340,1065,1084]" count="1" pageId="15" pageNumber="146" type="female">1♀</specimenCount>
/
<collectorName id="67175338EC555062FEB4B40BFE5CFBA7" box="[357,480,1064,1084]" pageId="15" pageNumber="146">Donaldson SF</collectorName>
<geoCoordinate id="AFD65029EC555062FE36B40AFDFDFBA7" box="[487,577,1063,1084]" degrees="28.3418" direction="south" orientation="latitude" pageId="15" pageNumber="146" precision="5" value="-28.3418">28.3418°S</geoCoordinate>
<geoCoordinate id="AFD65029EC555062FD99B40BFD11FBA7" box="[584,685,1063,1084]" degrees="152.6693" direction="east" orientation="longitude" pageId="15" pageNumber="146" precision="5" value="152.6693">152.6693°E</geoCoordinate>
<quantity id="0D1A9B0BEC555062FD64B40BFD51FBA7" box="[693,749,1064,1084]" metricMagnitude="2" metricUnit="m" metricValue="5.24" pageId="15" pageNumber="146" unit="m" value="524.0">
<elevation id="41CFD1DDEC555062FD64B40BFD51FBA7" box="[693,749,1064,1084]" metricMagnitude="2" metricUnit="m" metricValue="5.24" pageId="15" pageNumber="146" unit="m" value="524.0">524 m</elevation>
</quantity>
<collectingCounty id="233C4E62EC555062FFA8B463FF26FBC8" box="[121,154,1088,1107]" pageId="15" pageNumber="146">Site</collectingCounty>
113
<collectingMunicipality id="2A39AC94EC555062FF13B463FEE5FBCF" box="[194,345,1088,1108]" pageId="15" pageNumber="146">Summerland Way</collectingMunicipality>
nr
<location id="CF3D6035EC555062FEA2B463FE5FFBCF" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC555062FEA2B463FE5FFBCF" box="[371,483,1088,1108]" county="Site" latitude="-28.3418" longLatPrecision="6" longitude="152.6693" municipality="Summerland Way" name="Rabbit Fence" pageId="15" pageNumber="146" stateProvince="Fife">Rabbit Fence</location>
trail, burnt wet scler, macropod dung baited
<collectingMethod id="13A34EF9EC555062FF0EB47BFEADFBF0" box="[223,273,1112,1131]" pageId="15" pageNumber="146">pitfall</collectingMethod>
<date id="BE5C102EEC555062FEC4B47BFEC4FBF0" box="[277,376,1112,1131]" pageId="15" pageNumber="146" value="2021-04-02" valueMax="2021-04-03" valueMin="2021-04-02">
<collectingDate id="AE18E9C6EC555062FEC4B47BFEC4FBF0" box="[277,376,1112,1131]" pageId="15" pageNumber="146" value="2021-04-02" valueMax="2021-04-03" valueMin="2021-04-02">23.iv.2021</collectingDate>
</date>
<collectorName id="67175338EC555062FEADB47BFE18FBF0" box="[380,420,1112,1131]" pageId="15" pageNumber="146">Reid</collectorName>
&amp; Runagall-McNaull (
<collectionCode id="ACF3AE2BEC555062FDB5B47AFD2AFBF7" box="[612,662,1113,1132]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FD75B47AFD03FB07" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="M Gray G Cassis" elevation="845" latitude="-30.816668" location="Survey" longLatPrecision="1219" longitude="152.05" municipality="East Kunderang Trail" pageId="15" pageNumber="146" specimenCount="1" specimenCount-female="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FD75B47AFD07FBF7" box="[676,699,1113,1132]" count="1" pageId="15" pageNumber="146" type="female">1♀</specimenCount>
/
<collectingMunicipality id="2A39AC94EC555062FD19B47AFEBEFB18" pageId="15" pageNumber="146">East Kunderang Trail</collectingMunicipality>
39AR
<locationDeviation id="DBEF2FCCEC555062FEEDB452FDE9FB18" box="[316,597,1136,1156]" location="West Kunderang Trail" pageId="15" pageNumber="146">
<quantity id="0D1A9B0BEC555062FEEDB452FEC9FB18" box="[316,373,1136,1156]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="146" unit="km" value="2.1">2.1 km</quantity>
E of West Kunderang Trail
</locationDeviation>
<geoCoordinate id="AFD65029EC555062FD88B452FD27FB1F" box="[601,667,1137,1156]" degrees="30" direction="south" minutes="49" orientation="latitude" pageId="15" pageNumber="146" precision="925" value="-30.816668">30°49'S</geoCoordinate>
<geoCoordinate id="AFD65029EC555062FD4FB453FD50FB18" box="[670,748,1136,1155]" degrees="152" direction="east" minutes="03" orientation="longitude" pageId="15" pageNumber="146" precision="925" value="152.05">152°03'E</geoCoordinate>
<quantity id="0D1A9B0BEC555062FFA8B4ABFF0FFB07" box="[121,179,1160,1180]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="15" pageNumber="146" unit="m" value="845.0">
<elevation id="41CFD1DDEC555062FFA8B4ABFF0FFB07" box="[121,179,1160,1180]" metricMagnitude="2" metricUnit="m" metricValue="8.45" pageId="15" pageNumber="146" unit="m" value="845.0">845 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC555062FED6B4AAFEFAFB07" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC555062FED6B4AAFEFAFB07" box="[263,326,1161,1180]" latitude="-30.816668" longLatPrecision="1219" longitude="152.05" municipality="East Kunderang Trail" name="Survey" pageId="15" pageNumber="146" stateProvince="Fife">Survey</location>
