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<document id="7D67F8DAE6600BC915ECC0F0BAF32813" ID-CLB-Dataset="30010" ID-DOI="http://dx.doi.org/10.3897/zookeys.761.24214" ID-GBIF-Dataset="6f3db375-08bb-453e-b1ff-424f0354452a" ID-PMC="PMC5988806" ID-Pensoft-Pub="1313-2970-761-1" ID-PubMed="29875597" ID-ZooBank="9165835900AE4319ACBCE9C544599C5B" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-761-1" ModsDocOrigin="ZooKeys 761" ModsDocTitle="A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1527611678831" checkinUser="pensoft" docAuthor="Srisonchai, Ruttapon, Enghoff, Henrik, Likhitrakarn, Natdanai &amp; Panha, Somsak" docDate="2018" docId="41DC3F5916B6E098E9A8F2DEF6B46C5F" docLanguage="en" docName="ZooKeys 761: 1-177" docOrigin="ZooKeys 761" docSource="http://dx.doi.org/10.3897/zookeys.761.24214" docTitle="Desmoxytes Chamberlin 1923" docType="treatment" docVersion="6" lastPageNumber="1" masterDocId="FF8B4167FFBFFFA3F22FD43BFFCED866" masterDocTitle="A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1701377357523" updateUser="ExternalLinkService">
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<mods:title id="E7A8EBE74DA9E3C3FCF02BE2BA81C9F1">A revision of dragon millipedes I: genus Desmoxytes Chamberlin, 1923, with the description of eight new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:namePart id="5F4B62C473D977A34DE671D271EC3C70">Srisonchai, Ruttapon</mods:namePart>
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<mods:namePart id="C7E573E083CED28FDB1F303560DF0737">Likhitrakarn, Natdanai</mods:namePart>
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<mods:namePart id="D658A9A59718F56CC2C159E7A7D249F3">Panha, Somsak</mods:namePart>
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<treatment id="41DC3F5916B6E098E9A8F2DEF6B46C5F" ID-GBIF-Taxon="144172418" LSID="urn:lsid:plazi:treatment:41DC3F5916B6E098E9A8F2DEF6B46C5F" httpUri="http://treatment.plazi.org/id/41DC3F5916B6E098E9A8F2DEF6B46C5F" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection id="B2D0912159D0038A491BCC9FB827708E" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="9ABD9804B64593B6B0D74D4F40FEA423" pageId="0" pageNumber="1">
Genus
<taxonomicName id="55FD1679EDFC34037A5E7AF321DC721A" ID-CoL="7P8ZX" authority="Chamberlin, 1923" authorityName="Chamberlin" authorityYear="1923" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes Chamberlin, 1923</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="A966B0E7EEF76C579EAB6E83D25930CD" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="567F3DDB47165D801D3D23D408350EC8" pageId="0" pageNumber="1">
<taxonomicName id="176EA8EC0F0BD27B9A1402BE7046F88D" class="Diplopoda" family="Paradoxosomatidae" genus="Prionopeltis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prionopeltis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Prionopeltis</taxonomicName>
Pocock, 1895: 828 (preoccupied name).
<bibRefCitation id="2AC2C9AE326920607D59932545DE0357" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="651 - 670" title="The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida)." url="https://biodiversitylibrary.org/page/41420961" volume="87" year="1980 a">Jeekel 1980a</bibRefCitation>
: 652 (synonymised with
<taxonomicName id="E6F8F7457BD2EA970C4CC771C4585281" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
).
</paragraph>
<paragraph id="BF0BE9A66FF6902A8168B570A2B3C77E" pageId="0" pageNumber="1">
<taxonomicName id="D2E20A8F665326DE6B0845603176D371" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
Chamberlin, 1923: 165.
</paragraph>
<paragraph id="08C649F8C8F70F388AD0C38CF97E345B" pageId="0" pageNumber="1">
<taxonomicName id="FBE1A752572096C52ED41C1B80F99857" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hylomus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Hylomus</taxonomicName>
Cook &amp; Loomis, 1924: 105.
<bibRefCitation id="596DF54ABE56929530AD01598F97AF85" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff 1994</bibRefCitation>
: 46 (synonymised with
<taxonomicName id="CF833A55F03E89AA47CB346D58BE9061" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
).
</paragraph>
<paragraph id="68FFF42ABCE3007396FAEFDEEB2AE084" pageId="0" pageNumber="1">
<taxonomicName id="4CC20C285241361948617C09F3D548B2" class="Diplopoda" family="Paradoxosomatidae" genus="Pratinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pratinus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pratinus</taxonomicName>
Attems, 1937: 113 (replacement name for
<taxonomicName id="77F1B9B4DD418A08C9B54D1169C6EA30" class="Diplopoda" family="Paradoxosomatidae" genus="Prionopeltis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Prionopeltis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Prionopeltis</taxonomicName>
).
<bibRefCitation id="DB825432A7370A930259EFE0A527893E" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="651 - 670" title="The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida)." url="https://biodiversitylibrary.org/page/41420961" volume="87" year="1980 a">Jeekel 1980a</bibRefCitation>
: 652 (synonymised with
<taxonomicName id="03A6D04D5AACA3C7F711B43B22292AF5" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
).
</paragraph>
<paragraph id="24EBE9C505CC0D27C3030146F4D252D9" pageId="0" pageNumber="1">
<taxonomicName id="C866798E2B6D7F02A3A5F20ADC58C6FE" class="Diplopoda" family="Paradoxosomatidae" genus="Ceylonesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ceylonesmus" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Ceylonesmus</taxonomicName>
Chamberlin, 1941: 33.
<bibRefCitation id="047EAC57042CDE87F5EBA8A2E1AB3F37" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="651 - 670" title="The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida)." url="https://biodiversitylibrary.org/page/41420961" volume="87" year="1980 a">Jeekel 1980a</bibRefCitation>
: 652 (synonymised with
<taxonomicName id="0BE4344F662227F533A15F3D2AF57F3B" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
).
</paragraph>
<paragraph id="5AE0B33A14F1B047DC48C08AAAFF2993" pageId="0" pageNumber="1">
<taxonomicName id="6B3A5BD35EC2AB296407D4FCF0DE9857" class="Diplopoda" family="Paradoxosomatidae" genus="Pteroxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pteroxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Pteroxytes</taxonomicName>
Jeekel, 1980a: 655.
<bibRefCitation id="4E14295E8B45D78F3ABF227366758BA9" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff 1994</bibRefCitation>
: 46 (synonymised with
<taxonomicName id="A777531EA667E5478EEA0DCFDE0AD486" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="595F87BEBB888582F3F6E1C1E0BEDB1E" pageId="0" pageNumber="1" type="type species">
<paragraph id="2BA5EF6758DFE045D5AD863BAEACC341" pageId="0" pageNumber="1">Type species</paragraph>
<paragraph id="332D0E10F4F01992679DF169A43EB144" pageId="0" pageNumber="1">
Type species.
