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<document id="7296C30D792CE581421D2C92AD5CC53F" ID-CLB-Dataset="42161" ID-DOI="10.11646/zootaxa.3887.2.2" ID-GBIF-Dataset="395faf4e-7cb0-45b0-9f1a-348da32744f4" ID-ISSN="1175-5326" ID-Zenodo-Dep="251053" ID-ZooBank="D9275063-BD74-498E-B504-1421B50114A5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460975774690" checkinUser="plazi" docAuthor="Greenwalt, Dale E. &amp; Bechly, Günter" docDate="2014" docId="40446A46FFBF670AFF4C060A8D52FD81" docLanguage="en" docName="zt03887p156.pdf" docOrigin="Zootaxa 3887 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eolestes ramosus Greenwalt &amp; Bechly, 2014, n. sp." docType="treatment" docVersion="10" lastPageNumber="145" masterDocId="BC7D123EFFB86703FFDB02668923FFBC" masterDocTitle="A re-description of the fossil damselfly Eolestes syntheticus Cockerell, 1940 (Odonata: Zygoptera: Eolestidae n. fam.) with description of new taxa from the Eocene of North America" masterLastPageNumber="156" masterPageNumber="138" pageNumber="145" updateTime="1732711539088" updateUser="ExternalLinkService">
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<mods:title id="3F1EB40CD8031D921126C36D8920FF4C">A re-description of the fossil damselfly Eolestes syntheticus Cockerell, 1940 (Odonata: Zygoptera: Eolestidae n. fam.) with description of new taxa from the Eocene of North America</mods:title>
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<mods:namePart id="04C7E9EFAA091530B66FB8A402A1E1C1">Greenwalt, Dale E.</mods:namePart>
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<mods:namePart id="13526DF664749B26F09716863DC3B2AF">Bechly, Günter</mods:namePart>
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<mods:date id="1795C5E78F8165F3DC364F52E42F2F72">2014</mods:date>
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<treatment id="40446A46FFBF670AFF4C060A8D52FD81" ID-DOI="http://doi.org/10.5281/zenodo.6141526" ID-GBIF-Taxon="119593984" ID-Zenodo-Dep="6141526" LSID="urn:lsid:plazi:treatment:40446A46FFBF670AFF4C060A8D52FD81" httpUri="http://treatment.plazi.org/id/40446A46FFBF670AFF4C060A8D52FD81" lastPageId="9" lastPageNumber="145" pageId="7" pageNumber="145" scope_order="Odonata">
<subSubSection id="80F788DBFFBF6704FF4C060A8853FB1B" pageId="7" pageNumber="145" type="nomenclature">
<paragraph id="C852DB50FFBF6704FF4C060A8891FB3A" blockId="7.[151,434,1132,1192]" box="[151,434,1132,1158]" pageId="7" pageNumber="145">
<heading id="931A6C3CFFBF6704FF4C060A8891FB3A" bold="true" box="[151,434,1132,1158]" fontSize="11" level="1" pageId="7" pageNumber="145" reason="1">
<emphasis id="FA990742FFBF6704FF4C060A8891FB3A" bold="true" box="[151,434,1132,1158]" pageId="7" pageNumber="145">
<taxonomicName id="0FEDA0D3FFBF6704FF4C060A884BFB3A" box="[151,360,1132,1158]" class="Insecta" genus="Eolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="ramosus" status="sp. nov.">
<emphasis id="FA990742FFBF6704FF4C060A884BFB3A" bold="true" box="[151,360,1132,1158]" italics="true" pageId="7" pageNumber="145">Eolestes ramosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E1AABA39FFBF6704FEB4060A8891FB3A" box="[367,434,1132,1158]" pageId="7" pageNumber="145" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C852DB50FFBF6704FF4C06E98853FB1B" blockId="7.[151,434,1132,1192]" box="[151,368,1167,1192]" pageId="7" pageNumber="145">
(
