423 lines
53 KiB
XML
423 lines
53 KiB
XML
<document ID-CLB-Dataset="53339" ID-DOI="http://dx.doi.org/10.3897/vz.73.e104494" ID-GBIF-Dataset="6d5bfe91-627c-4dd3-93ef-cad0eb2c3f52" ID-Pensoft-Pub="2625-8498-73-499" ID-Pensoft-UUID="8ACE3A2EE7025F8CBB56686CA028BB3A" ID-ZooBank="8F9CC156D4BE4C61A2A6337B1DC0C23A" ModsDocID="2625-8498-73-499" checkinTime="1684941527018" checkinUser="pensoft" docAuthor="Narayanan, Surya, Christopher, Peter, Raman, Kothandapani, Mukherjee, Nilanjan, Prabhu, Ponmudi, Lenin, Maniezhilan, Vimalraj, Sivangnanaboopathidoss & Deepak, V." docDate="2023" docId="3F9574FC242D515BA53EFDD1EC3EDF85" docLanguage="en" docName="VertZool 73: 499-512" docOrigin="Vertebrate Zoology 73" docPubDate="2023-05-23" docSource="http://dx.doi.org/10.3897/vz.73.e104494" docTitle="Hemidactylus pakkamalaiensis Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak 2023, sp. nov." docType="treatment" docUuid="820BD788-BD9D-491B-BBBB-0E1C0649ED6E" docUuidSource="ZooBank" docVersion="4" id="8ACE3A2EE7025F8CBB56686CA028BB3A" lastPageNumber="499" masterDocId="8ACE3A2EE7025F8CBB56686CA028BB3A" masterDocTitle="A new species of rock-dwelling Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from the southern Eastern Ghats, India" masterLastPageNumber="512" masterPageNumber="499" pageNumber="499" updateTime="1732834532847" updateUser="ExternalLinkService">
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<mods:titleInfo id="7130BE935BF9E38592E846EE3A83782E">
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<mods:title id="A4A7BA30C019D1DF5F98B044174A2E30">A new species of rock-dwelling Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from the southern Eastern Ghats, India</mods:title>
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<mods:namePart id="5C9BF0920039EF27A5E9AB07B101087B">Narayanan, Surya</mods:namePart>
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<mods:affiliation id="5F4BAF5EDD10C4E77BA7537650FCD4DE">SM Sehgal Centre for Biodiversity and Conservation, Ashoka Trust for Research in Ecology and the Environment (ATREE), Royal Enclave, Srirampura, Bangalore, Karnataka, 560064, India</mods:affiliation>
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<mods:namePart id="990E4C5F1E4C0FC24C1D9A8B483C89BB">Christopher, Peter</mods:namePart>
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<mods:nameIdentifier id="96D1D4D42E3D9F79C1B44B7E4DEA1BC9" type="ORCID">https://orcid.org/0000-0001-9769-1319</mods:nameIdentifier>
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<mods:affiliation id="8DB62396085DC7A0CB585B143E31EC16">Indigeneous Biodiversity Foundation, Mission Street, Puducherry, 605001, India</mods:affiliation>
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<mods:namePart id="08BEEE45519F3C1EFD39005F205410FD">Raman, Kothandapani</mods:namePart>
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<mods:affiliation id="DFFCD6056BFF7C23DF4055E07AECD3FC">Indigeneous Biodiversity Foundation, Mission Street, Puducherry, 605001, India</mods:affiliation>
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<mods:namePart id="459C4B2B917068FE4BF5C5F5EE2D279B">Mukherjee, Nilanjan</mods:namePart>
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<mods:affiliation id="3933A8B40A021FA034C3D11234963BA9">Felis Creations, Sadashiva Nagar, Bengaluru, Karnataka, 560080, India</mods:affiliation>
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<mods:namePart id="496165096336A206075B7C6BDC734C56">Prabhu, Ponmudi</mods:namePart>
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<mods:affiliation id="A51D44F3330F11304034D8C9569B55A1">Indigeneous Biodiversity Foundation, Mission Street, Puducherry, 605001, India</mods:affiliation>
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<mods:namePart id="31B262302CC4E5709A66075874E96F66">Lenin, Maniezhilan</mods:namePart>
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<mods:affiliation id="E6CD05AF8B6D601079DF3A0AE7F5BA47">Kunnathur Road, Perundurai, Tamil Nadu, 638052, India</mods:affiliation>
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<mods:namePart id="3A3F0F28D3C50F4F9AA0752E9F91AC2E">Vimalraj, Sivangnanaboopathidoss</mods:namePart>
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<mods:affiliation id="9597E97FB81218960B8FB9469D17C5D6">Indigeneous Biodiversity Foundation, Mission Street, Puducherry, 605001, India</mods:affiliation>
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<mods:namePart id="0B81065A0621146B6B44449969E65A07">Deepak, V.</mods:namePart>
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<mods:nameIdentifier id="1E65A9B2BB6633E57847B153899C2F85" type="ORCID">https://orcid.org/0000-0002-8826-9367</mods:nameIdentifier>
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<mods:affiliation id="818399E28EA7C7FDA1E79BA3B98CCA73">Senckenberg Dresden, Koenigsbruecker Landstrasse 159, 01109 Dresden, Germany & Department of Life Sciences, The Natural History Museum, London SW 7 5 BD, UK & University of Wolverhampton, Wulfruna St, Wolverhampton WV 1 1 LY, UK</mods:affiliation>
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<mods:nameIdentifier id="07505059BDD579360EA3C7FBA550182D" type="email">veerappandeepak@gmail.com</mods:nameIdentifier>
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<mods:typeOfResource id="77B5324BF5B8BBD1E636D1322CD948B9">text</mods:typeOfResource>
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<mods:title id="808D80C34D81F44669EAB582BBF7854B">Vertebrate Zoology</mods:title>
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<mods:part id="8242E3A326B13F79AD2F66A59666F040">
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<mods:date id="358CC28933BB059B958F75D9DA678577">2023</mods:date>
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<mods:number id="9F04A744428B50DB46CE7065A0C7B12E">2023-05-23</mods:number>
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<mods:number id="E8D8ADD614CD14FE7A4CD8D4C8BBA8F5">73</mods:number>
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<mods:url id="1D95BC9ACD3DCB1C55F1A368438B2F00">http://dx.doi.org/10.3897/vz.73.e104494</mods:url>
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<mods:classification id="FA8C7414751292C161B4F5AB42C8F7C6">journal article</mods:classification>
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<mods:identifier id="74596AF410891AE3C2CF8F1E9637FF36" type="DOI">http://dx.doi.org/10.3897/vz.73.e104494</mods:identifier>
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<treatment id="3F9574FC242D515BA53EFDD1EC3EDF85" ID-GBIF-Taxon="209629479" LSID="urn:lsid:zoobank.org:act:820BD788-BD9D-491B-BBBB-0E1C0649ED6E" httpUri="http://treatment.plazi.org/id/3F9574FC242D515BA53EFDD1EC3EDF85" lastPageNumber="499" pageId="0" pageNumber="499" scope_class="Squamata" scope_family="Gekkonidae">