)
<date id="BE5C102EEC555062FE84B4ABFE72FB00" box="[341,462,1160,1179]" pageId="15" pageNumber="146" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC555062FE84B4ABFE72FB00" box="[341,462,1160,1179]" pageId="15" pageNumber="146" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii9.iv.1993</collectingDate>
</date>
<collectorName id="67175338EC555062FE06B4AAFDCAFB00" box="[471,630,1160,1180]" pageId="15" pageNumber="146">M Gray G Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FD54B4AAFD05FB07" box="[645,697,1161,1180]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FD1AB4AAFDEEFB57" collectingDate="1993-02-04" collectingDateMax="1993-04-09" collectingDateMin="1993-02-04" collectionCode="AMS" collectorName="Richmond Range SF &amp; AR Goanna Ck Rd &amp; Gray &amp; Cassis" elevation="545" latitude="-28.6" location="Survey" longLatPrecision="1231" longitude="152.68333" pageId="15" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FD1AB4AAFD58FB07" box="[715,740,1161,1180]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
/
<collectorName id="67175338EC555062FFA8B483FE97FB2F" box="[121,299,1184,1204]" pageId="15" pageNumber="146">Richmond Range SF</collectorName>
47A 04
<collectorName id="67175338EC555062FEBFB482FDAEFB28" box="[366,530,1184,1204]" pageId="15" pageNumber="146">AR Goanna Ck Rd</collectorName>
,
<geoCoordinate id="AFD65029EC555062FDCAB482FDE1FB28" box="[539,605,1183,1204]" degrees="28" direction="south" minutes="36" orientation="latitude" pageId="15" pageNumber="146" precision="925" value="-28.6">28°36'S</geoCoordinate>
<geoCoordinate id="AFD65029EC555062FDB2B483FD0DFB28" box="[611,689,1184,1203]" degrees="152" direction="east" minutes="41" orientation="longitude" pageId="15" pageNumber="146" precision="925" value="152.68333">152°41'E</geoCoordinate>
<quantity id="0D1A9B0BEC555062FD66B483FD51FB2F" box="[695,749,1184,1204]" metricMagnitude="2" metricUnit="m" metricValue="5.45" pageId="15" pageNumber="146" unit="m" value="545.0">
<elevation id="41CFD1DDEC555062FD66B483FD51FB2F" box="[695,749,1184,1204]" metricMagnitude="2" metricUnit="m" metricValue="5.45" pageId="15" pageNumber="146" unit="m" value="545.0">545 m</elevation>
</quantity>
(NPWS
<location id="CF3D6035EC555062FF6EB49AFF41FB57" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC555062FF6EB49AFF41FB57" box="[191,253,1209,1228]" latitude="-28.6" longLatPrecision="1231" longitude="152.68333" name="Survey" pageId="15" pageNumber="146" stateProvince="Fife">Survey</location>
)
<date id="BE5C102EEC555062FED6B49BFEC0FB50" box="[263,380,1208,1227]" pageId="15" pageNumber="146" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">
<collectingDate id="AE18E9C6EC555062FED6B49BFEC0FB50" box="[263,380,1208,1227]" pageId="15" pageNumber="146" value="1993-02-04" valueMax="1993-04-09" valueMin="1993-02-04">4.ii9.iv.1993</collectingDate>
</date>
M
<collectorName id="67175338EC555062FE46B49AFE7EFB57" box="[407,450,1209,1228]" pageId="15" pageNumber="146">Gray</collectorName>
G
<collectorName id="67175338EC555062FE08B49BFDB3FB50" box="[473,527,1208,1227]" pageId="15" pageNumber="146">Cassis</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FDC8B49AFDF7FB57" box="[537,587,1209,1228]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FD88B49AFD06FB7F" collectingDate="2001-05-10" collectionCode="AMS" collectorName="Francis Greenway HS &amp; Vacy" location="Sydney" pageId="15" pageNumber="146" specimenCount="1" specimenCount-female="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FD88B49AFDCCFB57" box="[601,624,1209,1228]" count="1" pageId="15" pageNumber="146" type="female">1♀</specimenCount>
/
<location id="CF3D6035EC555062FDAFB49BFD02FB50" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC555062FDAFB49BFD02FB50" box="[638,702,1208,1227]" name="Sydney" pageId="15" pageNumber="146" stateProvince="Fife">Sydney</location>
[sic],
<collectorName id="67175338EC555062FFA8B4F3FE88FB7F" box="[121,308,1232,1252]" pageId="15" pageNumber="146">Francis Greenway HS</collectorName>
, leaflitter, cowpasture
<date id="BE5C102EEC555062FE21B4F2FDFAFB7F" box="[496,582,1233,1252]" pageId="15" pageNumber="146" value="2001-05-10">
<collectingDate id="AE18E9C6EC555062FE21B4F2FDFAFB7F" box="[496,582,1233,1252]" pageId="15" pageNumber="146" value="2001-05-10">10.v.2001</collectingDate>
</date>
,