<figureCitation id="50D6C7D5FFBF6704FF4506E9882AFB14" box="[158,265,1167,1192]" captionStart="FIGURE 5" captionStartId="8.[151,250,1475,1497]" captionTargetBox="[151,1436,193,1453]" captionTargetId="figure@8.[151,1436,193,1454]" captionTargetPageId="8" captionText="FIGURE 5. Wings of Eolestes spp.: (A) left hindwing of E. syntheticus (holotype UCM 19170); (B) E. syntheticus (USNM 559049); (C) E. ramosus (holotype USNM 559047). Scale bars = 5 mm." httpUri="https://zenodo.org/record/251058/files/figure.png" pageId="7" pageNumber="145">Figures 5</figureCitation>
C, 6C, 7)
</paragraph>
</subSubSection>
<subSubSection id="80F788DBFFBF6704FF4C06B189FCFAE0" pageId="7" pageNumber="145" type="materials_examined">
<paragraph id="C852DB50FFBF6704FF4C06B18D1DFB53" blockId="7.[151,1437,1239,2020]" box="[151,1086,1239,1264]" pageId="7" pageNumber="145">
<emphasis id="FA990742FFBF6704FF4C06B18828FB4C" bold="true" box="[151,267,1239,1264]" pageId="7" pageNumber="145">
<typeStatus id="175665F2FFBF6704FF4C06B18825FB4C" box="[151,262,1239,1264]" pageId="7" pageNumber="145" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="AEFC4395FFBF6704FEC906B18840FB53" LSID="urn:lsid:biocol.org:col:34871" box="[274,355,1239,1263]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="7" pageNumber="145">USNM</collectionCode>
559047, National Museum of Natural History, Washington, D.C.
</paragraph>
<paragraph id="C852DB50FFBF6704FF1C069D89FCFAE0" blockId="7.[151,1437,1239,2020]" pageId="7" pageNumber="145">
<emphasis id="FA990742FFBF6704FF1C069D8B22FAA8" bold="true" box="[199,513,1275,1300]" pageId="7" pageNumber="145">
<typeStatus id="175665F2FFBF6704FF1C069D8821FAA8" box="[199,258,1275,1300]" pageId="7" pageNumber="145">Type</typeStatus>
locality and stratum.
</emphasis>
Disbrow Creek site, Middle Fork of the Flathead River, Pinnacle, Montana,
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. Coal Creek Member of the Kishenehn Formation, early Middle Eocene, 46.2±0.4 or 43.5±4.9 mya (
<bibRefCitation id="AC7CA6A1FFBF6704FACC077989F0FAE0" author="Constenius" pageId="7" pageNumber="145" refString="Constenius, K. N. (1996) Late Paleogene extensional collapse of the Cordilleran foreland fold and thrust belt. Geological Society of America Bulletin, 108, 20 - 39. http: // dx. doi. org / 10.1130 / 0016 - 7606 (1996) 108 &lt;0020: lpecot&gt; 2.3. co; 2" type="journal article" year="1996">Constenius, 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="80F788DBFFBF6704FF1C07018A7FFA18" pageId="7" pageNumber="145" type="etymology">
<paragraph id="C852DB50FFBF6704FF1C07018A7FFA18" blockId="7.[151,1437,1239,2020]" pageId="7" pageNumber="145">
<emphasis id="FA990742FFBF6704FF1C0701886DFA3C" bold="true" box="[199,334,1383,1408]" pageId="7" pageNumber="145">Etymology.</emphasis>
Species name
<taxonomicName id="0FEDA0D3FFBF6704FE22070E8B7BFAC3" box="[505,600,1384,1407]" class="Insecta" genus="Eolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="FA990742FFBF6704FE22070E8B7BFAC3" box="[505,600,1384,1407]" italics="true" pageId="7" pageNumber="145">ramosus</emphasis>
</taxonomicName>
from the Latin word
<taxonomicName id="0FEDA0D3FFBF6704FC95070E8A88FAC3" box="[846,939,1384,1407]" class="Insecta" genus="Eolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="ramosus">ramosus</taxonomicName>
(branching), an indication of the branching of the supplementary longitudinal sectors in the MP-CuA field.