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<subSubSection id="21EFD9B8C49A2264B4E28D7B81E7784B" pageId="0" pageNumber="499" type="nomenclature">
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<paragraph id="A73EA0667EF5C5F60578287356287938" pageId="0" pageNumber="499">
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<taxonomicName id="4BE98E7E1FF08E08BB9BA4B0286B2EA4" LSID="https://zoobank.org/820BD788-BD9D-491B-BBBB-0E1C0649ED6E" authority="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis" status="sp. nov.">Hemidactylus pakkamalaiensis Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak</taxonomicName>
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<taxonomicNameLabel id="4C7E904C3C26081C84D7EA80D66900B5" pageId="0" pageNumber="499">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="99F73034B895CAB842FB082E10D5CC6E" pageId="0" pageNumber="499" type="description">
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<paragraph id="A945A96CC7A015435A5EB8C4BF696281" pageId="0" pageNumber="499">
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<figureCitation id="D5B2C16886AB5F360BA656CFA7CF4374" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Hemidactylus pakkamalaiensis sp. nov. (BNHS 2907) in the dorsal (A) and ventral (B) view. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure3" httpUri="https://binary.pensoft.net/fig/853723" pageId="0" pageNumber="499">Figs 3</figureCitation>
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<figureCitation id="0FC8F935F8C960478D24E0D3D1BF36E5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">, 4</figureCitation>
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<figureCitation id="E7F07DFBBB2CA998F5B678CC31EFDDA2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paratypes of Hemidactylus pakkamalaiensis sp. nov. in dorsal view, A BNHS 2908, B BNHS 2909, C BNHS 2910, D BNHS 2911. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure5" httpUri="https://binary.pensoft.net/fig/853725" pageId="0" pageNumber="499">, 5</figureCitation>
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<figureCitation id="139AB97073F4373E0ADB110BB94AACD4" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsolateral view of the A holotype (BNHS 2907) and B one of the paratypes (BNHS 2911) of Hemidactylus pakkamalaiensis sp. nov. in life from Pakkamalai Hill, Tamil Nadu, India." figureDoi="10.3897/vz.73.e104494.figure6" httpUri="https://binary.pensoft.net/fig/853726" pageId="0" pageNumber="499">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B85425A96EFEFB29538D4557339EFD9B" pageId="0" pageNumber="499" type="holotype">
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<paragraph id="B1EA7C6D6754843C32132CF1AD1C3B1C" pageId="0" pageNumber="499">Holotype.</paragraph>
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<paragraph id="97F9D7A5362E6BDC0A326648B64CC17D" pageId="0" pageNumber="499">
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BNHS 2907 (Figs
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<figureCitation id="5AE620E16DCF801998CE510F6ED66B6F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Holotype of Hemidactylus pakkamalaiensis sp. nov. (BNHS 2907) in the dorsal (A) and ventral (B) view. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure3" httpUri="https://binary.pensoft.net/fig/853723" pageId="0" pageNumber="499">3</figureCitation>
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,
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<figureCitation id="9C5F2555ACC189C2725A969A29B80952" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">4</figureCitation>
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,
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<figureCitation id="7BF342C3C4255F05359073AB6FD9A229" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsolateral view of the A holotype (BNHS 2907) and B one of the paratypes (BNHS 2911) of Hemidactylus pakkamalaiensis sp. nov. in life from Pakkamalai Hill, Tamil Nadu, India." figureDoi="10.3897/vz.73.e104494.figure6" httpUri="https://binary.pensoft.net/fig/853726" pageId="0" pageNumber="499">6A</figureCitation>
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) adult male, collected from Pakkamalai Hill, Gingee Hills range, Villupuram district, Tamil Nadu, India (
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<geoCoordinate id="A5EFAF3C02FFD6887DBA1DCC0E271533" degrees="12.164819" direction="north" orientation="latitude" precision="1" value="12.164819">12.164819°N</geoCoordinate>
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,
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<geoCoordinate id="91044AB0371139102B294CC437920474" degrees="79.250909" direction="east" orientation="longitude" precision="1" value="79.25091">79.250909°E</geoCoordinate>
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, 335m asl) by Peter Christopher and Surya Narayanan on 27 November 2022.
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</paragraph>
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<caption id="5A57B67F37C526A84DF13D778B6F1AAD" doi="10.3897/vz.73.e104494.figure3" httpUri="https://binary.pensoft.net/fig/853723" pageId="0" pageNumber="499" start="Figure 3" startId="F3">
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<paragraph id="CA3C1C1A4FD27F6DDD0FE4056DFCDD3B" pageId="0" pageNumber="499">
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<emphasis id="873D717E598F6DC5D6315C94DA194127" bold="true" pageId="0" pageNumber="499">Figure 3.</emphasis>
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Holotype of
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<taxonomicName id="4857DF4A2C7B9A139D464D2649882111" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
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<emphasis id="35A1982003AF937C9E966BE09DCAC387" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
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</taxonomicName>
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<emphasis id="CB749825933DCFF1E681DE66B67BFAAA" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
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(BNHS 2907) in the dorsal (
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<emphasis id="9A76088829A324EF77DB8D057B2FC09B" bold="true" pageId="0" pageNumber="499">A</emphasis>
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) and ventral (
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<emphasis id="DB74FB2C50AFF6EB48F723977462CBF3" bold="true" pageId="0" pageNumber="499">B</emphasis>
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) view. Scale bar = 10mm.