<taxonomicName id="28A970EE9BB48F09DB2DA87DB299FED8" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes coniger" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="coniger">Desmoxytes coniger</taxonomicName>
Chamberlin, 1923 (MCZ, USA). This species was later synonymised with
<taxonomicName id="BA64A0CC0E85E6FC99DF08AD150DDF29" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes planata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="planata">Desmoxytes planata</taxonomicName>
by
<bibRefCitation id="82F7A35C446BADBAC35FD54F3FF253ED" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="651 - 670" title="The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida)." url="https://biodiversitylibrary.org/page/41420961" volume="87" year="1980 a">Jeekel (1980a)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="616EB74551B2A9D4F8A4F6ECC082FB53" pageId="0" pageNumber="1" type="included species">
<paragraph id="600A1CBBA0C52EB6A492D48B1FF12FE2" pageId="0" pageNumber="1">Included species</paragraph>
<paragraph id="8CAF45BBDD3AF91A717DD5FD5DB5F382" pageId="0" pageNumber="1">(18).</paragraph>
<paragraph id="BFF6420F54A4BDDB82E850042AC70426" pageId="0" pageNumber="1">
-
<taxonomicName id="ECCB7EE6E9D7F727F695FFB973841EE8" lsidName="D. aurata" pageId="0" pageNumber="1" rank="species" species="aurata">D. aurata</taxonomicName>
sp. n.
</paragraph>
<paragraph id="0C7337842BDCBAB5466CB783662B5213" pageId="0" pageNumber="1">
-
<taxonomicName id="396978D38B6552FCE653E1283C101D50" lsidName="D. breviverpa" pageId="0" pageNumber="1" rank="species" species="breviverpa">D. breviverpa</taxonomicName>
Srisonchai, Enghoff &amp; Panha, 2016
</paragraph>
<paragraph id="291BCC23E35BF1A85D2DC379663A1C27" pageId="0" pageNumber="1">
-
<taxonomicName id="F4E38C28701E26997AB4EE0BAD820277" lsidName="D. cervina" pageId="0" pageNumber="1" rank="species" species="cervina">D. cervina</taxonomicName>
(Pocock, 1895)
</paragraph>
<paragraph id="AB7545FC30EDF800D0D49E03F19B820E" pageId="0" pageNumber="1">
-
<taxonomicName id="95C458399358F6BE8705683864CFD10F" lsidName="D. corythosaurus" pageId="0" pageNumber="1" rank="species" species="corythosaurus">D. corythosaurus</taxonomicName>
sp. n.
</paragraph>
<paragraph id="A6296B08531F1135D5B16DEF55C1D2E5" pageId="0" pageNumber="1">
-
<taxonomicName id="93C6C6D32E3BD9E9FF4A1AC1CCC380CB" lsidName="D. delfae" pageId="0" pageNumber="1" rank="species" species="delfae">D. delfae</taxonomicName>
(Jeekel, 1964)
</paragraph>
<paragraph id="913D43683D80D52B7AF920FBDB28BB7F" pageId="0" pageNumber="1">
-
<taxonomicName id="E00EDE7A74012E92387A343B22483CA7" lsidName="D. des" pageId="0" pageNumber="1" rank="species" species="des">D. des</taxonomicName>
Srisonchai, Enghoff &amp; Panha, 2016
</paragraph>
<paragraph id="72531BA304E116729B0976167996B4A1" pageId="0" pageNumber="1">
-
<taxonomicName id="9D73D4AB8E36916F68C660FBB8A2E11E" lsidName="D. euros" pageId="0" pageNumber="1" rank="species" species="euros">D. euros</taxonomicName>
sp. n.
</paragraph>
<paragraph id="44EFC6EC8B9808880BA2479FCC5CB147" pageId="0" pageNumber="1">
-
<taxonomicName id="18B7601BD8F913B267D425557DACBA74" lsidName="D. flabella" pageId="0" pageNumber="1" rank="species" species="flabella">D. flabella</taxonomicName>
sp. n.
</paragraph>
<paragraph id="A2AF215B6DB9A6150E2C15A446960CE1" pageId="0" pageNumber="1">
-
<taxonomicName id="00CC4AF85F39B149AEFF77DDBE2082EE" lsidName="D. golovatchi" pageId="0" pageNumber="1" rank="species" species="golovatchi">D. golovatchi</taxonomicName>
sp. n.
</paragraph>
<paragraph id="AB93D2332D5D572FFFF7D024A767B990" pageId="0" pageNumber="1">
-
<taxonomicName id="DF12452796FB071E999A9BAA29CEA117" lsidName="D. octoconigera" pageId="0" pageNumber="1" rank="species" species="octoconigera">D. octoconigera</taxonomicName>
sp. n.
</paragraph>
<paragraph id="2E374E1C9EE0D7984A78E14FAE40AE6C" pageId="0" pageNumber="1">
-
<taxonomicName id="CF98F6875D944959E52C3A9117E2A5FD" lsidName="D. perakensis" pageId="0" pageNumber="1" rank="species" species="perakensis">D. perakensis</taxonomicName>
sp. n.
</paragraph>
<paragraph id="1015196A951D5CFC938977C40F1A95C3" pageId="0" pageNumber="1">
-
<taxonomicName id="795A512D33EA799B71B6BF670B230B73" lsidName="D. pinnasquali" pageId="0" pageNumber="1" rank="species" species="pinnasquali">D. pinnasquali</taxonomicName>
Srisonchai, Enghoff &amp; Panha, 2016
</paragraph>
<paragraph id="F5C84A56CDB4FD362DAD10D4A6B9BB52" pageId="0" pageNumber="1">
-
<taxonomicName id="468323A8BAB7DD4441ECF04276323250" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
(Pocock, 1895) (=
<taxonomicName id="2B474B441D46548CC84A42BFEEEC3A90" lsidName="D. coniger" pageId="0" pageNumber="1" rank="species" species="coniger">D. coniger</taxonomicName>
Chamberlin, 1923, type species)
</paragraph>
<paragraph id="ABA6C70FD23B2C5FA8543E3570DC5E8B" pageId="0" pageNumber="1">
-
<taxonomicName id="CC26DCFDDBB86BF15FEEC6F4B2CB6CE4" lsidName="D. purpurosea" pageId="0" pageNumber="1" rank="species" species="purpurosea">D. purpurosea</taxonomicName>
Enghoff, Sutcharit &amp; Panha, 2007
</paragraph>
<paragraph id="2CFBAE10501EC052645CA131D48C9398" pageId="0" pageNumber="1">
-
<taxonomicName id="2BDB58F6906EDF436E8FC45502F897EA" lsidName="D. takensis" pageId="0" pageNumber="1" rank="species" species="takensis">D. takensis</taxonomicName>
Srisonchai, Enghoff &amp; Panha, 2016
</paragraph>
<paragraph id="CBF42D082EF6289F5C025C96F4ED8C7B" pageId="0" pageNumber="1">
-
<taxonomicName id="7DF63389DE91438E5BC5597369E0DBB5" lsidName="D. taurina" pageId="0" pageNumber="1" rank="species" species="taurina">D. taurina</taxonomicName>
(Pocock, 1895)
</paragraph>
<paragraph id="0D24F87AF44766CC4544B9F2EC8FD7F6" pageId="0" pageNumber="1">
-
<taxonomicName id="BE69BE3BB14CCD482D1366CE5FFB610A" lsidName="D. terae" pageId="0" pageNumber="1" rank="species" species="terae">D. terae</taxonomicName>
(Jeekel, 1964)
</paragraph>
<paragraph id="25ADBDC1A8C7427C323CA674353A5A2E" pageId="0" pageNumber="1">
-
<taxonomicName id="D9C329F6A80B3696761E9918000BC7A1" lsidName="D. waepyanensis" pageId="0" pageNumber="1" rank="species" species="waepyanensis">D. waepyanensis</taxonomicName>
sp. n.