</paragraph>
</subSubSection>
<subSubSection id="80F788DBFFBF6704FF1C07C98BF9F9B3" pageId="7" pageNumber="145" type="diagnosis">
<paragraph id="C852DB50FFBF6704FF1C07C98BF9F9B3" blockId="7.[151,1437,1239,2020]" pageId="7" pageNumber="145">
<emphasis id="FA990742FFBF6704FF1C07C98862FA74" bold="true" box="[199,321,1455,1480]" pageId="7" pageNumber="145">Diagnosis.</emphasis>
Differs from
<taxonomicName id="0FEDA0D3FFBF6704FE3807D68BA2FA7B" box="[483,641,1455,1479]" class="Insecta" genus="Eolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="7" pageNumber="145" phylum="Arthropoda" rank="species" species="syntheticus">
<emphasis id="FA990742FFBF6704FE3807D68BA2FA7B" box="[483,641,1455,1479]" italics="true" pageId="7" pageNumber="145">E. syntheticus</emphasis>
</taxonomicName>
by the following characters: 1) CuP origin at instead of distal of the separation of AA and AA; 2) three supplementary longitudinal sectors in the cubital field vs. two; 3) 24 rows of cells between distal parts of IR2 and
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3/4 vs. 12.
</paragraph>
</subSubSection>
<subSubSection id="80F788DBFFBF670AFF1C047D8D52FD81" lastPageId="9" lastPageNumber="147" pageId="7" pageNumber="145" type="description">
<paragraph id="C852DB50FFBF670AFF1C047D8D52FD81" blockId="7.[151,1437,1239,2020]" lastBlockId="9.[151,1437,151,573]" lastPageId="9" lastPageNumber="147" pageId="7" pageNumber="145">
<emphasis id="FA990742FFBF6704FF1C047D887AF988" bold="true" box="[199,345,1563,1588]" pageId="7" pageNumber="145">Description.</emphasis>
An intact fossil damselfly wing, hyaline,
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long and
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and
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wide at nodus and widest point respectively (
<figureCitation id="50D6C7D5FFBF6704FE2104598B1CF9E4" box="[506,575,1599,1624]" captionStart="FIGURE 5" captionStartId="8.[151,250,1475,1497]" captionTargetBox="[151,1436,193,1453]" captionTargetId="figure@8.[151,1436,193,1454]" captionTargetPageId="8" captionText="FIGURE 5. Wings of Eolestes spp.: (A) left hindwing of E. syntheticus (holotype UCM 19170); (B) E. syntheticus (USNM 559049); (C) E. ramosus (holotype USNM 559047). Scale bars = 5 mm." httpUri="https://zenodo.org/record/251058/files/figure.png" pageId="7" pageNumber="145">Fig. 5</figureCitation>
C); pterostigma four cells and
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long,
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wide; pterostigmal brace oblique; distance between wing apex and pterostigma, pterostigma and nodus, nodus and arculus and arculus and base 2.43, 12.13, 5.8 and
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respectively; petiole length relative to distance from petiole to nodus = 0.44; post-pterostigmal cell area consists of two single cells proximally, three smaller cells distally and five double cells in between; two antenodal and two antesubnodal crossveins. Ax1 opposite separation of AA &amp; AA and separated from Ax2 by
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; Ax2 opposite arcular crossvein just distal to
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origin and anterior arcular crossvein;
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separates from
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+MA just below RA+
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so that
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+MA very short; supplementary antenodal crossveins absent; posterior arcular crossvein separating from discoidal cell
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below RA; nodus and subnodal bracket not preserved, subnodal crossvein apparently oblique; 14 postnodal crossveins exactly aligned with postsubnodal crossveins with exception of last four; lestine oblique vein O present four cells distal of base of
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2; base of IR1 four cells and
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distal of
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2 origin, IR1 zigzagged through first three cells, underlies a single row of cells that, proximal to pterostigma, are higher than they are wide and overlies a row of seven single cells that distally expand through six double cells and subsequently, an area of reticulated cells with two and three secondary longitudinal veins; base of
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2 two cells and
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distal of subnodus and six cells and