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</paragraph>
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</caption>
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<caption id="332A6B9F2A6A68923930574508EB1717" doi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499" start="Figure 4" startId="F4">
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<paragraph id="9C289837A358F554B5E59588882C576D" pageId="0" pageNumber="499">
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<emphasis id="B793F81B93F87F50F8497479ED31D651" bold="true" pageId="0" pageNumber="499">Figure 4.</emphasis>
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<taxonomicName id="B6CB7815CE3886930341C13A5F299462" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
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<emphasis id="79591D0EBD559757EDCB6D38BD47B90A" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
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</taxonomicName>
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<emphasis id="D68AE93CC1408C6524588A8345E1C18F" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
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(Holotype, BNHS 2907),
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<emphasis id="FD13BAAC37B8BB5F74F90640293F8A96" bold="true" pageId="0" pageNumber="499">A</emphasis>
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head in dorsal view,
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<emphasis id="839D42648A2DFA444F879986B86B31D6" bold="true" pageId="0" pageNumber="499">B</emphasis>
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ventral view,
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<emphasis id="217D8308F93BE4F3AAE12B4776CEA8F6" bold="true" pageId="0" pageNumber="499">C</emphasis>
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lateral view,
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<emphasis id="84F18C3FE8ED6C19EA8BCEF5C0D0F0F8" bold="true" pageId="0" pageNumber="499">D</emphasis>
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view of the femoral region,
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<emphasis id="BFD5A2505712EB221AABD82A29420C76" bold="true" pageId="0" pageNumber="499">E</emphasis>
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ventral view of right manus and
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<emphasis id="08C8946100EA380624ECA72B6276B39E" bold="true" pageId="0" pageNumber="499">F</emphasis>
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right pes. Scale bar = 10mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="4FA1BA7BBD5AB6003A3DEDDC0A570A9E" pageId="0" pageNumber="499" type="paratypes (n = 4)">
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<paragraph id="507A6B89373DBD8C5DD5B80D7B373878" pageId="0" pageNumber="499">Paratypes (n = 4).</paragraph>
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<paragraph id="44AFBAE9617FDD477608355D9527C186" pageId="0" pageNumber="499">
|
|
BNHS 2908 & BNHS 2909 (adult males), BNHS 2910 & BNHS 2911 (adult females), collections details same as the holotype (Figs
|
|
<figureCitation id="0D97F44EB515908A87B71E064DF8F612" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Paratypes of Hemidactylus pakkamalaiensis sp. nov. in dorsal view, A BNHS 2908, B BNHS 2909, C BNHS 2910, D BNHS 2911. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure5" httpUri="https://binary.pensoft.net/fig/853725" pageId="0" pageNumber="499">5</figureCitation>
|
|
,
|
|
<figureCitation id="B8D69C0A870F09D9C9788F4EB8589AD6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsolateral view of the A holotype (BNHS 2907) and B one of the paratypes (BNHS 2911) of Hemidactylus pakkamalaiensis sp. nov. in life from Pakkamalai Hill, Tamil Nadu, India." figureDoi="10.3897/vz.73.e104494.figure6" httpUri="https://binary.pensoft.net/fig/853726" pageId="0" pageNumber="499">6B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="4DBB8CBAE63902198BA17C81B026D2DD" doi="10.3897/vz.73.e104494.figure5" httpUri="https://binary.pensoft.net/fig/853725" pageId="0" pageNumber="499" start="Figure 5" startId="F5">
|
|
<paragraph id="8A8AB0FD8B3D98860BBF677C1C9E2E42" pageId="0" pageNumber="499">
|
|
<emphasis id="87C405AC07C9E158A60183BA726FF426" bold="true" pageId="0" pageNumber="499">Figure 5.</emphasis>
|
|
Paratypes of
|
|
<taxonomicName id="F66B637D0EE61A17F3787A48FFD2F1BB" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
|
|
<emphasis id="4FA36645E4E7BE170935247BB94D30BB" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="40791CA69E5531D8761685035D038114" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
|
|
in dorsal view,
|
|
<emphasis id="D6C48C712EDE2E30B86CBCFBA4C8B4B9" bold="true" pageId="0" pageNumber="499">A</emphasis>
|
|
BNHS 2908,
|
|
<emphasis id="585B56A7A49EF94E222D8FF86E7F2471" bold="true" pageId="0" pageNumber="499">B</emphasis>
|
|
BNHS 2909,
|
|
<emphasis id="BF95A5735DD81B56B8A61DF4D80DF0B4" bold="true" pageId="0" pageNumber="499">C</emphasis>
|
|
BNHS 2910,
|
|
<emphasis id="4BDDDADFA6335314E40A492F6346674A" bold="true" pageId="0" pageNumber="499">D</emphasis>
|
|
BNHS 2911. Scale bar = 10mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="9FC90C3978DC5E3D40165196E3D514FF" doi="10.3897/vz.73.e104494.figure6" httpUri="https://binary.pensoft.net/fig/853726" pageId="0" pageNumber="499" start="Figure 6" startId="F6">
|
|
<paragraph id="9C08EA519B3C2E3E85D33CCABB8169B9" pageId="0" pageNumber="499">
|
|
<emphasis id="3FFAF6B101CF3A48D6941E70802A0EFA" bold="true" pageId="0" pageNumber="499">Figure 6.</emphasis>
|
|
Dorsolateral view of the
|
|
<emphasis id="21A09523764560FE1AA504DA4D7B3361" bold="true" pageId="0" pageNumber="499">A</emphasis>
|
|
holotype (BNHS 2907) and
|
|
<emphasis id="B3E5B033D147D4C6234335C147E81720" bold="true" pageId="0" pageNumber="499">B</emphasis>
|
|
one of the paratypes (BNHS 2911) of
|
|
<taxonomicName id="7A239E1EB90E443C4C4C220696F16EEB" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
|
|
<emphasis id="CD7F1685D3973FA1AACBFD926F83FE68" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="E86A0C7745322DD9049B6C0AEA181BC2" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