<collectorName id="67175338EC555062FD9CB4F2FDC4FB7F" box="[589,632,1233,1252]" pageId="15" pageNumber="146">Vacy</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FD50B4F2FD0FFB7F" box="[641,691,1233,1252]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FD10B4F2FF0BFAB7" collectingDate="2004-11-23" collectingDateMax="2004-12-09" collectingDateMin="2004-11-23" collectionCode="AMS" collectorName="Denman S &amp; J Gollan" county="Upper Hunter River" latitude="-32.34333" location="Denman" longLatPrecision="20" longitude="150.73834" municipality="Denman Vineyard" pageId="15" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FD10B4F2FD66FB7F" box="[705,730,1233,1252]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
*/
<collectingCounty id="233C4E62EC555062FFA8B4CAFE9BFB60" box="[121,295,1256,1276]" pageId="15" pageNumber="146">Upper Hunter River</collectingCounty>
, mature riparian strip,
<collectingMunicipality id="2A39AC94EC555062FE3DB4CAFD34FB60" box="[492,648,1256,1276]" pageId="15" pageNumber="146">Denman Vineyard</collectingMunicipality>
at
<location id="CF3D6035EC555062FD75B4CAFD50FB67" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC555062FD75B4CAFD50FB67" box="[676,748,1257,1276]" county="Upper Hunter River" latitude="-32.34333" longLatPrecision="20" longitude="150.73834" municipality="Denman Vineyard" name="Denman" pageId="15" pageNumber="146" stateProvince="Fife">Denman</location>
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<geoCoordinate id="AFD65029EC555062FF0EB523FEF0FA8F" box="[223,332,1279,1300]" degrees="150" direction="east" minutes="44" orientation="longitude" pageId="15" pageNumber="146" precision="15" seconds="18" value="150.73834">150°44'18&quot;E</geoCoordinate>
<collectorName id="67175338EC555062FE81B522FE14FA8F" box="[336,424,1281,1300]" pageId="15" pageNumber="146">Denman S</collectorName>
1/2P
<collectingMethod id="13A34EF9EC555062FE05B523FDACFA8F" box="[468,528,1280,1300]" pageId="15" pageNumber="146">pit trap</collectingMethod>
<date id="BE5C102EEC555062FDC5B523FD1DFA88" box="[532,673,1280,1299]" pageId="15" pageNumber="146" value="2004-11-23" valueMax="2004-12-09" valueMin="2004-11-23">
<collectingDate id="AE18E9C6EC555062FDC5B523FD1DFA88" box="[532,673,1280,1299]" pageId="15" pageNumber="146" value="2004-11-23" valueMax="2004-12-09" valueMin="2004-11-23">23.xi9.xii.2004</collectingDate>
</date>
<collectorName id="67175338EC555062FD74B522FD50FA88" box="[677,748,1280,1300]" pageId="15" pageNumber="146">J Gollan</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FFAEB53AFF0DFAB7" box="[127,177,1305,1324]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FF6FB53AFD2BFAB7" box="[190,663,1304,1324]" collectingDate="2004-11-23" collectingDateMax="2004-12-09" collectingDateMin="2004-11-23" collectionCode="AMS" collectorName="Denman S" county="Upper Hunter River" latitude="-32.34333" location="Denman" longLatPrecision="20" longitude="150.73834" municipality="Denman Vineyard" pageId="15" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Fife">
<specimenCount id="DCE4FD67EC555062FF6FB53AFF66FAB7" box="[190,218,1305,1324]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
, ditto except
<collectorName id="67175338EC555062FE97B53AFE20FAB7" box="[326,412,1305,1324]" pageId="15" pageNumber="146">Denman S</collectorName>
1/1
<collectingMethod id="13A34EF9EC555062FE62B53AFDEAFAB7" box="[435,598,1304,1324]" pageId="15" pageNumber="146">YP yellow pan trap</collectingMethod>
(
<collectionCode id="ACF3AE2BEC555062FD8EB53AFD2DFAB7" box="[607,657,1305,1324]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="7A8A3CB3EC555062FD4FB53BFD8EFADF" collectingDate="1974-02-03" collectionCode="AMS" collectorName="Gullan" location="Victoria" pageId="15" pageNumber="146" specimenCount="1" specimenCount-male="1" stateProvince="Victoria">
<emphasis id="F896EAFCEC555062FD4FB53BFD5BFAB0" bold="true" box="[670,743,1304,1323]" pageId="15" pageNumber="146">
<collectingRegion id="0826F80CEC555062FD4FB53BFD5BFAB0" box="[670,743,1304,1323]" country="Australia" name="Victoria" pageId="15" pageNumber="146">Victoria</collectingRegion>
</emphasis>
:
<specimenCount id="DCE4FD67EC555062FFA8B512FF2EFADF" box="[121,146,1329,1348]" count="1" pageId="15" pageNumber="146" type="male">1♂</specimenCount>
*/
<quantity id="0D1A9B0BEC555062FF7BB512FF54FAD8" box="[170,232,1328,1348]" metricMagnitude="3" metricUnit="m" metricValue="6.437376" pageId="15" pageNumber="146" unit="mi" value="4.0">
4