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distal of origin of IR2; IR2 and PR3/4 originate much closer to arculus than nodus. IR2 arched abruptly toward
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at base with crossvein opposite, one cell and
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from origin of
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3/4; IR2 underlies a row of 18 single cells basally, followed by a gradual increase in number of secondary veins until there are 11 small cells at wing margin;
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3/4 originates one cell and
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from arculus and underlies a row of single cells that transitions from cells longer than high to higher than long before double cells appear just proximal to beginning of pterostigma; MA slightly zigzagged after a point about midway between arculus and nodus, and underlies a row of cells that transitions to double cells at level of IR1 origin and then gradually to 17 small cells at wing margin; MP underlies a single row of cells that gradually transitions from square-shaped to cells higher than wide; CuA a prominent vein that leaves subdiscoidal cell
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below MP and transitions to a zigzagged pattern at about level of origin of
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2; it underlies a broad cubital field
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in height that starts with a single cell below base of MP and quickly transitions through one, two and then three secondary longitudinal veins and then back to two secondary veins at wing margin (
<figureCitation id="50D6C7D5FFB1670AFEEF02F18855FF0C" box="[308,374,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1226,1248]" captionTargetBox="[340,1247,628,1199]" captionTargetId="figure@9.[340,1248,628,1205]" captionTargetPageId="9" captionText="FIGURE 6. Line drawings of the cubital and cubito-anal fields of: (A) left hindwing of E. syntheticus (holotype UCM 19170); (B) E. syntheticus (USNM 559049); (C) E. ramosus (holotype USNM 559047). Scale bars = 5 mm." httpUri="https://zenodo.org/record/251059/files/figure.png" pageId="9" pageNumber="147">Fig. 6</figureCitation>
C); cubito-anal field consists of a single row of cells (12 distal of subnodus) that, except for the first, are higher than wide, and terminates at a point halfway between nodus and pterostigma; cubito-anal field
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in height; CuP origin at separation of AA &amp; AA (
<figureCitation id="50D6C7D5FFB1670AFCD102B98A6DFF44" box="[778,846,223,248]" captionStart="FIGURE 6" captionStartId="9.[151,250,1226,1248]" captionTargetBox="[340,1247,628,1199]" captionTargetId="figure@9.[340,1248,628,1205]" captionTargetPageId="9" captionText="FIGURE 6. Line drawings of the cubital and cubito-anal fields of: (A) left hindwing of E. syntheticus (holotype UCM 19170); (B) E. syntheticus (USNM 559049); (C) E. ramosus (holotype USNM 559047). Scale bars = 5 mm." httpUri="https://zenodo.org/record/251059/files/figure.png" pageId="9" pageNumber="147">Fig. 6</figureCitation>
), directly below Ax1; petiole well defined,
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long (from base to separation of AA and AA) and 9.3% of wing length; discoidal cell closed basally,
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long (end of posterior arcular vein to origin of MP) and
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wide (origin of arcular vein to origin of MAb) with an acute posterior internal angle of 27 degrees (Note: given the curved nature of the posterior portions of MAb and MP &amp; CuA, the angle is that between lines that align with the majority of these two veins.); distal side of discoidal cell (MAb) nearly perpendicular to RA (7 degrees from vertical relative to RA), and
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in length; anterior, posterior and basal (posterior arculus) sides
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,
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and
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long; Ratio of lengths of anterior and posterior sides of discoidal cell = 0.36; subdiscoidal cell elongate with no fusion of CuP &amp; AA to posterior wing margin distal of CuP; MP only partially preserved but with apparently no or very little arch as it leaves discoidal cell at angles of 93 and 76 degrees from MAb and vertical respectively.
</paragraph>
</subSubSection>
</treatment>
</document>