|
|
in life from Pakkamalai Hill, Tamil Nadu, India.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
<subSubSection id="77C318485DA22707F0B4E401C32708BC" pageId="0" pageNumber="499" type="etymology">
|
|
<paragraph id="C21F85D3FCAC56D4E32FA8815427FB3A" pageId="0" pageNumber="499">Etymology.</paragraph>
|
|
<paragraph id="48165F8B8B0CED5ACCD70D84FB584E70" pageId="0" pageNumber="499">
|
|
The specific epithet is a toponym named after its type locality Pakkamalai Hill. We suggest an English name Pakkamalai rock gecko and a Tamil name
|
|
<normalizedToken id="A5ADC71785A4FDC7B558383A7836FDFC" originalValue="ப">ப</normalizedToken>
|
|
ாக்கமைல
|
|
<normalizedToken id="B1A142887914CFE237107A93C99BE8C0" originalValue="ப">ப</normalizedToken>
|
|
ாைறப்பல்லி.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="F71DE3183D22D03B7CA616DD8332DD93" pageId="0" pageNumber="499" type="diagnosis">
|
|
<paragraph id="1ECA9613EE928E75E8DE30CDEE939275" pageId="0" pageNumber="499">Diagnosis.</paragraph>
|
|
<paragraph id="DBAF97BF3EAA0E63334F0A51B826E1E1" pageId="0" pageNumber="499">
|
|
A large-sized gecko of the genus
|
|
<taxonomicName id="4E845B5244EBD0F016AB5A0F6AA57943" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus" order="Squamata" pageId="0" pageNumber="499" rank="genus">
|
|
<emphasis id="AB03680B6122F046DD23C4106EC73B9A" italics="true" pageId="0" pageNumber="499">Hemidactylus</emphasis>
|
|
</taxonomicName>
|
|
, snout-vent length up to a maximum SVL of 109 mm (n = 5). Dorsal pholidosis heterogeneous, composed of roughly circular, granular scales intermixed with much enlarged, fairly regularly arranged longitudinal rows of 19-22 striated subtrihedral tubercles at midbody. Enlarged tubercles on the two most medial parasagittal rows are small, subconical, strongly keeled and rounded, gradually increasing in size and becoming conical towards flanks, last two to three rows on flanks smaller, conical. Two well-developed pairs of postmentals, the inner pair longer than the outer pair and mental, and in broad contact behind the mental. Paravertebral rows with 21-24 tubercles; 34-38 transverse ventral scale rows at mid-body. Digits with enlarged scansors, lamellae in straight transverse series; two or three undivided basal lamellae beneath first finger and two or three beneath first toe; one or two undivided basal lamellae beneath fourth toe; ten or 11 lamellae (including undivided and divided) beneath first finger and nine or ten beneath first toe; 12 or 13 lamellae (including undivided and divided) beneath fourth finger and 12-14 beneath fourth toe; males with series of 19-21 femoral pores on each side separated by four or five pore-less scales. Original tail depressed, oval in transverse section with a median dorsal furrow; scales on the tail slightly larger than dorsals, striated, with longitudinal series of 4-8 large, keeled, striated, posteriorly pointed tubercles. Dorsal colouration dull-brown with a series of four or five transverse pale saddles from occiput to sacrum, tail with distinct alternating light and dark bands.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="1AA8D84DF630C99B781D7820016A4329" pageId="0" pageNumber="499" type="comparisons">
|
|
<paragraph id="B3A3F0B27821F5A045E98A99E775111A" pageId="0" pageNumber="499">Comparison with other members of prashadi group.</paragraph>
|
|
<paragraph id="21961CBCE423BDCD43DC3134B297BDA2" pageId="0" pageNumber="499">
|
|
<taxonomicName id="563AA186F72D8009B7243A624CE0F6CF" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
|
|
<emphasis id="E099E0F66AFAB3DF80FEE51B1679843E" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="C3EB1BD21DA11E32D7938D0AB8F9710F" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
|
|
differs from the other large-bodied congeners by several non-overlapping morphological characters. It can be distinguished by the presence of 19-21 femoral pores (FP) separated by 4 or 5 pore-less scales in males (vs. 23-28 FP separated by 1-3 pore-less scales in
|
|
<taxonomicName id="6F3D6E0DE0182B2B319F733BD3E06A40" lsidName="H. graniticolus" pageId="0" pageNumber="499" rank="species" species="graniticolus">
|
|
<emphasis id="8D0998B15E89E18FF87CF79BCFDF8607" italics="true" pageId="0" pageNumber="499">H. graniticolus</emphasis>
|
|
</taxonomicName>
|
|
; 19-21 FP separated by 13 or 14 pore-less scales in
|
|
<taxonomicName id="9A957A32E14ADAFEF07AE0561CB42750" lsidName="H. acanthopholis" pageId="0" pageNumber="499" rank="species" species="acanthopholis">
|
|
<emphasis id="4761699CCCA00BBA5A1D9A4BF8C49C78" italics="true" pageId="0" pageNumber="499">H. acanthopholis</emphasis>
|
|
</taxonomicName>
|
|
Mirza & Sanap, 2014; 26-28 FP separated by 7-9 pore-less scales in
|
|
<taxonomicName id="EA3C77729E606EB387C8C2C7D3BA90FD" lsidName="H. hunae" pageId="0" pageNumber="499" rank="species" species="hunae">
|
|
<emphasis id="FA6AEEE508CAF422343473D3889C7E38" italics="true" pageId="0" pageNumber="499">H. hunae</emphasis>
|
|
</taxonomicName>
|
|
Deraniyagala, 1937; 16-18 FP separated by 13 or 14 pore-less scales in
|
|
<taxonomicName id="EF1E3E96CF7D25A6F11CCA84DBE5DADA" lsidName="H. sirumalaiensis" pageId="0" pageNumber="499" rank="species" species="sirumalaiensis">
|
|
<emphasis id="EC5547448DD707BFCE6E1C6225BE02C2" italics="true" pageId="0" pageNumber="499">H. sirumalaiensis</emphasis>
|
|
</taxonomicName>
|
|
<bibRefCitation id="199257D67F977F4426508FE1E77294F0" DOI="https://doi.org/10.11646/zootaxa.4852.1.4" author="Khandekar, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="499" pagination="83 - 100" refId="B19" refString="Khandekar, A, Thackeray, T, Pawar, S, Agarwal, I, 2020. A new medium-bodied rupicolous Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from the Sirumalai massif, Tamil Nadu, India. Zootaxa 4852: 83 - 100, DOI: https://doi.org/10.11646/zootaxa.4852.1.4" title="A new medium-bodied rupicolous Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) from the Sirumalai massif, Tamil Nadu, India." url="https://doi.org/10.11646/zootaxa.4852.1.4" volume="4852" year="2020">Khandekar et al., 2020</bibRefCitation>
|
|