</paragraph>
<paragraph id="D37C5553F49F5FAE3156ED2D0C32EAF0" pageId="0" pageNumber="1">
<taxonomicName id="156FC950C2BEE58788F4B373FB5AD469" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
s.l. was reviewed by
<bibRefCitation id="50FBC41827449CDD8A90AF7919D6D328" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff (1994)</bibRefCitation>
, and the diagnosis of the genus was treated in detail again by
<bibRefCitation id="759134FA7BA994FB2D97B14C167333EA" author="Golovatch, SI" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 17" title="Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.185.3082" volume="185" year="2012">Golovatch et al. (2012)</bibRefCitation>
. Here we propose a restricted diagnosis for
<taxonomicName id="B7FA92C44E145256E64919B0CDFE5FC4" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
s.s. based on morphological characters (gonopod, tegument, paraterga, metaterga, sterna, femora, epiproct, hypoproct) which have been extracted from previous taxonomic works (
<bibRefCitation id="E184379EDA057916C26C06DB7217D3EB" author="Pocock, RI" journalOrPublisher="Annali del Museo civico di Storia Naturale Giacomo Doria" pageId="0" pageNumber="1" pagination="787 - 834" title="The Myriopoda of Burma, Pt. IV. Report upon the Polydesmoidea collected by Sig. L. Fea, Mr. E. W. Oates and others, by R. I. Pocock of the British Museum of Natural History." url="https://archive.org/details/cbarchive_104716_themyriapodaofburma1870" volume="14" year="1895">Pocock 1895</bibRefCitation>
,
<bibRefCitation id="6437221C79B381DD0ED9286144F6CEEE" author="Chamberlin, RV" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="1" pagination="165 - 168" title="Two diplopod immigrants taken at Honolulu." url="https://biodiversitylibrary.org/page/34510694" volume="36" year="1923">Chamberlin 1923</bibRefCitation>
,
<bibRefCitation id="F14CE2F70FC2ADADEDB69A92331983C8" author="Chamberlin, RV" journalOrPublisher="Proceedings of the Entomological Society of Washington" pageId="0" pageNumber="1" pagination="32 - 35" title="New polydesmoid diplopods intercepted at quarantine." url="https://biodiversitylibrary.org/page/16130128" volume="43" year="1941">1941</bibRefCitation>
,
<bibRefCitation id="13AB087C01F7525057D03A104EFF9ED2" author="Jeekel, CAW" journalOrPublisher="Beaufortia" pageId="0" pageNumber="1" pagination="61 - 73" title="Two new species of Pratinus Attems, with taxonomic notes on the genus and a redescription of its type-species (Diplopoda, Polydesmida)." url="http://www.repository.naturalis.nl/document/548475" volume="11" year="1964">Jeekel 1964</bibRefCitation>
,
<bibRefCitation id="FE60ACB9A6AC36C79C58A1115BD5E2FA" author="Jeekel, CAW" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="1" pagination="651 - 670" title="The generic allocation of some little-known Paradoxosomatidae from South-East Asia (Diplopoda, Polydesmida)." url="https://biodiversitylibrary.org/page/41420961" volume="87" year="1980 a">1980a</bibRefCitation>
,
<bibRefCitation id="894823F69EBA64E51A0B949CF8ECA435" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff 1994</bibRefCitation>
,
<bibRefCitation id="02ED7EF7B2BDC6E0CDFB7A0BD9140A80" author="Enghoff, H" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="31 - 36" title="The shocking pink dragon millipede, Desmoxytespurpurosea, a colourful new species from Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://dx.doi.org/10.11646/zootaxa.1563.1.3" volume="1563" year="2007">Enghoff et al. 2007</bibRefCitation>
,
<bibRefCitation id="688159FAD4C699F168FE6DD1C6DD4654" author="Srisonchai, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="93 - 113" title="Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://dx.doi.org/10.11646/zootaxa.4170.1.4" volume="4710" year="2016">Srisonchai et al. 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="D4FFB1812768FEF3DCBFD47D76B91FB1" pageId="0" pageNumber="1" type="the name “dragon millipede”">
<paragraph id="386CB5DD2B31320491757F0F6801889B" pageId="0" pageNumber="1">The name &quot;dragon millipede&quot;.</paragraph>
<paragraph id="A844D677737E322FB53EBB7E55E41C2A" pageId="0" pageNumber="1">
For
<taxonomicName id="DA007DEB83BDE218561AEA99F933B314" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
sensu
<bibRefCitation id="475A397D4C3BE8C594F431E9409C1BDF" author="Golovatch, SI" journalOrPublisher="Steenstrupia" pageId="0" pageNumber="1" pagination="45 - 71" title="Review of the dragon millipedes, genus Desmoxytes Chamberlin, 1923 (Diplopoda, Polydesmida, Paradoxosomatidae)." volume="20" year="1994">Golovatch and Enghoff (1994)</bibRefCitation>
we will retain the name &quot;dragon millipede&quot;, originally coined by
<bibRefCitation id="BED486160C3EAF08414CABFE5B482D79" author="Cook, OF" journalOrPublisher="Journal of the Washington Academy of Sciences" pageId="0" pageNumber="1" pagination="103 - 108" title="A new family of spined millipeds from central China." url="https://biodiversitylibrary.org/page/39702295" volume="14" year="1924">Cook and Loomis (1924)</bibRefCitation>
for
<taxonomicName id="1BFE280308EFA475D5E6BD01625B919C" class="Diplopoda" family="Paradoxosomatidae" genus="Hylomus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hylomus draco" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="draco">Hylomus draco</taxonomicName>
.
<bibRefCitation id="2E7F1999E139CE432FD7AC8E2B092DD6" author="Golovatch, SI" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 17" title="Three new cavernicolous species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from southern China, with notes on a formal congener from the Philippines (Diplopoda, Polydesmida, Paradoxosomatidae)." url="https://doi.org/10.3897/zookeys.185.3082" volume="185" year="2012">Golovatch et al. (2012)</bibRefCitation>
outlined the history of the name &quot;dragon millipede&quot; and further argued that
<taxonomicName id="40BE15A98D835FB98517E22325CB1EBA" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes philippina" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="philippina">Desmoxytes philippina</taxonomicName>
Nguyen &amp; Sierwald, 2010 (Philippine Isl.), and
<taxonomicName id="11C70E2E6CEFD8674EB83C614B3BF59A" class="Diplopoda" genus="Desmoxytoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytoides hasenpuschorum" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="hasenpuschorum">Desmoxytoides hasenpuschorum</taxonomicName>
Mesibov, 2006 (Australia) have been wrongly assigned to
<taxonomicName id="A8181DE95DE143B987973AF49FBA4B00" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
and &quot;dragon millipedes&quot;, respectively.
</paragraph>
</subSubSection>
<subSubSection id="E53C8F644E4CDA16A3D3A8FA66613EC8" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="902868C1611321B20741ECB5E4D388D9" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="092CD6DBAF484F72F285C29CD1B913D1" pageId="0" pageNumber="1">
<taxonomicName id="4BC593CF4B508B5B07209B0FE4311F0F" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
s.s. differs from other genera of
<taxonomicName id="8C89A4F60521BE22A22B9A8BE6143A0B" lsidName="" pageId="0" pageNumber="1" rank="tribe" tribe="Orthomorphini">Orthomorphini</taxonomicName>
by the combination of the following characters:
</paragraph>
<paragraph id="BED56A7F5084F3D37F58DB8294D01E7D" pageId="0" pageNumber="1">1. Gonopod suberect: solenophore strongly condensed; lamina lateralis developed, lobe-like, without process or spine; lamina medialis distinctly demarcated from lamina lateralis, bearing process and lobes.</paragraph>
<paragraph id="53B88E4E6E3C77D8D8BA176D462D3C84" pageId="0" pageNumber="1">2. Metaterga with 2 transverse rows of setiferous setae/ tubercles/ cones/ spines.</paragraph>
<paragraph id="76843B6DB84BA377432C9C10B2AE07A5" pageId="0" pageNumber="1">3. Paraterga wing-shaped, well-elevated.</paragraph>
<paragraph id="9E73D7D6F10D602A6EDAF31DBD49FD08" pageId="0" pageNumber="1">4. Sternal cone between male coxae 4 present; subtrapeziform/ subquadrate/ subsemicircular/ incompletely bilobed.</paragraph>
<paragraph id="EEB1F20C0A03530485C862E92E13BD4B" pageId="0" pageNumber="1">
5. Male femora 5 and 6 modified; swollen/humped (exception:
<taxonomicName id="42D338A3C2057FCA25BFBDB26306017E" lsidName="D. terae" pageId="0" pageNumber="1" rank="species" species="terae">D. terae</taxonomicName>
without modification).
</paragraph>
</subSubSection>
<subSubSection id="0EB3406C7958C5910CC9A127DB42965A" pageId="0" pageNumber="1" type="description">
<paragraph id="0F53E11F969B3F69B9A29DCDD210D57C" pageId="0" pageNumber="1">
General description of the genus
<taxonomicName id="6CC49A5C2696CC6B276DD049997F9B61" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
s.s.
</paragraph>
<paragraph id="B650AA7E7FCEC548F858A1D3856D0301" pageId="0" pageNumber="1">
The description applies to adult males and females, except for the gonopods section and when
<normalizedToken id="9829A525BC5CCBF1363515B7C2AFF97E" originalValue="“male”">&quot;male&quot;</normalizedToken>
is specified (Figs 5, 6, 7, 8). The description hereunder is mainly based on illustrations of
<taxonomicName id="87808835F5A3474981D5275ABF7BB62D" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
.
</paragraph>
<caption id="C95D864A0AB4CF66CE5CD67047B14948" pageId="0" pageNumber="1">
<paragraph id="9A10CA10F9D6DEFA53C08C72208D2380" pageId="0" pageNumber="1">
Figure 4. Common habitats for
<taxonomicName id="E75973473B165DC6E4B544C93E8E9975" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
Chamberlin, 1923. A, B limestone mountain C rock wall and plants D rock wall and small cave.
</paragraph>
</caption>
<caption id="A04C2D407F9B4AD0A6DE20B523710A2C" pageId="0" pageNumber="1">
<paragraph id="0E68F0D5CB40FB486D02DBF3F7E89DE9" pageId="0" pageNumber="1">
Figure 5. General body characters of
<taxonomicName id="52984CCA0802E429E8984A223894B459" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
-
<taxonomicName id="706F3E786896AE5C5C501CC7F49EEFE0" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
(Pocock, 1895), male specimen from Wat Puang Malai. A whole body B anterior body part C head region D body ring E body rings 5-7, showing sternal lobe between coxae 4 and gonopods on ring 7 F male femora 5 or 6.
</paragraph>
</caption>
<caption id="439E869A629090838145C637C54B66C9" pageId="0" pageNumber="1">
<paragraph id="F43520F36079E708FBC9AB7CA5F19C09" pageId="0" pageNumber="1">
Figure 6. General body characters of
<taxonomicName id="86E5A90562017C9E369322DBB04873E7" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
(
<taxonomicName id="B608AF1FA289CCFB125B6507421D4F42" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
(Pocock, 1895), male specimen from Wat Puang Malai) - SEM images. A, B anterior body part C collum D, E body rings 9-10 F last ring and telson G posteriormost rings H body ring 10 I mouth parts J leg 13 (right) K sternal lobe between male coxae 4 L gonopods on ring 7 M tip of tarsus and claw of leg 13 N tip of epiproct.
</paragraph>
</caption>
<caption id="EABEC89445EC12665721194A2DB5A499" pageId="0" pageNumber="1">
<paragraph id="9C0F870AD0245BD45C397D24827306B8" pageId="0" pageNumber="1">
Figure 7. Diagrammatic drawings of right gonopod of
<taxonomicName id="4B7AAABA859900238BED59189768F918" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
-
<taxonomicName id="37C8B8C832FE27AD10F7D54107A0369E" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
(Pocock, 1895), specimen from Wat Puang Malai. A mesal view B dorsal view C ventral view.
</paragraph>
</caption>
<caption id="C0A54CABAE0DD800A462F59E35551DE5" pageId="0" pageNumber="1">
<paragraph id="23E37701A53863EF2FA0052F2F912ECE" pageId="0" pageNumber="1">
Figure 8. SEM images of right gonopod of
<taxonomicName id="13390AD1CB464F11E29122BB07A255FB" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes planata" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="planata">Desmoxytes planata</taxonomicName>
(Pocock, 1895), specimen from Wat Puang Malai. A lateral view B mesal view C submesal view D ventral view.
</paragraph>
</caption>
<paragraph id="F69F8DAB02720D49853681DBAD68A0B1" pageId="0" pageNumber="1">SIZE: Body length 16-35 mm (male) 20-38 mm (female), width ca. 1.7-2.5 mm (male) 2.0-3.6 mm (female); size varies between species, usually female wider and longer than male.</paragraph>
<paragraph id="B3DE65A7DA4FCFF157636BFFE58B7AF5" pageId="0" pageNumber="1">
COLOUR: Most species in life with aposematic colouration: purplish pink, red, orange, brown, black, brownish black (piceous), brownish red (testaceous). Colour in alcohol: all specimens partly faded after one
<normalizedToken id="A9E0266FF2433946448B59209DD4CE40" originalValue="year’s">year's</normalizedToken>
preservation in alcohol; specimens kept in darkness faded more slowly.
</paragraph>
<paragraph id="408B2A4A9F4E09F332BB1D82991220B4" pageId="0" pageNumber="1">HEAD (Figs 5C, 6A, B, I): Sparsely setose; vertex bare; labrum and genae sparsely setose; epicranial suture conspicuous as a brown or black stripe.</paragraph>
<paragraph id="9DEC02B643C918B9134F010AC826CF83" pageId="0" pageNumber="1">
ANTENNAE (Fig. 5
<normalizedToken id="CE8A942108791BDB3397DC130CBFEB27" originalValue="A–C">A-C</normalizedToken>
): Often long and slender, rarely short, covered by delicate setation, usually reaching backwards to body ring 3-8 (male) and 3-6 (female) when stretched dorsally. Antennomere lengths 3 = 4 = 5&gt; 2&gt; 6&gt; 1&gt; 7&gt; 8.
</paragraph>
<paragraph id="520A937E8CA60A6737DCD91E6F30ADA4" pageId="0" pageNumber="1">COLLUM (Fig. 6A, C): With 1-3 transverse rows of setae/tubercles, the exact number in each row varying between species (sometimes lateral setae/tubercles of anterior row located nearly at the base of paraterga, on anterior margin). Paraterga wing-shaped, usually elevated at ca. 0°-30°.</paragraph>
<paragraph id="D7A97259D27D48AA762D5AA4C19505C3" pageId="0" pageNumber="1">
TEGUMENT (Fig. 6
<normalizedToken id="385D56F79C80688D34F7F4CC8C7D7ABC" originalValue="A–E">A-E</normalizedToken>
, G): Often shining, sometimes dull; collum and metaterga often microgranulate; surface below paraterga coarsely or finely microgranulate; prozona finely shagreened. Suture between prozona and metazona usually shallow, quite narrow.
</paragraph>
<paragraph id="335B3DA89620DF95B5299FA3C04D1B2B" pageId="0" pageNumber="1">METATERGA (Figs 5A, 6A, C, D, G): With 2 transverse rows of setae/tubercles/cones/spines, number in each row varying between species, lateral cones/spines of posterior row often longer and larger than inner and/or mesal ones. Suture (transverse sulcus) on metaterga present, usually conspicuous on body rings 5-17 in all species. Mid-dorsal (axial) line missing. Pleurosternal carinae forming complete, tooth-liked crests on ring 2, small ridges on ring 3, missing on remaining body rings.</paragraph>
<paragraph id="3EC4D0B4898A69783D7F0B44ED64BF85" pageId="0" pageNumber="1">
PARATERGA (Figs 5A, B, D, 6
<normalizedToken id="03672B9584594AD9FF3334328145D71D" originalValue="A–E">A-E</normalizedToken>
, G, H): Wing-shaped, well-elevated, directed caudolaterad. Callus and shoulder present, usually conspicuous, sometimes inconspicuous. Anterior margin with two distinct notches; on body rings 9, 10, 12, 13, 15-18 a tiny denticle present on lateral margin, near tip (but denticle virtually absent in some species). Elevation of paraterga in male usually higher than in female. Posterior edge concave. Tip pointed and sharp. Ozopore visible from above, round.
</paragraph>
<paragraph id="7AEF4DB6EE2EE8C1D1FA781E1D5D131A" pageId="0" pageNumber="1">TELSON (Fig. 6F, G, N): Epiproct often flattened ventrally, tip often truncate or subtruncate, sometimes emarginate; lateral tubercles often conspicuous, sometimes inconspicuous; apical tubercles usually evident, sometimes indistinct, apically with two pairs of setae. Paraprocts convex. Hypoproct often subtrapeziform, sometimes subsemicircular, rarely subtriangular; caudal margin often round, sometimes emarginate, with two small setiferous tubercles.</paragraph>
<paragraph id="7ADDF6A269AD18E0C073C9C1A77A1897" pageId="0" pageNumber="1">STERNA (Figs 5E, 6K): Sparsely setose; cross-impressions faint, usually shallow, rarely quite deep. Sternal lobe between male coxae 4 usually subtrapeziform, subrectangular, subsemicircular or round, varying between species and sometimes varying between populations, usually with two pores on posterior side.</paragraph>
<paragraph id="37270DB3B1EEFF2F5C713A8775E30558" pageId="0" pageNumber="1">
LEGS (Figs 5F, 6J, M): Very long and slender. Podomere lengths 3 = 6&gt; 5&gt; 2 = 4&gt; 1&gt; 7. Male femora 5 and 6 modified, often humped ventrally in middle part, sometimes swollen (exception:
<taxonomicName id="DBCA36C3B19DC4DC9461FEA8B6BB7736" lsidName="D. terae" pageId="0" pageNumber="1" rank="species" species="terae">D. terae</taxonomicName>
, without modification).
</paragraph>
<paragraph id="D84F91AC4C9E682393DD658673E0BD84" pageId="0" pageNumber="1">GONOPODS (Figs 6L, 7, 8): Coxa (cx) usually subequal in length to or longer than prefemur, with distoanterior group of seta. Prefemur (pfe) almost half or 2/3 as long as femur. Femur often long. Seminal groove (sg) running entirely on mesal surface of femur. Mesal sulcus (ms) conspicuous, deep; lateral sulcus (ls) conspicuous, usually deep, sometimes shallow. Postfemur shorter than femur, conspicuous. Solenophore (sph) variable between species: lamina lateralis (ll) swollen, sometimes with furrow(s) anterolaterally, sometimes with conspicuous ventral lobe (vll) or ventral ridge (vrl); lamina medialis (lm) variously species-specifically modified, consisting of one process (plm) and two lobes (distal lobe = dlm and broad lobe = blm, dlm and blm demarcated by indentation). Solenomere (sl) usually long, sometimes short.</paragraph>
</subSubSection>
<subSubSection id="48F136789B02CBCEF5732D5456139BF6" pageId="0" pageNumber="1" type="distribution">
<paragraph id="E78E6A7AF419A683FD1C5C053E4CFF38" pageId="0" pageNumber="1">Distribution and habitat.</paragraph>
<paragraph id="0C23CFD6114B3FA36AB25698A89AC4C6" pageId="0" pageNumber="1">
From south China to Malaysia. Most species seem to be local endemics (only
<taxonomicName id="52935CFE5F1C68402B9CF8E8995BB9AB" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
is dispersed, certainly by anthropochory, through mainland Southeast Asia and in many islands).
<taxonomicName id="170659F066F05987FBA15A1EB0EBA2F9" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
specimens were usually found by ASRU personnel in limestone habitats or on granitic mountains, and some were seen crawling on rocks or vegetation or tree branches (Figure 4).
</paragraph>
</subSubSection>
<subSubSection id="ECD4ED091E662022E32812BE8F2DE2B3" pageId="0" pageNumber="1" type="key">
<paragraph id="9338860B203E67C358DA0E9225A75FAD" pageId="0" pageNumber="1">
Key to species of
<taxonomicName id="4B76EA7BDE893E151A7ADE3525C18F5E" class="Diplopoda" family="Paradoxosomatidae" genus="Desmoxytes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Desmoxytes" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">Desmoxytes</taxonomicName>
s.s.
</paragraph>
<paragraph id="3D9E01AB9BC3923F82948A394A422AC3" pageId="0" pageNumber="1">
<table id="48F38F04D0C0F9B73B820F0F56A9559C" pageId="0" pageNumber="1">
<tr id="814742D010DD392FB566DD89627390AD" pageId="0" pageNumber="1">
<td id="C321500159B1A5E11242B2F66682E889" colspan="1" pageId="0" pageNumber="1" rowspan="1">90H, I</td>
<td id="108BBB0414CC4A768BD6F5CB9F838060" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="1983E14845EAC3D4C988FA4F036AB27E" lsidName="D. terae" pageId="0" pageNumber="1" rank="species" species="terae">D. terae</taxonomicName>
(Jeekel, 1964)
</td>
</tr>
<tr id="88FED402C86F260D5928EA21F89170F9" pageId="0" pageNumber="1">
<td id="9063DF7B907EB53E2766B120CF24B530" colspan="1" pageId="0" pageNumber="1" rowspan="1">11H, I</td>
<td id="5CFC79E48A8EAC7A143116B5FD70826C" colspan="1" pageId="0" pageNumber="1" rowspan="1">2</td>
</tr>
<tr id="276E927B2002E83E49CBC9763788828A" pageId="0" pageNumber="1">
<td id="360BE47F92173156991444B9B69AC67B" colspan="1" pageId="0" pageNumber="1" rowspan="1">18A, B38A, B18B, C38B, C</td>
<td id="5BD9387CAB18FE44AEFF5A9F55B35A87" colspan="1" pageId="0" pageNumber="1" rowspan="1">3</td>
</tr>
<tr id="80BACD5966F994A3D3AA9A65AE6BB715" pageId="0" pageNumber="1">
<td id="C3B6622834579396E4212BF7F67436FB" colspan="1" pageId="0" pageNumber="1" rowspan="1">56A, B76A, B56B, C76B, C</td>
<td id="086D99C4AED3303777B9F9CD9F0EFC4E" colspan="1" pageId="0" pageNumber="1" rowspan="1">12</td>
</tr>
<tr id="00ABB122F3DBAD08621D4202F75BECB9" pageId="0" pageNumber="1">
<td id="031EC178F2D2D56F02F4E7B7529EAE56" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation id="40350719599D862E142F1DBA7357CE0C" author="Srisonchai, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="93 - 113" title="Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://dx.doi.org/10.11646/zootaxa.4170.1.4" volume="4710" year="2016">Srisonchai et al. 2016</bibRefCitation>
</td>
<td id="261FE620FE958BE2E7FE43683C49786D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="04ECA5D8E8DC608B47640CE2E85D6E21" lsidName="D. des" pageId="0" pageNumber="1" rank="species" species="des">D. des</taxonomicName>
Srisonchai et al., 2016
</td>
</tr>
<tr id="F298E32C5073191573D08BA471CFD9A9" pageId="0" pageNumber="1">
<td id="883EFCE404621EE6CA8BB5785531F780" colspan="1" pageId="0" pageNumber="1" rowspan="1">
3
<normalizedToken id="278C1765587456A90D92A20A8D22E10F" originalValue="A–D">A-D</normalizedToken>
</td>
<td id="BC3E52D02DFA505F89A3DFA071962F0F" colspan="1" pageId="0" pageNumber="1" rowspan="1">4</td>
</tr>
<tr id="06D567FE8BE7FA69C740CB4228ADF8A8" pageId="0" pageNumber="1">
<td id="5711F40AEEE31E5A4C6296EA509185E7" colspan="1" pageId="0" pageNumber="1" rowspan="1">
18A24A44A18
<normalizedToken id="DD5C70E88FA39F8C0C3327996DB18AD4" originalValue="A–Cll">A-Cll</normalizedToken>
21B, D22F47B48E
</td>
<td id="1C1358A2D9276B925CA1C4DBC6CE5638" colspan="1" pageId="0" pageNumber="1" rowspan="1">5</td>
</tr>
<tr id="6778CEDA254729271780AD9B134A37DF" pageId="0" pageNumber="1">
<td id="B9E7C53D556E188D246FF7C9DC18DE8B" colspan="1" pageId="0" pageNumber="1" rowspan="1">
38A50A56A38
<normalizedToken id="9868A70E9C5574F16A35A1B3E41400C5" originalValue="A–Cll">A-Cll</normalizedToken>
41B53B
</td>
<td id="4ABE27A26BFE9A4A9A6DEE372B287860" colspan="1" pageId="0" pageNumber="1" rowspan="1">10</td>
</tr>
<tr id="DBE889C76E969DA4544BCD45C6E0EA91" pageId="0" pageNumber="1">
<td id="6AD36DB3F47D245D229860831E949E7B" colspan="1" pageId="0" pageNumber="1" rowspan="1">
llvrl27Cdlm27
<normalizedToken id="B06988ED6D495569CE21A0095407C896" originalValue="B–D">B-D</normalizedToken>
28Edlmblm27D28E
</td>
<td id="4F911ED4616C636F8C3289938BEA083D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="9B12DAE6ADBFC9C60E718955EA194FE0" lsidName="D. corythosaurus" pageId="0" pageNumber="1" rank="species" species="corythosaurus">D. corythosaurus</taxonomicName>
sp. n.
</td>
</tr>
<tr id="ABE10D33FF4FDD44DE05B1C202E11F15" pageId="0" pageNumber="1">
<td id="F3FE9B103A44EE3FB0F3B91FA0119D0A" colspan="1" pageId="0" pageNumber="1" rowspan="1">llvrl21C22Cdlm21B22E47D48Edlmblm21D22D47D48E</td>
<td id="515404582A1DF5D24C5DDD323642093E" colspan="1" pageId="0" pageNumber="1" rowspan="1">6</td>
</tr>
<tr id="9338A06F5AF68B45C28FE7795A7ADDC2" pageId="0" pageNumber="1">
<td id="D1908CC631DA4B80679EF4377F14A2EC" colspan="1" pageId="0" pageNumber="1" rowspan="1">ll47B48E</td>
<td id="283447E2C40C53F0F2C051C15E297093" colspan="1" pageId="0" pageNumber="1" rowspan="1">7</td>
</tr>
<tr id="4A705B2764C67464D83F030802D382D6" pageId="0" pageNumber="1">
<td id="3FB04C654A49E0FD4D153E6EF256CB6B" colspan="1" pageId="0" pageNumber="1" rowspan="1">ll34E35F</td>
<td id="DA46502DEB5E6D11A52681D33636A863" colspan="1" pageId="0" pageNumber="1" rowspan="1">8</td>
</tr>
<tr id="B95F75E6F63B9820FD2914F648252984" pageId="0" pageNumber="1">
<td id="E71646A501BFF5C66126130B20D98D1B" colspan="1" pageId="0" pageNumber="1" rowspan="1">61Bplmdlm64B, C65C, Ddlm64B65D</td>
<td id="63A90FD25A23B9B2E0A7549EEACBDD54" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="2EDB52858A0370A24180D2B919D4D71C" lsidName="D. perakensis" pageId="0" pageNumber="1" rank="species" species="perakensis">D. perakensis</taxonomicName>
sp. n.
</td>
</tr>
<tr id="92C772A4F5929BF53DD98356359296BF" pageId="0" pageNumber="1">
<td id="FF93A99E9401AADB5683808BDB93A7FE" colspan="1" pageId="0" pageNumber="1" rowspan="1">44Bplmdlm47B48Edlm47B48E</td>
<td id="55CA3D1664FB2BDD4D3B035CC5A3E4EF" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="4CA2A0421B6A3597AA6EBE9E1FC76E01" lsidName="D. flabella" pageId="0" pageNumber="1" rank="species" species="flabella">D. flabella</taxonomicName>
sp. n.
</td>
</tr>
<tr id="199C7B7F813D4377D2679215343FED61" pageId="0" pageNumber="1">
<td id="F37FE8873C42E35EACF85B838DC1050A" colspan="1" pageId="0" pageNumber="1" rowspan="1">18E, F</td>
<td id="9353447933EFB90E456C25BE0FB34D21" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="4D29E07DF4BED94EBE2DDE7C5A175FB1" lsidName="D. cervina" pageId="0" pageNumber="1" rank="species" species="cervina">D. cervina</taxonomicName>
(Pocock, 1895)
</td>
</tr>
<tr id="3720D864AA44D5652FB200F925F5DB5E" pageId="0" pageNumber="1">
<td id="C9F755C03E8A1D04319395A0E0ECFEE6" colspan="1" pageId="0" pageNumber="1" rowspan="1">10E, F</td>
<td id="257C870F389BB43D0CDFA144E20AA445" colspan="1" pageId="0" pageNumber="1" rowspan="1">9</td>
</tr>
<tr id="AA5B0891802959B7CBCE1AEC757EB899" pageId="0" pageNumber="1">
<td id="1B4E9A006B79CACE83E7184E44D95027" colspan="1" pageId="0" pageNumber="1" rowspan="1">10Als13Ell13B, C14C, Evrl13C14Cdlm13B14E12A, B</td>
<td id="734BAF43124462D99F9C2C3CA284ED8C" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="DED3D2E451F464800B948B64CF112F6A" lsidName="D. aurata" pageId="0" pageNumber="1" rank="species" species="aurata">D. aurata</taxonomicName>
sp. n.
</td>
</tr>
<tr id="2A519A3186402B22D0B06D3310CA20A0" pageId="0" pageNumber="1">
<td id="D9B2329A785FAA91F7D5450111EDF774" colspan="1" pageId="0" pageNumber="1" rowspan="1">31Als34Ell34B, C35C, Evrl34C35Cdlm34B35E33A</td>
<td id="720C417D40295F653C2C74E9E5DA25B9" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="4A92DDD5426D1A81B7C82CAE39C54CAE" lsidName="D. delfae" pageId="0" pageNumber="1" rank="species" species="delfae">D. delfae</taxonomicName>
(Jeekel, 1964)
</td>
</tr>
<tr id="3BFA1F78EF0D9FB083E4881058A67B5E" pageId="0" pageNumber="1">
<td id="CCFA6037ADA263AF288ED2C58908FD52" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<bibRefCitation id="BB0BD4583C651410901F2EFFFF4AC81A" author="Srisonchai, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="93 - 113" title="Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://dx.doi.org/10.11646/zootaxa.4170.1.4" volume="4710" year="2016">Srisonchai et al 2016</bibRefCitation>
<bibRefCitation id="D7C5C5E1F8221C103DD28D7AEA64577C" author="Srisonchai, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="93 - 113" title="Four colorful new species of dragon millipedes, genus Desmoxytes Chamberlin, 1923, from northern Thailand (Diplopoda: Polydesmida: Paradoxosomatidae)." url="http://dx.doi.org/10.11646/zootaxa.4170.1.4" volume="4710" year="2016">Srisonchai et al. 2016</bibRefCitation>
</td>
<td id="C427F52D302FB5AEC5B7875887B72530" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="8EE1551C2729703645F35B8E7BDF02D3" lsidName="D. pinnasquali" pageId="0" pageNumber="1" rank="species" species="pinnasquali">D. pinnasquali</taxonomicName>
Srisonchai et al., 2016
</td>
</tr>
<tr id="A519F86742F9EB3FE0C3A8FC652CB431" pageId="0" pageNumber="1">
<td id="057D6B1D9C6385B8BBE21A0864A29B07" colspan="1" pageId="0" pageNumber="1" rowspan="1">38A69A39F, G70F, G</td>
<td id="2CA21A7BEF8050A740D2B6A5C849C50E" colspan="1" pageId="0" pageNumber="1" rowspan="1">11</td>
</tr>
<tr id="55201929DD9D931D35C327DF20901E2F" pageId="0" pageNumber="1">
<td id="6F243F0643496B9DBD795619839C5209" colspan="1" pageId="0" pageNumber="1" rowspan="1">70C, D</td>
<td id="ABDF677AD833DAC041AD0CA049300DFA" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="F30C2086B76406E5E806E27DD77F18A9" lsidName="D. planata" pageId="0" pageNumber="1" rank="species" species="planata">D. planata</taxonomicName>
(Pocock, 1895)
</td>
</tr>
<tr id="AB37A85D6F41FD090A4D677617369C2B" pageId="0" pageNumber="1">
<td id="4A7B7262933C3D84C71EBBCAED47FF78" colspan="1" pageId="0" pageNumber="1" rowspan="1">39C, D</td>
<td id="2312DD3B44B1F6DFEAFAA665803856A3" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="9E5E3BF233ADA6000DE1131309B3D194" lsidName="D. euros" pageId="0" pageNumber="1" rank="species" species="euros">D. euros</taxonomicName>
sp. n.
</td>
</tr>
<tr id="7945E9BBA3504E549965AA3D92F169AE" pageId="0" pageNumber="1">
<td id="0B7DC2B927F6658AA3667FCABF2C120B" colspan="1" pageId="0" pageNumber="1" rowspan="1">76B, C</td>
<td id="71A03A653BF6B99BFBAD3048A6C63C05" colspan="1" pageId="0" pageNumber="1" rowspan="1">13</td>
</tr>
<tr id="02DB19590867E30922BA1A16DD6A3F9B" pageId="0" pageNumber="1">
<td id="B6CEE365C760533E252DC84773C58534" colspan="1" pageId="0" pageNumber="1" rowspan="1">50B, C56B, C</td>
<td id="E214525D0AB1676B3EBF662FD9C91B63" colspan="1" pageId="0" pageNumber="1" rowspan="1">16</td>
</tr>
<tr id="5BE958A659F53B7466DE6F3320D98932" pageId="0" pageNumber="1">
<td id="9DF62488C91645D977E633E15ED705F1" colspan="1" pageId="0" pageNumber="1" rowspan="1">llvll87plm87E</td>
<td id="AF41DD46682905A7E2080D43331B7F83" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="EB4112F7ADB64B6C1ACE5780AE431838" lsidName="D. taurina" pageId="0" pageNumber="1" rank="species" species="taurina">D. taurina</taxonomicName>
(Pocock, 1895)
</td>
</tr>
<tr id="316C657605D3A0E97074C1C18C446F7E" pageId="0" pageNumber="1">
<td id="376B7A39A8D158337D4AB55BCB50C14C" colspan="1" pageId="0" pageNumber="1" rowspan="1">llvll79F81Cplm15B79E83B</td>
<td id="6BB33B54006AB91766D5A32EE278805D" colspan="1" pageId="0" pageNumber="1" rowspan="1">14</td>
</tr>
<tr id="FEE883A7A55ACFFE4A916DBE2E9DAEEA" pageId="0" pageNumber="1">
<td id="0F9F675335A761E78DF2546179BBFADB" colspan="1" pageId="0" pageNumber="1" rowspan="1">vll83C, Ddlm83B</td>
<td id="B68C3DB035FF51811703ABFF1DE1BBC4" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="C7E05F0F5684B03B3AB1800E140C9958" lsidName="D. takensis" pageId="0" pageNumber="1" rank="species" species="takensis">D. takensis</taxonomicName>
Srisonchai et al., 2016
</td>
</tr>
<tr id="106A84FD4DAE8E73A41C6C89555F376B" pageId="0" pageNumber="1">
<td id="5DA350D92EA5EA84915DEE4BADE26AC0" colspan="1" pageId="0" pageNumber="1" rowspan="1">vll15C79Fdlm15B79E</td>
<td id="8B38D677FF1097B89E32ABFA0B92B7E9" colspan="1" pageId="0" pageNumber="1" rowspan="1">15</td>
</tr>
<tr id="2C4D65789426B5E89ED3CA5B70D5DA3B" pageId="0" pageNumber="1">
<td id="BCD95C1C03004FC4CC51A73987EFCCED" colspan="1" pageId="0" pageNumber="1" rowspan="1">ll79E81E79D81Fplm80</td>
<td id="701F871DA485100E4401408734F879B7" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="CD45765625B2F3D5758BE4BCDCBC80E8" lsidName="D. purpurosea" pageId="0" pageNumber="1" rank="species" species="purpurosea">D. purpurosea</taxonomicName>
Enghoff et al., 2007
</td>
</tr>
<tr id="1AD3E56F2BDEF861D9B01B512C54E411" pageId="0" pageNumber="1">
<td id="0A4A939244046597115E9F4B5398411F" colspan="1" pageId="0" pageNumber="1" rowspan="1">ll15B15Eplm15B</td>
<td id="D71CD1D1E6283541EEBD62F79D11D16A" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="4E85CCF60211B7F728686528D1FAAA34" lsidName="D. breviverpa" pageId="0" pageNumber="1" rank="species" species="breviverpa">D. breviverpa</taxonomicName>
Srisonchai et al., 2016
</td>
</tr>
<tr id="0FD11CFEED5F2FFC5B9F895FC99187E6" pageId="0" pageNumber="1">
<td id="8DF278A1B5CEF89AA7F090C55DA08009" colspan="1" pageId="0" pageNumber="1" rowspan="1">50B, Cblm53B54E</td>
<td id="123D17D3BF730219116D1409E7B1134D" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="0A66AD17B32731EF8211E06CB6374F00" lsidName="D. golovatchi" pageId="0" pageNumber="1" rank="species" species="golovatchi">D. golovatchi</taxonomicName>
sp. n.
</td>
</tr>
<tr id="7A0C2916796F17EBD95CB10A3E332F9E" pageId="0" pageNumber="1">
<td id="B8AAFDA81BDE7D6AD0287C0E9224F9B7" colspan="1" pageId="0" pageNumber="1" rowspan="1">56B, C94B, Cblm59B97B</td>
<td id="F6ACD3286F36D0F4D407E7EDBD308A99" colspan="1" pageId="0" pageNumber="1" rowspan="1">17</td>
</tr>
<tr id="A6FFCC029C05858AE5197BCBB031AE11" pageId="0" pageNumber="1">
<td id="40C32148B8E4E5DCF0972942D27F72B3" colspan="1" pageId="0" pageNumber="1" rowspan="1">
55
<normalizedToken id="151DB0DDF5252F8DB65E56BFF85B4230" originalValue="A–C56A–C">A-C56A-C</normalizedToken>
57H, I58
</td>
<td id="E8BAD8A7E48FECD715A9381CBC999243" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="8DD41C1F616B98321DF0C8473CC010AC" lsidName="D. octoconigera" pageId="0" pageNumber="1" rank="species" species="octoconigera">D. octoconigera</taxonomicName>
sp. n.
</td>
</tr>
<tr id="2DE21B25BA7A29C013B646FF4F0C4876" pageId="0" pageNumber="1">
<td id="94371E0BC1B0761BB3DED68286A15019" colspan="1" pageId="0" pageNumber="1" rowspan="1">
93A, B, D94
<normalizedToken id="0926149FF58B803E86897350759C54DD" originalValue="A–C">A-C</normalizedToken>
95H, I96
</td>
<td id="3CBA6746DEE09A05DAD95AC0540BF363" colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName id="39299D9377D4089D1D935ACECFE997C6" lsidName="D. waepyanensis" pageId="0" pageNumber="1" rank="species" species="waepyanensis">D. waepyanensis</taxonomicName>
sp. n.
</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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