<taxonomicName id="0DE24D6DEC555062FF6BB513FF54FAD8" authority="W Melton" authorityName="W Melton" box="[186,232,1328,1347]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="146" phylum="Arthropoda" rank="species" species="miles">miles</taxonomicName>
</quantity>
W Melton
<date id="BE5C102EEC555062FE98B513FE28FAD8" box="[329,404,1328,1347]" pageId="15" pageNumber="146" value="1974-02-03">
<collectingDate id="AE18E9C6EC555062FE98B513FE28FAD8" box="[329,404,1328,1347]" pageId="15" pageNumber="146" value="1974-02-03">3.ii.1974</collectingDate>
</date>
PJ
<collectorName id="67175338EC555062FE62B513FE51FAD8" box="[435,493,1328,1347]" pageId="15" pageNumber="146">Gullan</collectorName>
(
<collectionCode id="ACF3AE2BEC555062FE28B512FD90FADF" box="[505,556,1329,1348]" pageId="15" pageNumber="146">AMS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="CA5D36EEEC555062FCCBB0B4FB9EFE01" blockId="15.[792,1422,151,411]" pageId="15" pageNumber="146">
<emphasis id="F896EAFCEC555062FCCBB0B4FC7CFF31" bold="true" box="[794,960,151,170]" pageId="15" pageNumber="146">Additional records</emphasis>
(not included in
<bibRefCitation id="AE734B1FEC555062FB99B0B4FB6DFF31" author="Matthews, E." box="[1096,1233,151,170]" pageId="15" pageNumber="146" pagination="3 - 330" refId="ref13173" refString="Matthews, E. 1972. A revision of the scarabaeine dung beetles of Australia. I. tribe Onthophagini. Australian Journal of Zoology. Supplementary Series 9: 3 - 330. https: // doi. org / 10.1071 / AJZS 009" type="book chapter" year="1972">Matthews, 1972</bibRefCitation>
).
<emphasis id="F896EAFCEC555062FB0CB0B4FCD7FF59" bold="true" pageId="15" pageNumber="146">Australian National Territory</emphasis>
: 1/ 6k NE Piccadilly Circus (
<collectionCode id="ACF3AE2BEC555062FB8BB08CFB2EFF59" box="[1114,1170,175,194]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="146">ANIC</collectionCode>
);
<emphasis id="F896EAFCEC555062FB4EB08CFA84FF59" bold="true" box="[1183,1336,175,194]" pageId="15" pageNumber="146">
<collectingRegion id="0826F80CEC555062FB4EB08CFA84FF59" box="[1183,1336,175,194]" country="Australia" name="New South Wales" pageId="15" pageNumber="146">New South Wales</collectingRegion>
</emphasis>
: 1/ Byrrill Ck (
<collectionCode id="ACF3AE2BEC555062FCEEB0E4FCC8FF41" box="[831,884,199,218]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Calosoma HS (
<collectionCode id="ACF3AE2BEC555062FBCEB0E4FBE4FF41" box="[1055,1112,199,218]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="146">ANIC</collectionCode>
); 3/ W end Coolah Tops (
<collectionCode id="ACF3AE2BEC555062FAE6B0E4FAD0FF41" box="[1335,1388,199,218]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ 2.5K W Minyon Falls (
<collectionCode id="ACF3AE2BEC555062FC09B0FCFBB0FF69" box="[984,1036,223,242]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ 4k NE Mt Wog Wog (
<collectionCode id="ACF3AE2BEC555062FB0EB0FCFAABFF69" box="[1247,1303,223,242]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="146">ANIC</collectionCode>
); 1/ Styx R SF (
<collectionCode id="ACF3AE2BEC555062FCF1B0D4FCE4FE91" box="[800,856,247,266]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="146">ANIC</collectionCode>
); 1/ Unumgar SF (
<collectionCode id="ACF3AE2BEC555062FC3FB0D4FB9AFE91" box="[1006,1062,247,266]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="146">ANIC</collectionCode>
);
<emphasis id="F896EAFCEC555062FBE2B0D4FB22FE91" bold="true" box="[1075,1182,247,266]" pageId="15" pageNumber="146">
<collectingRegion id="0826F80CEC555062FBE2B0D4FB22FE91" box="[1075,1182,247,266]" country="Australia" name="Queensland" pageId="15" pageNumber="146">Queensland</collectingRegion>
</emphasis>
: 1/ Belmont Hills (
<collectionCode id="ACF3AE2BEC555062FAEAB0D4FAD3FE91" box="[1339,1391,247,266]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 8/ Buhot Creek (
<collectionCode id="ACF3AE2BEC555062FC41B12CFC79FEB9" box="[912,965,271,290]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Bulimba Creek (
<collectionCode id="ACF3AE2BEC555062FBA4B12CFB16FEB9" box="[1141,1194,271,290]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Burleigh Headland NP (
<collectionCode id="ACF3AE2BEC555062FCF1B104FCE9FEA1" box="[800,853,295,314]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 2/ 14k &amp; 21k SSW Canungra (
<collectionCode id="ACF3AE2BEC555062FBBAB104FB1CFEA1" box="[1131,1184,295,314]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Levers Plateau (
<collectionCode id="ACF3AE2BEC555062FA9EB104FA38FEA1" box="[1359,1412,295,314]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Mt Cotton (
<collectionCode id="ACF3AE2BEC555062FC48B11CFC72FEC9" box="[921,974,319,338]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Mt Gannon (
<collectionCode id="ACF3AE2BEC555062FBB6B11CFB20FEC9" box="[1127,1180,319,338]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 9/ Mt Huntley (
<collectionCode id="ACF3AE2BEC555062FAE6B11CFAD0FEC9" box="[1335,1388,319,338]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Mt
<taxonomicName id="0DE24D6DEC555062FCEAB174FC62FEF1" authority="(QDAF)" baseAuthorityName="QDAF" box="[827,990,343,362]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="146" phylum="Arthropoda" rank="species" species="superbus">Superbus (QDAF)</taxonomicName>
; 8/ Passchendaele SF (
<collectionCode id="ACF3AE2BEC555062FB7DB174FB5DFEF1" box="[1196,1249,343,362]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 2/ Sankeys Scrub (
<collectionCode id="ACF3AE2BEC555062FCF1B14CFCE9FE19" box="[800,853,367,386]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 3/ 6k WNW Stanthorpe (
<collectionCode id="ACF3AE2BEC555062FBEDB14CFBCDFE19" box="[1084,1137,367,386]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 1/ Tallebudgera Valley (
<collectionCode id="ACF3AE2BEC555062FA9EB14CFA38FE19" box="[1359,1412,367,386]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
); 7/ Vickerman Reserve (
<collectionCode id="ACF3AE2BEC555062FC33B1A4FBABFE01" box="[994,1047,391,410]" country="Australia" httpUri="http://grbio.org/cool/awdh-9p9c" name="Queensland Museum, Brisbane" pageId="15" pageNumber="146" type="Museum">QMB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="82F86565EC55507DFCCBB192FB66FE01" lastPageId="16" lastPageNumber="147" pageId="15" pageNumber="146" type="description">
<paragraph id="CA5D36EEEC555062FCCBB192FAF8FDC0" blockId="15.[793,1422,433,603]" pageId="15" pageNumber="146">
<emphasis id="F896EAFCEC555062FCCBB192FC05FE52" bold="true" box="[794,953,433,457]" pageId="15" pageNumber="146">Redescription</emphasis>
. Mature specimens dull blackish-grey, head and pronotum shiner than duller and densely microsculptured elytra and pygidium; teneral (or recently emerged specimens) with reddish-brown elytral apices, sides of apical ventrites and legs; antennae reddish-brown, with orange to dark brown clubs. Length, male
<quantity id="0D1A9B0BEC555062FC23B260FBDEFDC0" box="[1010,1122,579,603]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" metricValueMax="7.0" metricValueMin="4.5" pageId="15" pageNumber="146" unit="mm" value="5.75" valueMax="7.0" valueMin="4.5">4.57 mm</quantity>
; female
<quantity id="0D1A9B0BEC555062FB6DB260FAFCFDC0" box="[1212,1344,579,603]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="15" pageNumber="146" unit="mm" value="5.5" valueMax="6.5" valueMin="4.5">4.56.5 mm</quantity>
.
</paragraph>
<paragraph id="CA5D36EEEC555062FCCBB252FAB2FC73" blockId="15.[793,1422,625,1353]" pageId="15" pageNumber="146">
<emphasis id="F896EAFCEC555062FCCBB252FCEFFD12" bold="true" box="[794,851,625,649]" pageId="15" pageNumber="146">Male</emphasis>
. Head (
<figureCitation id="52D92A6BEC555062FC73B252FC53FD12" box="[930,1007,625,649]" captionStart="Figures 2124" captionStartId="10.[166,243,1231,1253]" captionTargetBox="[166,1466,149,1195]" captionTargetId="figure-11@10.[166,1466,149,1219]" captionTargetPageId="10" captionText="Figures 2124. Onthophagus species, head, male (left), and female (right), fresh specimens. (21, 22) O. pexatus Harold; (23, 24) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228985" httpUri="https://zenodo.org/record/13228985/files/figure.png" pageId="15" pageNumber="146">Fig. 23</figureCitation>
). Surface smooth, shallowly microreticulate, dull to slightly shiny in basal half and shinier in apical half, with close punctures, separated by about 1× diameters at base becoming smaller and sparser towards apex, minute stubble on base of frons and short semi-erect setae on apex of clypeus. Clypeal apical margin uptilted, arcuately excavate, each side of excavation triangularly produced, side margins slightly rounded; clypeal suture entirely effaced at frontal portion, genal suture present (completely effaced on
<specimenCount id="DCE4FD67EC555062FAB7B37FFC3FFC0A" count="1" pageId="15" pageNumber="146" type="generic">one specimen</specimenCount>
) but not raised; frons flat, without elevations, slightly medially depressed; eyes narrow, 57 facet rows in width, separated by 1520 eye widths, canthus narrowly complete; mentum shallowly excavate at apex.
</paragraph>
<paragraph id="CA5D36EEEC55507DFCE9B3CDFE6DFB99" blockId="15.[793,1422,625,1353]" lastBlockId="16.[166,795,151,1349]" lastPageId="16" lastPageNumber="147" pageId="15" pageNumber="146">
Thorax (
<figureCitation id="52D92A6BEC555062FC47B3CDFBBBFB9D" box="[918,1031,1006,1030]" captionStart="Figures 9, 10" captionStartId="6.[447,524,1048,1070]" captionTargetBox="[225,1362,170,1001]" captionTargetId="figure-496@6.[166,1466,148,1025]" captionTargetPageId="6" captionText="Figures 9, 10. Onthophagus squalidus Lea, male dorsal, lateral, and ventral." figureDoi="http://doi.org/10.5281/zenodo.13228975" httpUri="https://zenodo.org/record/13228975/files/figure.png" pageId="15" pageNumber="146">Figs 910</figureCitation>
,
<figureCitation id="52D92A6BEC555062FBC4B3CDFB8EFB9D" box="[1045,1074,1006,1030]" captionStart="Figures 2529" captionStartId="11.[121,198,1565,1587]" captionTargetBox="[121,748,151,1551]" captionTargetId="figure-106@11.[121,748,151,1553]" captionTargetPageId="11" captionText="Figures 2529. Onthophagus species, venter of right metafemur. (25) O. bulga sp. nov.; (26) O. longipes Paulian; (27) O. nammuldi Matthews; (28) O. pexatus Harold; (29) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228987" httpUri="https://zenodo.org/record/13228987/files/figure.png" pageId="15" pageNumber="146">29</figureCitation>
,
<figureCitation id="52D92A6BEC555062FB90B3CDFBE3FB9D" box="[1089,1119,1006,1030]" captionStart="Figures 3034" captionStartId="12.[167,244,762,784]" captionTargetBox="[178,1466,144,748]" captionTargetId="figure-10@12.[166,1466,130,748]" captionTargetPageId="12" captionText="Figures 3034. Onthophagus species, left protibia. (30) O. bulga sp. nov.; (31) O. longipes Paulian; (32) O. nammuldi Matthews; (33) O. pexatus Harold; (34) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228991" httpUri="https://zenodo.org/record/13228991/files/figure.png" pageId="15" pageNumber="146">34</figureCitation>
). Pronotum. Broad in large males (as wide as elytra), narrower and flatter in small males. Moderately convex, slightly tumid at middle of anterior slope in larger specimens; surface smooth, midline not obviously elevated or depressed, entirely strongly and closely punctured, punctures not ocellate, slightly elongate at middle, becoming rounder and larger towards sides, intervals shiny and not microreticulate, less than half diameter of punctures, and covered in dense short curved and thickened setae (lengths less than diameter of punctures); anterior angles 8090°; anterior margined, sides margined and crenulate with curved short setae between crenulations, base with or without thin raised edge at middle; pronotal hypomeron dull and densely microsculptured, strongly punctured on outer half, punctures separated by 12 diameters, with elevated rims, increasing in diameter towards sides, outermost punctures with long semi-erect setae. Elytra. Semi-ovate in dorsal view, with prominent humeri and preapical swelling, longer than pronotum along midline. Surface including striae densely microreticulate and duller than pronotum, except numerous shiny tubercles on intervals and shiny apical half of sutural margins; all intervals flat except for distinct shiny tubercles, apex of 5th interval with irregular shiny area formed by coalesced tubercles, similar but much smaller patches may be present at apices of 3rd and 4th intervals; tubercles in 23 irregular rows in intervals 27, denser on outermost interval; each tubercle with minute short curved scale-like seta, not in distinct rows, outermost interval more densely setose; strial punctures weak, slightly broader and deeper than striae; epipleura with single row of curved setae. Metaventrite medially shiny, anteriorly and laterally (lateral to mesocoxae) dull and microreticulate, with scattered semi-recumbent setae on anterior half, anterior smoothly convex, midline grooved; wing fully developed, folded twice in repose. Legs. Protibiae elongated and narrowed, inner apical angle with compact dense tuft of yellow setae about twice length of apical spur; distal face of apical tooth with loose sparse tuft of yellow setae about as long as apical spur, 4th (smallest) lateral tooth at about midpoint; metafemoral ventral surface smooth and mostly shiny, distinctly microreticulate on apical third, almost evenly punctured with moderately large punctures separated by 13 diameters.
</paragraph>
<caption id="9E9D6666EC555062FF0AB78BFA90F826" ID-DOI="http://doi.org/10.5281/zenodo.13229001" ID-Zenodo-Dep="13229001" box="[219,1324,1960,1982]" httpUri="https://zenodo.org/record/13229001/files/figure.png" pageId="15" pageNumber="146" startId="15.[219,296,1960,1982]" targetBox="[247,1247,1395,1932]" targetPageId="15" targetType="figure">
<paragraph id="CA5D36EEEC555062FF0AB78BFA90F826" blockId="15.[219,1324,1960,1982]" box="[219,1324,1960,1982]" pageId="15" pageNumber="146">
<emphasis id="F896EAFCEC555062FF0AB78BFED6F826" bold="true" box="[219,362,1960,1982]" pageId="15" pageNumber="146">Figures 49, 50</emphasis>
.
<taxonomicName id="0DE24D6DEC555062FEAAB78BFD2BF825" authority="Harold" authorityName="Harold" box="[379,663,1960,1982]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="146" phylum="Arthropoda" rank="species" species="pexatus">
<emphasis id="F896EAFCEC555062FEAAB78BFDF0F825" box="[379,588,1960,1982]" italics="true" pageId="15" pageNumber="146">Onthophagus pexatus</emphasis>
Harold
</taxonomicName>
, apices of parameres. (
<emphasis id="F896EAFCEC555062FCA2B78BFC37F826" box="[883,907,1960,1981]" italics="true" pageId="15" pageNumber="146">49</emphasis>
) Dairy Park, NSW; (
<emphasis id="F896EAFCEC555062FB87B78BFBD2F826" box="[1110,1134,1960,1981]" italics="true" pageId="15" pageNumber="146">50</emphasis>
), Woolomin, NSW.
</paragraph>
</caption>
<paragraph id="CA5D36EEEC4A507DFF14B424FDA6FB76" blockId="16.[166,795,151,1349]" pageId="16" pageNumber="147">
Abdomen (
<figureCitation id="52D92A6BEC4A507DFEE6B424FE3EFBBB" box="[311,386,1031,1056]" captionStart="Figures 3741" captionStartId="13.[819,896,1603,1625]" captionTargetBox="[121,1422,151,1679]" captionTargetId="figure-10@13.[121,1421,151,1679]" captionTargetPageId="13" captionText="Figures 3741. Onthophagus species, pygidium.(37) O. bulga sp. nov.; (38) O. longipes Paulian; (39) O. nammuldi Matthews; (40) O. pexatus Harold; (41) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228995" httpUri="https://zenodo.org/record/13228995/files/figure.png" pageId="16" pageNumber="147">Fig. 41</figureCitation>
). Ventrites dull, densely microreticulate, obscuring puncturation, each with single transverse row of short semi-recumbent pale setae, set on shiny tubercles on ventrites 56; pygidium flat or almost so, densely and evenly microreticulate except shallowly microreticulate and shinier raised margins; punctures distinct or obscured by microsculpture, sparse, separated by 13 diameters; setae short and thick, length 58× width.
</paragraph>
<paragraph id="CA5D36EEEC4A507DFF14B4D1FE7BFADE" blockId="16.[166,795,151,1349]" pageId="16" pageNumber="147">
Aedeagus (
<figureCitation id="52D92A6BEC4A507DFEEAB4D1FE36FA91" box="[315,394,1266,1290]" captionStart="Figures 4248" captionStartId="14.[167,244,1399,1421]" captionTargetBox="[167,1459,164,1379]" captionTargetId="figure-65@14.[166,1465,151,1381]" captionTargetPageId="14" captionText="Figures 4248. Onthophagus species, aedeagus, lateral. (42) O. bulga sp. nov.; (43) O. longipes Paulian; (44) O. nammuldi Matthews; (45) O. squalidus Lea; (46) O. pexatus, Dairy Park, NSW; (47) O. pexatus, Wombeyan Caves, NSW; (48) O. pexatus, Woolomin, NSW." figureDoi="http://doi.org/10.5281/zenodo.13228999" httpUri="https://zenodo.org/record/13228999/files/figure.png" pageId="16" pageNumber="147">Fig. 45</figureCitation>
). In lateral view, parameres “crested”, with an angular apico-dorsal elevation and prominent but short apico-ventral “beak”.
</paragraph>
<paragraph id="CA5D36EEEC4A507DFF77B579FD6CF9A4" blockId="16.[166,794,1370,1599]" pageId="16" pageNumber="147">
<emphasis id="F896EAFCEC4A507DFF77B579FF40FAE9" bold="true" box="[166,252,1370,1394]" pageId="16" pageNumber="147">Female</emphasis>
. Head (
<figureCitation id="52D92A6BEC4A507DFE8CB579FE0AFAE9" box="[349,438,1370,1394]" captionStart="Figures 2124" captionStartId="10.[166,243,1231,1253]" captionTargetBox="[166,1466,149,1195]" captionTargetId="figure-11@10.[166,1466,149,1219]" captionTargetPageId="10" captionText="Figures 2124. Onthophagus species, head, male (left), and female (right), fresh specimens. (21, 22) O. pexatus Harold; (23, 24) O. squalidus Lea." figureDoi="http://doi.org/10.5281/zenodo.13228985" httpUri="https://zenodo.org/record/13228985/files/figure.png" pageId="16" pageNumber="147">Fig. 24</figureCitation>
). Sides more rounded, surface rougher than male, mostly shiny, smooth at base becoming transversely rugose on clypeus, with dense large punctures throughout; median depression of vertex not deeper than male; clypeal suture slightly elevated in frontal portion, effaced before reaching genal sutures, straight, genal sutures often slightly raised; pronotum slightly narrower than elytra; protibiae short and straight, with broad lateral teeth.
</paragraph>
<paragraph id="CA5D36EEEC4A507DFF77B676FD44F911" blockId="16.[166,794,1621,1674]" pageId="16" pageNumber="147">
<emphasis id="F896EAFCEC4A507DFF77B676FE9BF9F6" bold="true" box="[166,295,1621,1645]" pageId="16" pageNumber="147">Minor male</emphasis>
(
<specimenCount id="DCE4FD67EC4A507DFEE2B676FE7AF9F6" box="[307,454,1621,1645]" count="1" pageId="16" pageNumber="147" type="generic">one specimen</specimenCount>
).As major male but genal sutures slightly elevated and protibiae short, without setal tufts.
</paragraph>
<paragraph id="CA5D36EEEC4A507DFF77B683FB97FF71" blockId="16.[166,794,1695,1867]" lastBlockId="16.[839,1467,151,410]" pageId="16" pageNumber="147">
<emphasis id="F896EAFCEC4A507DFF77B683FF59F923" bold="true" box="[166,229,1696,1720]" pageId="16" pageNumber="147">Notes</emphasis>
.
<materialsCitation id="7A8A3CB3EC4A507DFF20B6BCFB9BFF71" county="He" location="The" municipality="Dividing Range" pageId="16" pageNumber="147" specimenCount="172" specimenCount-male="172" stateProvince="Queensland" typeStatus="holotype">
Matthews redescription was based on
<specimenCount id="DCE4FD67EC4A507DFD55B683FD53F92C" box="[644,751,1695,1720]" count="172" pageId="16" pageNumber="147" type="male">172 males</specimenCount>
and females, including 141 from a single locality (Canberra), and the female
<typeStatus id="1559884CEC4A507DFECAB6F9FEC4F969" box="[283,376,1754,1778]" pageId="16" pageNumber="147" type="holotype">holotype</typeStatus>
from south
<collectingRegion id="0826F80CEC4A507DFE29B6F9FDC5F969" box="[504,633,1754,1778]" country="Australia" name="Queensland" pageId="16" pageNumber="147">Queensland</collectingRegion>
.
<collectingCounty id="233C4E62EC4A507DFD53B6F8FD1DF969" box="[642,673,1755,1778]" pageId="16" pageNumber="147">He</collectingCounty>
recorded it along the coast and
<collectingMunicipality id="2A39AC94EC4A507DFEADB6D4FD9AF88B" box="[380,550,1783,1808]" pageId="16" pageNumber="147">Dividing Range</collectingMunicipality>
from
<collectingRegion id="0826F80CEC4A507DFDB6B6D4FD01F894" box="[615,701,1783,1807]" country="Australia" name="Victoria" pageId="16" pageNumber="147">Victoria</collectingRegion>
to south
<collectingRegion id="0826F80CEC4A507DFF77B736FE95F8B6" box="[166,297,1813,1837]" country="Australia" name="Queensland" pageId="16" pageNumber="147">Queensland</collectingRegion>
.
<location id="CF3D6035EC4A507DFEE7B736FEDCF8B6" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC4A507DFEE7B736FEDCF8B6" box="[310,352,1813,1837]" county="He" municipality="Dividing Range" name="The" pageId="16" pageNumber="147" stateProvince="Queensland">The</location>
material examined here fits within this range and conforms to
<location id="CF3D6035EC4A507DFE45B711FDB8F8D1" LSID="urn:lsid:plazi:treatment:424B87F8EC55507DFF69B26FFB66FE01:CF3D6035EC4A507DFE45B711FDB8F8D1" box="[404,516,1842,1866]" county="He" municipality="Dividing Range" name="Matthews'" pageId="16" pageNumber="147" stateProvince="Queensland">Matthews</location>
description, except that he noted a slightly different size range (
<quantity id="0D1A9B0BEC4A507DFB15B0B4FAA7FF34" box="[1220,1307,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="16" pageNumber="147" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">58 mm</quantity>
long), the eyes are slightly more variable in size and the striae are distinctly but feebly punctured
</materialsCitation>
.
</paragraph>
<paragraph id="CA5D36EEEC4A507DFCB4B0CCFB66FE01" blockId="16.[839,1467,151,410]" pageId="16" pageNumber="147">
<taxonomicName id="0DE24D6DEC4A507DFCB4B0CCFBDDFE9C" box="[869,1121,239,263]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="147" phylum="Arthropoda" rank="species" species="squalidus">
<emphasis id="F896EAFCEC4A507DFCB4B0CCFBDDFE9C" box="[869,1121,239,263]" italics="true" pageId="16" pageNumber="147">Onthophagus squalidus</emphasis>
</taxonomicName>
appears to be tolerant of a wider range of habitats (Reid
<emphasis id="F896EAFCEC4A507DFB8DB12DFB22FEBE" box="[1116,1182,269,293]" italics="true" pageId="16" pageNumber="147">et al.,</emphasis>
2022) than indicated by Matthews, who regarded it as a woodland species. The sites recorded here vary from closed forest to open grassland. The species appears to be rare where it occurs in
<collectingRegion id="0826F80CEC4A507DFA94B146FC36FE01" country="Australia" name="New South Wales" pageId="16" pageNumber="147">New South Wales</collectingRegion>
, most records being singletons.
</paragraph>
</subSubSection>
</treatment>
</document>