; 17-22 FP separated by 10 or 11 pore-less scales in
|
|
<taxonomicName id="69986DC2318C3C91A407FB2722439746" lsidName="H. vanam" pageId="0" pageNumber="499" rank="species" species="vanam">
|
|
<emphasis id="68D86366CA89674D171C9EFE31E8FF9F" italics="true" pageId="0" pageNumber="499">H. vanam</emphasis>
|
|
</taxonomicName>
|
|
<bibRefCitation id="802C8333EFF55DD305104C4723182B68" DOI="doi.org/10.11646/zootaxa.4374.1.3" author="Chaitanya, R" journalOrPublisher="Zootaxa" pageId="0" pageNumber="499" pagination="49 - 70" refId="B9" refString="Chaitanya, R, Lajmi, A, Giri, VB, 2018. A new cryptic, rupicolous species of Hemidactylus Oken, 1817 (Squamata: Gekkonidae) from Meghamalai, Tamil Nadu, India. Zootaxa 4374: 49 - 70, DOI: doi.org/10.11646/zootaxa.4374.1.3" title="A new cryptic, rupicolous species of Hemidactylus Oken, 1817 (Squamata: Gekkonidae) from Meghamalai, Tamil Nadu, India." url="doi.org/10.11646/zootaxa.4374.1.3" volume="4374" year="2018">Chaitanya et al., 2018</bibRefCitation>
|
|
; 15-19 FP separated by three pore-less scales in
|
|
<taxonomicName id="A12F776FF9F06F7D03EDD6AD7BEE7D83" lsidName="H. depressus" pageId="0" pageNumber="499" rank="species" species="depressus">
|
|
<emphasis id="3AE764C62EF46F5BACF6B4A9AB933D08" italics="true" pageId="0" pageNumber="499">H. depressus</emphasis>
|
|
</taxonomicName>
|
|
Gray, 1842; 17 or 18 FP separated by five pore-less scales in
|
|
<taxonomicName id="005D48609F2C193C3EF9984247E2DAFC" lsidName="H. siva" pageId="0" pageNumber="499" rank="species" species="siva">
|
|
<emphasis id="7E8DE2D17606396F43433ACE746F1ED3" italics="true" pageId="0" pageNumber="499">H. siva</emphasis>
|
|
</taxonomicName>
|
|
Srinivasalu et al., 2018; 7-9 FP separated by 1-3 pore-less scales in
|
|
<taxonomicName id="06D7B48648666D295BD66E396F60CA3E" lsidName="H. triedrus" pageId="0" pageNumber="499" rank="species" species="triedrus">
|
|
<emphasis id="DD2A6215FE581906431581E380F2C258" italics="true" pageId="0" pageNumber="499">H. triedrus</emphasis>
|
|
</taxonomicName>
|
|
(Daudin, 1802); 11-15 FP separated by 1-3 pore-less scales in
|
|
<taxonomicName id="5112BFBFF05172928D1BB640A6182DF4" lsidName="H. sahgali" pageId="0" pageNumber="499" rank="species" species="sahgali">
|
|
<emphasis id="71607A58F4A6DB8A553A5EEACBDE666B" italics="true" pageId="0" pageNumber="499">H. sahgali</emphasis>
|
|
</taxonomicName>
|
|
<bibRefCitation id="22806AB9E1D6056D02FAA4039BC1CE84" DOI="https://doi.org/10.7717/peerj.5341" author="Mirza, ZA" journalOrPublisher="Taprobanica" pageId="0" pageNumber="499" refId="B27" refString="Mirza, ZA, Gowande, GG, Patil, R, Ambekar, M, Patel, H, 2018. First appearance deceives many: Disentangling the Hemidactylus triedrus species complex using an integrated approach. PeerJ 6: e5341. https://doi.org/10.7717/peerj.5341" title="First appearance deceives many: Disentangling the Hemidactylus triedrus species complex using an integrated approach. PeerJ 6: e 5341." url="https://doi.org/10.7717/peerj.5341" year="2018">Mirza et al., 2018</bibRefCitation>
|
|
; 7 or 8 FP separated by three pore-less scales in
|
|
<taxonomicName id="5CD6921240E6DA989F93A5D8F0D40A7D" lsidName="H. whitakeri" pageId="0" pageNumber="499" rank="species" species="whitakeri">
|
|
<emphasis id="FC0491BFFCF2BF0DB27684C5F4B5EA0A" italics="true" pageId="0" pageNumber="499">H. whitakeri</emphasis>
|
|
</taxonomicName>
|
|
<bibRefCitation id="451CFC1B4DE1D494C5BA41A42941C970" DOI="https://doi.org/10.7717/peerj.5341" author="Mirza, ZA" journalOrPublisher="Taprobanica" pageId="0" pageNumber="499" refId="B27" refString="Mirza, ZA, Gowande, GG, Patil, R, Ambekar, M, Patel, H, 2018. First appearance deceives many: Disentangling the Hemidactylus triedrus species complex using an integrated approach. PeerJ 6: e5341. https://doi.org/10.7717/peerj.5341" title="First appearance deceives many: Disentangling the Hemidactylus triedrus species complex using an integrated approach. PeerJ 6: e 5341." url="https://doi.org/10.7717/peerj.5341" year="2018">Mirza et al., 2018</bibRefCitation>
|
|
; 15-19 FP separated by six pore-less scales in
|
|
<taxonomicName id="BA18B989DEFE9ED2CAB19DA043E71036" lsidName="H. aaronbaueri" pageId="0" pageNumber="499" rank="species" species="aaronbaueri">
|
|
<emphasis id="62D76ADE6C08F278E917B96B35ED091D" italics="true" pageId="0" pageNumber="499">H. aaronbaueri</emphasis>
|
|
</taxonomicName>
|
|
<bibRefCitation id="E1C99F9454ECB66D4EF4ACAEA97D9510" author="Giri, VB" journalOrPublisher="Hamadryad" pageId="0" pageNumber="499" pagination="25 - 33" refId="B14" refString="Giri, VB, 2008. A new rock dwelling Hemidactylus (Squamata: Gekkonidae) from Maharashtra, India. Hamadryad 32: 25 - 33" title="A new rock dwelling Hemidactylus (Squamata: Gekkonidae) from Maharashtra, India." volume="32" year="2008">Giri, 2008</bibRefCitation>
|
|
; 17 or 18 FP separated by seven pore-less scales in
|
|
<taxonomicName id="7780CBC4E1232662B1FEEE82936943DB" lsidName="H. tamhiniensis" pageId="0" pageNumber="499" rank="species" species="tamhiniensis">
|
|
<emphasis id="952FEF2CA3253F859627F32281435D85" italics="true" pageId="0" pageNumber="499">H. tamhiniensis</emphasis>
|
|
</taxonomicName>
|
|
<bibRefCitation id="0C74F443A54151EF9172B6DB0F35818B" DOI="https://doi.org/10.11646/zootaxa.5020.3.2" author="Khandekar, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="499" pagination="434 - 456" refId="B20" refString="Khandekar, A, Thackeray, T, Agarwal, I, 2021. A cryptic new species of rupicolous Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) allied to H. aaronbaueri Giri, 2008 from the northern Western Ghats of Maharashtra, India. Zootaxa 5020: 434 - 456, DOI: https://doi.org/10.11646/zootaxa.5020.3.2" title="A cryptic new species of rupicolous Hemidactylus Goldfuss, 1820 (Squamata: Gekkonidae) allied to H. aaronbaueri Giri, 2008 from the northern Western Ghats of Maharashtra, India." url="https://doi.org/10.11646/zootaxa.5020.3.2" volume="5020" year="2021">Khandekar et al., 2021</bibRefCitation>
|
|
; 18-21 FP separated by four pore-less scales in
|
|
<taxonomicName id="DD6F5C15DD7DBD232C14060FBB559618" lsidName="H. kangerensis" pageId="0" pageNumber="499" rank="species" species="kangerensis">
|
|
<emphasis id="419B55A3833531EF22230A0E402EA4C5" italics="true" pageId="0" pageNumber="499">H. kangerensis</emphasis>
|
|
</taxonomicName>
|
|
Mirza et al., 2017; 24-30 FP separated by 2-4 pore-less scales in
|
|
<taxonomicName id="A4D8C88B6617F7E765D4D271D4A17C84" lsidName="H. easai" pageId="0" pageNumber="499" rank="species" species="easai">
|
|
<emphasis id="D7E8E63CBA45258EDF178D9A7D34100A" italics="true" pageId="0" pageNumber="499">H. easai</emphasis>
|
|
</taxonomicName>
|
|
; 14-15 FP separated by 11 pore-less scales in
|
|
<taxonomicName id="88A23AC0999AFAF4C66BD0F41DE30C5B" lsidName="H. hegdei" pageId="0" pageNumber="499" rank="species" species="hegdei">
|
|
<emphasis id="9677CE656353ECD20D32E6B0CB942DDF" italics="true" pageId="0" pageNumber="499">H. hegdei</emphasis>
|
|
</taxonomicName>
|
|
Pal & Mirza, 2022); by the presence of 19-22 rows of moderately keeled subtrihedral tubercles in both sex (vs. 16 or 17 in
|
|
<taxonomicName id="DFE3565C547922B88F128FE6B55E6AD2" lsidName="H. sushilduttai" pageId="0" pageNumber="499" rank="species" species="sushilduttai">
|
|
<emphasis id="3CE7B6D54497CD531803F2DF881D4771" italics="true" pageId="0" pageNumber="499">H. sushilduttai</emphasis>
|
|
</taxonomicName>
|
|
Giri et al., 2017; 12-14 in
|
|
<taxonomicName id="1DC37F2A4C8996BB72AC65A84D5DF830" lsidName="H. kimbulae" pageId="0" pageNumber="499" rank="species" species="kimbulae">
|
|
<emphasis id="511891F27C8176804E220427087B3A5D" italics="true" pageId="0" pageNumber="499">H. kimbulae</emphasis>
|
|
</taxonomicName>
|
|
; 15 or 16 in
|
|
<taxonomicName id="7973C72667404FB68ACFF556649B5D8F" lsidName="H. kolliensis" pageId="0" pageNumber="499" rank="species" species="kolliensis">
|
|
<emphasis id="C2DB3A9499BB067FCE00625DF156B127" italics="true" pageId="0" pageNumber="499">H. kolliensis</emphasis>
|
|
</taxonomicName>
|
|
; 17 or 18 in
|
|
<taxonomicName id="771EA98658FB32B23DDF506F57A1FF93" lsidName="H. easai" pageId="0" pageNumber="499" rank="species" species="easai">
|
|
<emphasis id="50A83FD3DC2D840443BFC99B457220BA" italics="true" pageId="0" pageNumber="499">H. easai</emphasis>
|
|
</taxonomicName>
|
|
; 18-20 in
|
|
<taxonomicName id="D1D812C40D3CC11C96193586CB295623" lsidName="H. kangerensis" pageId="0" pageNumber="499" rank="species" species="kangerensis">
|
|
<emphasis id="2D4C5DD21EFB4D2AF7FE65A8ADD80ADD" italics="true" pageId="0" pageNumber="499">H. kangerensis</emphasis>
|
|
</taxonomicName>
|
|
; 14-16 in
|
|
<taxonomicName id="469535A957F1DDE589A8EBE1EF81A927" lsidName="H. hunae" pageId="0" pageNumber="499" rank="species" species="hunae">
|
|
<emphasis id="669E2387EB8AFB52AFABCA607AF005AF" italics="true" pageId="0" pageNumber="499">H. hunae</emphasis>
|
|
</taxonomicName>
|
|
; 16 in
|
|
<taxonomicName id="C016AD556FAF2797B3B53C2F1CEBB23C" lsidName="H. siva" pageId="0" pageNumber="499" rank="species" species="siva">
|
|
<emphasis id="8D30458435833BF6DCDA6921DEA90821" italics="true" pageId="0" pageNumber="499">H. siva</emphasis>
|
|
</taxonomicName>
|
|
; 16-18 in
|
|
<taxonomicName id="63DD6C48B0FA6772FE7E06284902A373" lsidName="H. sushilduttai" pageId="0" pageNumber="499" rank="species" species="sushilduttai">
|
|
<emphasis id="1CFE1DC43FD21E88A0930334DF977B71" italics="true" pageId="0" pageNumber="499">H. sushilduttai</emphasis>
|
|
</taxonomicName>
|
|
); presence of 34-38 transverse ventral scale rows at mid-body in both sex (vs. 40-13 in
|
|
<taxonomicName id="0022F8722A869584BE61F33DD517130B" lsidName="H. hunae" pageId="0" pageNumber="499" rank="species" species="hunae">
|
|
<emphasis id="2727D8722AAA574B1DA69935851891A1" italics="true" pageId="0" pageNumber="499">H. hunae</emphasis>
|
|
</taxonomicName>
|
|
, 41 or 42 in
|
|
<taxonomicName id="5AC174367275F1C6BDF5C64035994C26" lsidName="H. easai" pageId="0" pageNumber="499" rank="species" species="easai">
|
|
<emphasis id="BC98CDB7AA85F72C9C795780BC7460F4" italics="true" pageId="0" pageNumber="499">H. easai</emphasis>
|
|
</taxonomicName>
|
|
; 40-46 in
|
|
<taxonomicName id="E12BF477449EB232BB14C80FF2738488" lsidName="H. graniticolus" pageId="0" pageNumber="499" rank="species" species="graniticolus">
|
|
<emphasis id="99C1887F682A189DFFDFFF9A77004466" italics="true" pageId="0" pageNumber="499">H. graniticolus</emphasis>
|
|
</taxonomicName>
|
|
; 28-30 in
|
|
<taxonomicName id="395DA3A369CCC5E1D4F6D13C80707751" lsidName="H. kangerensis" pageId="0" pageNumber="499" rank="species" species="kangerensis">
|
|
<emphasis id="9347FA4689AA016F0AEA4E9A813F0127" italics="true" pageId="0" pageNumber="499">H. kangerensis</emphasis>
|
|
</taxonomicName>
|
|
; 30-33 in
|
|
<taxonomicName id="33CE31BD68012D226B3B70B8739AE4B1" lsidName="H. sushilduttai" pageId="0" pageNumber="499" rank="species" species="sushilduttai">
|
|
<emphasis id="43B65BE3E46539A9F7B7E09393043116" italics="true" pageId="0" pageNumber="499">H. sushilduttai</emphasis>
|
|
</taxonomicName>
|
|
; 27-30 in
|
|
<taxonomicName id="9E6311637BB96694EE04AF7F8AA076AB" lsidName="H. siva" pageId="0" pageNumber="499" rank="species" species="siva">
|
|
<emphasis id="7A333B4D580C1769DF48110118967238" italics="true" pageId="0" pageNumber="499">H. siva</emphasis>
|
|
</taxonomicName>
|
|
); dorsal pholidosis heterogeneous with enlarged tubercles (vs. dorsal pholidosis homogeneous with no enlarged tubercles in
|
|
<taxonomicName id="786101DD5A2971441EFEF13E13378AC8" lsidName="H. scabriceps" pageId="0" pageNumber="499" rank="species" species="scabriceps">
|
|
<emphasis id="EA2D366DD2A33FB5724E582F34D2DB35" italics="true" pageId="0" pageNumber="499">H. scabriceps</emphasis>
|
|
</taxonomicName>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9C11E2DF7D809C5D8302A5A27C4834D1" pageId="0" pageNumber="499" type="description">
|
|
<paragraph id="075C66253C84E3C2437A7200277860D3" pageId="0" pageNumber="499">Description of holotype (BNHS 2907).</paragraph>
|
|
<paragraph id="1FEFA21658D05168D8B9970CE897F8B1" pageId="0" pageNumber="499">
|
|
The holotype, an adult male, (SVL 104 mm) is well preserved. Head short (HL/SVL 0.29), slightly elongated (HW/HL 0.66), not strongly depressed (HH/HL 0.29), distinct from neck. Loreal region slightly inflated, and canthus rostralis indistinct (Fig.
|
|
<figureCitation id="B7EAB5E1D3ADDCA5D6703A173BA50E78" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">4C</figureCitation>
|
|
). Snout short (SE/HL 0.40); slightly longer than twice eye diameter (OD/SE 0.45); scales on the snout, canthus rostralis, forehead and inter-orbital region heterogeneous, mostly granular and conical; scales on the snout and canthus rostralis much larger than those on occiput, forehead and inter-orbital regions; skin in the frontal slightly damaged. Eye small (OD/HL 0.18); pupil vertical with crenulated margins; supraciliaris small, mucronate, gradually increasing in size towards the front of the orbit, largest about one-third the way from the anterior edge of orbit; a few supraciliaris in the suprocular region in the left eye damaged and missing. Ear opening elliptical (2.8 mm); a row of small keeled tubercles above the ear.
|
|
</paragraph>
|
|
<paragraph id="9EF962C4935146D0DFF33785E9637866" pageId="0" pageNumber="499">Rostral scale wider than deep (RL/RW 0.68), partially divided dorsally by a weakly developed rostral groove; nasals, enlarged and separated by three small scales; one supranasal on each side, smaller than internasals; two diminutive postnasals on each side; rostral in contact with nasal scale, supralabial I, internasals and the anterior small scale separating the internasals; nostrils small (0.8 mm), oval; nasal scale surrounded by supranasal, internasal, rostral, supralabial I and three postnasals on either side; five (on the anterior) and three (on the posterior) rows of scales separate orbit from supralabials.</paragraph>
|
|
<paragraph id="4D2CE8F0B43C983CC77CE0D944492E93" pageId="0" pageNumber="499">
|
|
Mental triangular; two well-developed pair of postmentals, the inner pair smaller (3.4 mm) than the mental (4.6 mm), and in strong contact with each other (1.2 mm) behind mental, outer pair shorter (2.2 mm) than the inner pair and separated from each other by inner pair (Fig.
|
|
<figureCitation id="BD90210BB284700EA4BE3615C1E898E2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">4B</figureCitation>
|
|
). Inner postmentals bordered by mental, infralabial I and II on left and right, outer postmental and 10 small gular scales on each side; outer postmentals bordered by infralabial II on the right and left, inner postmental, and five gular scales on the right and six gular scales in the left along with two other large scales between the gular scales and the infralabials; one to three additional rows of scales below infralabials III to VIII are enlarged and weakly imbricate. Supralabials (on both sides) to mid orbital position 10, to angle of jaw 12 on right and left; infralabials 10 and nine on right and left side, respectively. Body relatively stout (BW/SVL 0.22), ventrolateral folds not very distinct.
|
|
</paragraph>
|
|
<paragraph id="4FAB5E8BA293952CBB146161DE57873D" pageId="0" pageNumber="499">Dorsal pholidosis heterogeneous, composed of subcircular granular scales intermixed with enlarged, fairly regularly arranged strongly keeled. Pointed tubercles in 21 longitudinal rows, extending from occiput to tail, that are heterogeneous in shape and size; enlarged tubercles on the two most medial parasagittal rows slightly smaller than rest on dorsum and the rows most broadly spaced from one another, gradually increasing in size and becoming conical towards flanks, last two rows on flanks slightly smaller than medial parasagittal rows and strongly conical; each enlarged tubercle surrounded by a rosette of 13-16 small granules with 2-5 granules between two longitudinally adjacent enlarged tubercles (5-11 between parasagittal rows at midbody); enlarged tubercles on nape and shoulder smaller and conical, those on occiput and the temporal region still smaller, conical.</paragraph>
|
|
<paragraph id="7B6126865F1DE7F7FFD96F128988135B" pageId="0" pageNumber="499">Ventral scales larger than granular scales on the dorsum, smooth, imbricate, slightly larger on precloacal and femoral region than on chest and abdominal region; midbody scale rows across belly 34; gular region with small, granular scales, becoming slightly larger and imbricate on anterior and lateral aspect. Scales on palm and sole smooth, imbricate, subcircular; scales on the dorsal aspect of upper arm slightly smaller than the last row of enlarged tubercles on dorsum, flat, weakly pointed, imbricate, keeled; dorsal aspect of forearm with smaller, granular scales, intermixed with a few enlarged, conical tubercles, those on anterior aspect are smooth, flat, imbricate; scales on dorsal part of thigh and shank granular, imbricate, intermixed with enlarged, conical tubercles, which are larger on thigh compared to shank that are subequal to those tubercles in the dorsal aspect of the body; anterior aspect of thigh with flatter scales, posterior aspect with granular scales.</paragraph>
|
|
<paragraph id="5EE0EDBAC8436D9DD1CC978AB8B79889" pageId="0" pageNumber="499">
|
|
Twenty pores in an enlarged row of femoral scales on both sides, separated medially by a diastema of four pore-less scales; subequal row of enlarged scales anterior to pore-bearing scales (Fig.
|
|
<figureCitation id="B290DFCCEE3CAF1FFA7BF39918454324" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">4D</figureCitation>
|
|
). Fore and hind limbs relatively short, stout; forearm short (FL/SVL 0.13); tibia short (CL/SVL 0.15); digits moderately short, strongly clawed; all digits of manus and digits I-IV of pes indistinctly webbed; terminal phalanx of all digits curved, arising angularly from distal portion of expanded lamellar pad, half or more than half as long as associated toepad; scansors beneath each toe in a straight transverse series, divided except for distal and four to five basal scansors on digit I: 10-13-13-12-13 (left manus), 10-12-13-12-13 (right manus) (Fig.
|
|
<figureCitation id="7B3B291C47ED0F6963201F3C44A376D3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">4E</figureCitation>
|
|
), 10-13-13-13-13 (left pes), 10-14-13-13-13 (right pes) (Fig.
|
|
<figureCitation id="E6D85BA7E4440D12FE6721FB63396FA5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hemidactylus pakkamalaiensis sp. nov. (Holotype, BNHS 2907), A head in dorsal view, B ventral view, C lateral view, D view of the femoral region, E ventral view of right manus and F right pes. Scale bar = 10 mm." figureDoi="10.3897/vz.73.e104494.figure4" httpUri="https://binary.pensoft.net/fig/853724" pageId="0" pageNumber="499">4F</figureCitation>
|
|
). Relative length of digits (measurements in mm in parentheses): I (6.8) <II (8.7) <III (8.3) <IV (8.2) <V (8) (left manus); I (6.8) <V (9.6) <IV (8.7) <II (9.3) <III (6.6 damaged) (left pes).
|
|
</paragraph>
|
|
<paragraph id="1CAC4ECEDB509D9E490BEF356E600FA1" pageId="0" pageNumber="499">Tail complete regenerated (about 30% of the tail original); depressed, flat beneath, verticillate, with well-defined median furrow; scales on the dorsal aspect of tail subimbricate, larger than granules on dorsum, with a series of 4-6 much enlarged, strongly pointed, moderately keeled tubercles; ventral scales enlarged, imbricate, median row (subcaudal plates) covering almost entire base of the tail, bordered laterally by two or three rows of larger pointed, smooth, imbricate scales; those close to vent small, smooth, flat and imbricate. Two indistinct postcloacal spurs on each side, much smaller than dorsal tubercles at midbody.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="6DBB49012259F1636B40F41E09DABF6B" pageId="0" pageNumber="499" type="variation">
|
|
<paragraph id="3EAF599B4C7AF859755874443F61A615" pageId="0" pageNumber="499">Variations in the paratypes.</paragraph>
|
|
<paragraph id="38B18885AF83F18C6A1EE8D41536413B" pageId="0" pageNumber="499">Variations among the meristic and morphometric characters in the paratypes are provided in the Table. 2. Two paratypes BNHS 2910 & 2911 are females. BNHS 2909 has a very short tail among all the type series. Up to eight enlarged tubercles in the tail in paratype BNHS 2909. postcloacal spur distinct in paratypes BNHS 2908 & BNHS 2909. Other differences include the number of undivided lamellae at the base of pes and manus that varies between three (BNHS 2908) to five (BNHS 2910 & BNHS 2911) in the paratypes and occasionally in the middle in BNHS 2910 & 2909. Gular scales that are in contact with the outer postmental much larger on the right side in BNHS 2909. The total number of the gular scales contacting the outer postmentals vary between eight and thirteen.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CADFE851DB1C27EAC9C331FC8ACE93F5" pageId="0" pageNumber="499" type="colour in life and preservative (based on holotype)">
|
|
<paragraph id="CA59A6F44E76C0A118712D222A04B84E" pageId="0" pageNumber="499">Colour in life and preservative (based on holotype).</paragraph>
|
|
<paragraph id="0B0CF050DF0128576BD22DD005699C47" pageId="0" pageNumber="499">
|
|
Dorsal aspect of the body, uniformly brownish with three indistinct transverse bands from the neck to the hind limb insertion and one on the tail, bordered by discontinuous brownish patches extending towards the lateral side of the body (Fig.
|
|
<figureCitation id="1FB6D52B50F63DDC631CEA7FAEA11890" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Dorsolateral view of the A holotype (BNHS 2907) and B one of the paratypes (BNHS 2911) of Hemidactylus pakkamalaiensis sp. nov. in life from Pakkamalai Hill, Tamil Nadu, India." figureDoi="10.3897/vz.73.e104494.figure6" httpUri="https://binary.pensoft.net/fig/853726" pageId="0" pageNumber="499">6A</figureCitation>
|
|
). Dorsal aspect of the head is brownish with inconspicuous white markings on the parietal and neck region. Ventral aspect of the head is mostly cream-coloured with light brown mottling on the gular region and mental, infralabials, post mentals and a few other adjacent scales mottled with dark brown. Fore and hindlimbs are brownish speckled with irregular black patches and a few scattered brownish-yellow spots on the dorsal side. Ventral aspect of the body, forelimbs and hindlimbs predominantly cream coloured with sprinkled and or mottled with brown colour. In preservative, the overall colouration is the same as in life except in dark transverse bands on the dorsum that became less prominent.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0BD3704D38A9D2300D7BA9B269DF5C82" pageId="0" pageNumber="499" type="distribution">
|
|
<paragraph id="966332E059ED76F62ADDDF05A5F0DFFF" pageId="0" pageNumber="499">Distribution and natural history.</paragraph>
|
|
<paragraph id="18BC9849610EDDB0E388F31A8A75A94F" pageId="0" pageNumber="499">
|
|
<taxonomicName id="BADD1BB929813D4042C54B857121E3E3" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
|
|
<emphasis id="18D98114FF183B5A8AF2A494401DBA13" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="26E905FCE2C8FD1A457330CDA5F3BAA0" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
|
|
is currently known from the Gingee Hills cluster. This species is nocturnal and appears to be locally abundant. In addition to the collected specimens, we encountered 56 live individuals in and around the collection site during our four-day fieldwork. All the specimens in the type series were gathered from rock boulders within a limited area at the type locality, mostly after 19:00 hrs. These hills are predominantly formed of granitic rock boulders and feature native vegetation, including thorny scrub jungle, moist deciduous forest, and tropical dry evergreen forest (
|
|
<bibRefCitation id="28449083B1F98F26D65480A33C3ADA0D" author="Arulappan, M" journalOrPublisher="American Journal of Ethnomedicine" pageId="0" pageNumber="499" pagination="84 - 102" refId="B8" refString="Arulappan, M, Britto, SJ, Ruckmani, K, Mohankumar, R, 2015. An ethnobotanical study of medicinal plants used by ethnic people in Gingee, Villupuram District, Tamil Nadu, India. American Journal of Ethnomedicine 2: 84 - 102" title="An ethnobotanical study of medicinal plants used by ethnic people in Gingee, Villupuram District, Tamil Nadu, India." volume="2" year="2015">Arulappan et al. 2015</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="445C906A436502052D735943893572F5" doi="10.3897/vz.73.e104494.figure7" httpUri="https://binary.pensoft.net/fig/853727" pageId="0" pageNumber="499" start="Figure 7" startId="F7">
|
|
<paragraph id="1E323AED89257E150A8D2BC991AF2B72" pageId="0" pageNumber="499">
|
|
<emphasis id="4B3F67A48C48FDC8CA87876C0B31E102" bold="true" pageId="0" pageNumber="499">Figure 7.</emphasis>
|
|
Habitat at the type locality (Pakkamalai Hill, Tamil Nadu, India) of
|
|
<taxonomicName id="F91953C4B2EBEEEE6E565D0C44A86C66" authorityName="Narayanan, Christopher, Raman, Mukherjee, Vimalraj, Deepak" authorityYear="2023" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" lsidName="Hemidactylus pakkamalaiensis" order="Squamata" pageId="0" pageNumber="499" rank="species" species="pakkamalaiensis">
|
|
<emphasis id="3630A49EF87332442D0C8BEAA2467B06" italics="true" pageId="0" pageNumber="499">Hemidactylus pakkamalaiensis</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="D69A6A0661A0296ACF150C8E143ACAA0" bold="true" pageId="0" pageNumber="499">sp. nov.</emphasis>
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |