310 lines
29 KiB
XML
310 lines
29 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.644.10413" ID-GBIF-Dataset="9dd88063-e93b-4f8e-8f61-9612ff6680ac" ID-PMC="PMC5242256" ID-Pensoft-Pub="1313-2970-644-1" ID-PubMed="28144182" ID-ZBK="AC258A039F0C45DCB87F6100E6E936E4" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-644-1" ModsDocOrigin="ZooKeys 644" ModsDocTitle="Baetis (Baetis) cypronyx sp. n., a new species of the Baetisalpinus species-group (Insecta, Ephemeroptera, Baetidae) from Cyprus, with annotated checklist of Baetidae in the Mediterranean islands" checkinTime="1484230734844" checkinUser="pensoft" docAuthor="Godunko, Roman J., Soldan, Tomas & Staniczek, Arnold H." docDate="2017" docId="3EEBBB533AC50D3BFE148F1C5DD86D2D" docLanguage="en" docName="ZooKeys 644: 1-32" docOrigin="ZooKeys 644" docSource="http://dx.doi.org/10.3897/zookeys.644.10413" docTitle="Baetis (Baetis) cypronyx Godunko, Soldan & Staniczek, 2017, sp. n." docType="treatment" docUuid="0B11F59C-97F2-42F2-AFD4-A68B6E3D3DCB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="11" masterDocId="7011FF86FF8D63093E33FF82FFC7BA0B" masterDocTitle="Baetis (Baetis) cypronyx sp. n., a new species of the Baetisalpinus species-group (Insecta, Ephemeroptera, Baetidae) from Cyprus, with annotated checklist of Baetidae in the Mediterranean islands" masterLastPageNumber="32" masterPageNumber="1" pageNumber="3" updateTime="1668163866340" updateUser="ExternalLinkService">
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<mods:titleInfo>
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<mods:title>Baetis (Baetis) cypronyx sp. n., a new species of the Baetisalpinus species-group (Insecta, Ephemeroptera, Baetidae) from Cyprus, with annotated checklist of Baetidae in the Mediterranean islands</mods:title>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Godunko, Roman J.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Soldan, Tomas</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Staniczek, Arnold H.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>644</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>32</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.644.10413</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.644.10413</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-644-1</mods:identifier>
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<mods:identifier type="ZBK">AC258A039F0C45DCB87F6100E6E936E4</mods:identifier>
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<mods:identifier type="ZooBank">AC258A039F0C45DCB87F6100E6E936E4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127875939" LSID="urn:lsid:zoobank.org:act:0B11F59C-97F2-42F2-AFD4-A68B6E3D3DCB" httpUri="http://treatment.plazi.org/id/3EEBBB533AC50D3BFE148F1C5DD86D2D" lastPageId="10" lastPageNumber="11" pageId="2" pageNumber="3">
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<subSubSection pageId="2" pageNumber="3" type="nomenclature">
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<paragraph pageId="2" pageNumber="3">
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<taxonomicName LSID="http://zoobank.org/0B11F59C-97F2-42F2-AFD4-A68B6E3D3DCB" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis (Baetis) cypronyx" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="cypronyx" subGenus="Baetis">Baetis (Baetis) cypronyx</taxonomicName>
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<taxonomicNameLabel pageId="2" pageNumber="3">sp. n.</taxonomicNameLabel>
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Figs 1-2, 3-4, 5, 6-8, 9-12, 13-14, 15, 16-18, 19-22, 23-24
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="3" type="type material">
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<paragraph pageId="2" pageNumber="3">Type material.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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Holotype: mature larva, CYPRUS, Limassol [Lemesos;
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<normalizedToken originalValue="Λεμεσός">Λεμεσός</normalizedToken>
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] District, Troodos [Τρόοδος] Mts., Kryos River [Κρύος
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<normalizedToken originalValue="ποταμός">ποταμός</normalizedToken>
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], Kalidonia Waterfalls, app. 1250 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="34.892685">N34 53.561</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.867382">E32 52.043</geoCoordinate>
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, 22.v.2004, leg. T.
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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.
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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Paratypes: 75 larvae, the same date and place as holotype; 14 larvae, CYPRUS, Limassol [Lemesos;
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<normalizedToken originalValue="Λεμεσός">Λεμεσός</normalizedToken>
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] District, Troodos [Τρόοδος] Mts., Diplos River [Διπλός
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<normalizedToken originalValue="ποταμός">ποταμός</normalizedToken>
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], Chantara [Xantara] Waterfalls, near Trooditissa [Μοναστήρι
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<normalizedToken originalValue="Τροοδίτισσας">Τροοδίτισσας</normalizedToken>
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] Monastery, app. 1300 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="34.90715">N34 54.429</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.838383">E32 50.303</geoCoordinate>
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, 23.v.2004, leg. T.
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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;
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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4 larvae, ibid., Paphos District [
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<normalizedToken originalValue="Επαρχία">Επαρxία</normalizedToken>
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<normalizedToken originalValue="πάφου">πάφου</normalizedToken>
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], Gialia River [Γιαλιά], in the forest
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<normalizedToken originalValue="“Pochalantra”">"Pochalantra"</normalizedToken>
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, app. 5 km upstream from Gialia [Γιαλιά] village, app. 400-410 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="35.072735">N35 04.364</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.55958">E32 33.575</geoCoordinate>
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, 12.xi.2005, leg. A. Buffagni;
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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10 larvae, United Nations Buffer Zone in Cyprus, Nicosia District [
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<normalizedToken originalValue="Επαρχία">Επαρxία</normalizedToken>
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<normalizedToken originalValue="Λευκωσίας">Λευκωσίας</normalizedToken>
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], upstream of Kargotis River [Καρκώτη], vicinity of Kakopetriya [Κακοπετριά] village, Mitro place, app. 150-200 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="34.983532">N34 59.012</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.9">E32 54.000</geoCoordinate>
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, 22.iii.2006, leg. A. Buffagni;
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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2 larvae, ibid., Agios Nikolaos Lefkas [
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<normalizedToken originalValue="Άγιος">'aγιος</normalizedToken>
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<normalizedToken originalValue="Νικόλαος">Νικόλαος</normalizedToken>
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<normalizedToken originalValue="Λεύκας">Λεύκας</normalizedToken>
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] village (abandoned), app. 100-120 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="35.088">N35 5.280</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="1" value="32.891666">E32 53.500</geoCoordinate>
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, 24.iii.2006, leg. A. Buffagni.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph pageId="2" pageNumber="3">Comparative material.</paragraph>
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<paragraph pageId="2" pageNumber="3">
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis baroukianus" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="baroukianus">Baetis baroukianus</taxonomicName>
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Thomas & Dia, 1984: 1 male and 1 female mature larvae, LEBANON, Chouf District, type locality of
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis baroukianus" order="Ephemeroptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="baroukianus">Baetis baroukianus</taxonomicName>
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, branch of Salam (
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<normalizedToken originalValue="Râs">Ras</normalizedToken>
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el
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<normalizedToken originalValue="Mâ">Ma</normalizedToken>
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) spring near
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<normalizedToken originalValue="Harêt">Haret</normalizedToken>
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Jandal Municipality, app. 800 m a.s.l., 25.vii.1979, leg. Dia A. (see
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<bibRefCitation author="Thomas, AGB" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de Toulouse" pageId="19" pageNumber="20" pagination="7 - 12" title="Baetisbaroukianus n. sp., Ephemeroptere nouveau du Liban (Baetidae)." url="https://doi.org/10.1051/limn/1984036" volume="120" year="1984">Thomas and Dia 1984</bibRefCitation>
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: 10).
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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28 larvae (10 males, 18 females), IRAN [new record], Elburz Mts., Gilan Province, Rudbar County, Central District, unnamed brook in Divresh village, right tributary of Siah Rud River (SE upstream of Shirkooh village), app. 285 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="36.893166">N36 53.59</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="49.58433">E49 35.06</geoCoordinate>
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, 13.v.2016, leg.
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<normalizedToken originalValue="Bojková">Bojkova</normalizedToken>
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J.,
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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T. & J. Imanpour Namin, det. Sroka P.
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</paragraph>
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<paragraph pageId="2" pageNumber="3">
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2 larvae (1 male, 1 female), ibid, Fuman County, Sardar-e Jangal District, unnamed brook below of Masuleh City (right tributary of Rudkhan River), app. 710 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="37.157">N37 09.42</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="49.0195">E49 01.17</geoCoordinate>
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, 22.v.2016, leg.
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<normalizedToken originalValue="Bojková">Bojkova</normalizedToken>
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J.,
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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T. & J. Imanpour Namin, det. Sroka P.
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<pageBreakToken pageId="3" pageNumber="4" start="start">3</pageBreakToken>
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larvae (1 male, 2 females), ibid, Rudbar County, Central District, unnamed brook, left tributary of
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<normalizedToken originalValue="Sefīd-Rūd">Sefīd-Rūd</normalizedToken>
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River, below Rostamabad City, app. 155 m a.s.l.,
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<geoCoordinate direction="north" orientation="latitude" precision="9" value="37.157833">N37 09.47</geoCoordinate>
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="49.002834">E49 00.17</geoCoordinate>
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, 22.v.2016, leg.
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<normalizedToken originalValue="Bojková">Bojkova</normalizedToken>
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J.,
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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T. & J. Imanpour Namin, det. Sroka P.
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</paragraph>
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<paragraph pageId="3" pageNumber="4">
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis melanonyx" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="melanonyx">Baetis melanonyx</taxonomicName>
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(Pictet, 1843): 30 larvae (7 larvae mounted with Liquide de Faure), Czech Republic,
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<normalizedToken originalValue="Ústí">Usti</normalizedToken>
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nad Labem district, Elbe river-basin,
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<normalizedToken originalValue="Divoká">Divoka</normalizedToken>
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Orlice River,
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<normalizedToken originalValue="Líšnice">Lisnice</normalizedToken>
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village, 432 m a.s.l., 2.vii.1972, leg. T.
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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(for details see
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<bibRefCitation author="Soldan, T" journalOrPublisher="Acta Entomologica Bohemoslovaca" pageId="18" pageNumber="19" pagination="319 - 329" title="Mayflies (Ephemeroptera) new to the fauna of Czechoslovakia found in 1972 − 1977." volume="75" year="1978">
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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1978
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</bibRefCitation>
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); 24 larvae (8 larvae mounted with Euparal), Germany:
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<normalizedToken originalValue="Baden-Württemberg">Baden-Wuerttemberg</normalizedToken>
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, Boll, vor Tannegger Wasserfall, Wutach River, 623 m a.s.l., 03.vi.2008, leg. B. Frey. For other comparative material of
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis melanonyx" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="melanonyx">Baetis melanonyx</taxonomicName>
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see
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<bibRefCitation author="Godunko, RJ" journalOrPublisher="Vestnik Zoologii" pageId="17" pageNumber="18" pagination="23 - 28" title="Species of the Alpinus Group of the Genus Baetis (Ephemeroptera, Baetidae) from Ukraine." volume="33" year="1999">Godunko (1999)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4" type="diagnosis">
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<paragraph pageId="3" pageNumber="4">Diagnosis.</paragraph>
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<paragraph lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis cypronyx" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="cypronyx">Baetis cypronyx</taxonomicName>
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sp. n. differs from all other representatives of the
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis alpinus" order="Ephemeroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="alpinus">Baetis alpinus</taxonomicName>
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species-group by the following combination of larval characters (see Table 1): (i) labrum of distinctly oblong shape, nearly rectangular (Fig. 6
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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), (іі) outer mandibular incisor group distinctly fused, narrow and triangular (Fig. 8); (iii) segment 2 of maxillary palps usually with single seta, exceptionally with two stout apical setae
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<pageBreakToken pageId="4" pageNumber="5" start="start">(</pageBreakToken>
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Figs 11
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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, 19); (vi) paraglossae with two irregular rows of long, stout bristles apically (Fig. 9); (v) segment 3 of labial palps not elongated, nearly symmetrical and evenly rounded (Fig. 12
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<normalizedToken originalValue="a–c">a-c</normalizedToken>
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); (vi) sternal protuberances on meso- and metathorax pointed apically; (vii) outer margin of femora with 2-3 rows of long, apically obtuse to bluntly pointed bristles proximally and centrally (Figs 13, 20, 21); (viii) irregular row of small submarginal spines on abdominal terga
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<normalizedToken originalValue="III–VIII">III-VIII</normalizedToken>
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(IX) (Fig. 22); (ix) surface of abdominal terga with few scales in sockets, scales triangular to tongue-shaped, not elongated, mostly lacking on tergum X (Figs 23, 24); (x) paraproct plate with bluntly pointed stout setae near to inner margin (Figs 16, 17); (xi) paracercus strongly reduced, 2-16 segmented (Fig. 24).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="11" pageId="4" pageNumber="5" type="description">
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<paragraph pageId="4" pageNumber="5">Description.</paragraph>
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<paragraph pageId="4" pageNumber="5">Mature larva: female body length: 7.5−8.0 mm, length of cerci: 9.0−11.5 mm; male body length: 6.0−8.0 mm; length of cerci: 7.0−10.0 mm; paracercus vestigial or strongly reduced.</paragraph>
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<paragraph pageId="5" pageNumber="6">
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<pageBreakToken pageId="5" pageNumber="6" start="start">Cuticular</pageBreakToken>
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coloration (Figs 1-5). Due to ten to twelve years of material storage in ethanol, the herein described colour pattern might be slightly paler compared to fresh material.
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</paragraph>
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<paragraph pageId="5" pageNumber="6">General colour yellowish brown to brown. Head light brown with paler genae; clypeus light brown; frons with several small, isolated brown spots. Antennae light brown, flagellum paler than scape and pedicel.</paragraph>
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<paragraph pageId="5" pageNumber="6">Pronotum yellowish brown with two pairs of oblique brownish bands; mesonotum yellowish brown to brown, with longitudinal brown bands centrally, and several spots of the same colour centrally and laterally; metanotum brown with darker smudge centrally (Figs 1, 4). Lateral sides of thorax with brown pleurites (Fig. 2). Ventral side of thorax paler than dorsal side; sterna yellowish (Fig. 3). Legs pale. Femora yellowish brown with two darker, usually isolated longitudinal spots along outer margin; tibia light brown; base and apex of tarsi brown, darker than middle part; tarsal claw brown (Figs 2, 3).</paragraph>
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<paragraph pageId="6" pageNumber="7">
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<pageBreakToken pageId="6" pageNumber="7" start="start">Abdominal</pageBreakToken>
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terga (Figs 1, 4) yellowish brown to brown; terga
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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(IV) and
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<normalizedToken originalValue="VI–VIII">VI-VIII</normalizedToken>
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darker. Terga
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<normalizedToken originalValue="I–III">I-III</normalizedToken>
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(IV) brownish centrally, with broad pale area laterally; median brown spot on terga III and IV occasionally divided into two longitudinal spots; all terga with more or less well visible brownish band along anterior margin of segment; a pair of diffuse brownish maculae near posterior margin of terga
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<normalizedToken originalValue="V–VIII">V-VIII</normalizedToken>
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; a pair of brownish paramedian dots on terga
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<normalizedToken originalValue="II–X">II-X</normalizedToken>
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, terga
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<normalizedToken originalValue="III–VI">III-VI</normalizedToken>
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occasionally with additional oblique streaks fused with paramedian dots and forming a diffuse brownish U-shaped pattern in anterior half of segment. Abdominal sterna with a pair of sublateral elongated spots. Cerci yellowish brown to brown, 3-5 first segments slightly darker.
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</paragraph>
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<paragraph pageId="6" pageNumber="7">Hypodermal coloration. Hypoderm without contrasting markings.</paragraph>
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<paragraph pageId="7" pageNumber="8">
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<pageBreakToken pageId="7" pageNumber="8" start="start">Head</pageBreakToken>
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. Surface of clypeus and frons covered with solitary FT, B, and Hr setae. Larval turbinate eyes brown to intensively brown apically. Antennae slightly longer than 1/2 of body length. Scape and pedicel with solitary FT and Hr, and more abundant B setae only, without any particular cuticular ornamentation (e.g. corrugation/chagrin; see Bauernfeind and
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<normalizedToken originalValue="Soldán">Soldan</normalizedToken>
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2012), which is present in some representatives of the
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<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis alpinus" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="alpinus">Baetis alpinus</taxonomicName>
|
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species-group and in the closely related
|
||
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis lutheri" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lutheri">Baetis lutheri</taxonomicName>
|
||
and
|
||
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis pavidus" order="Ephemeroptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pavidus">Baetis pavidus</taxonomicName>
|
||
species-groups.
|
||
</paragraph>
|
||
<paragraph pageId="7" pageNumber="8">
|
||
Mouthparts. Labrum (Figs 6
|
||
<normalizedToken originalValue="a–c">a-c</normalizedToken>
|
||
) distinctly wider than long, nearly rectangular, with width/length ratio 1.80-1.88; dorsal surface with 1 + 11-18 long submarginal setae, arranged in a single irregular row (occasionally 1-4 bristles form an additional weekly defined row); 6-8 smaller setae laterally on both margins; dorsal surface of labrum covered with sparsely arranged B and only a few FT setae grouped mainly posterolaterally; ventral side with 2-5 small pointed setae anterolaterally. Median incurvation of middle part of anterior margin of labrum clearly shallow and wide.
|
||
</paragraph>
|
||
<paragraph pageId="7" pageNumber="8">Outer mandibular incisor group narrow and triangular, distinctly fused; inner incisor group not prominent, with 3-4 small teeth (of which most distal tooth is the biggest), both groups separated by a shallow incision. Right and left prostheca of same size, nearly symmetrical, with 8-10 apical teeth (Fig. 8).</paragraph>
|
||
<paragraph lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
|
||
Maxillary palp two-segmented; segment 1 shorter than second segment; segment 2 asymmetrical apically, with pronounced tip (conical protuberance), and one dis
|
||
<pageBreakToken pageId="8" pageNumber="9" start="start">tinct</pageBreakToken>
|
||
, stout seta; one additional stout seta occasionally near apex of segment 2; surface of both segments with B setae [uniporous sensillum basiconicum sensu
|
||
<bibRefCitation author="Gaino, E" editor="Gaino, E" journalOrPublisher="Universita di Perugia, Perugia" pageId="17" pageNumber="18" pagination="445 - 451" title="The sensilla on the labial and maxillary palps of the nymph of Baetisrhodani (Ephemeroptera: Baetidae)." volumeTitle="Research update on Ephemeroptera & Plecoptera" year="2003">Gaino and Rebora (2003</bibRefCitation>
|
||
: 449, figs 19−21)] which are clearly dense on distal part of segment 2 (Figs 11
|
||
<normalizedToken originalValue="a–c">a-c</normalizedToken>
|
||
, 19).
|
||
</paragraph>
|
||
<paragraph pageId="8" pageNumber="9">Hypopharynx relatively slender, anterior side laterodistally covered with fine, elongated setae along outer margins of lingua and superlinguae, lingua with prominent central lobe, superlinguae with marked hump (Fig. 7).</paragraph>
|
||
<paragraph pageId="8" pageNumber="9">
|
||
Labium with relatively slender glossae, slightly shorter than paraglossae (Figs 9, 10); glossae each with 8-10 stout bristles on inner margin, and 3-5 bristles on outer margin; 5-7 pairs of long, stout bristles form two irregular rows on tip of paraglossae; additionally 5-10 long bristles along outer margin and 3-5 medium sized setae on ventral side of paraglossae. Segment 2 of labial palp 1.30-1.42 longer than segment 3, covered only with sparse B and Hr setae; segment 3 not elongated, nearly symmetrical and evenly rounded, only slightly broader than long (width/length ratio 1.03-1.07); surface of segment 3 with 18-25 slender, pointed, stout setae [long and short hairs sensu
|
||
<bibRefCitation author="Gaino, E" editor="Gaino, E" journalOrPublisher="Universita di Perugia, Perugia" pageId="17" pageNumber="18" pagination="445 - 451" title="The sensilla on the labial and maxillary palps of the nymph of Baetisrhodani (Ephemeroptera: Baetidae)." volumeTitle="Research update on Ephemeroptera & Plecoptera" year="2003">Gaino and Rebora (2003)</bibRefCitation>
|
||
], and long Hr setae; quotient q changes from 0.76 to 0.88 (see
|
||
<bibRefCitation author="Sroka, P" journalOrPublisher="Zootaxa" pageId="19" pageNumber="20" pagination="27 - 49" title="Morphological and genetic variability of Baetis (Rhodobaetis) braaschi Zimmermann, 1980 (Ephemeroptera: Baetidae)." volume="3323" year="2012">Sroka et al. 2012</bibRefCitation>
|
||
, 29, 31: fig. 2) (Fig. 12
|
||
<normalizedToken originalValue="a–c">a-c</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">
|
||
<pageBreakToken pageId="9" pageNumber="10" start="start">Thorax</pageBreakToken>
|
||
. Surface of pronotum with few FT and Hr setae only. Sternal protuberances on meso- and metathorax well visible, pointed apically, yellowish brown to brown.
|
||
</paragraph>
|
||
<paragraph pageId="9" pageNumber="10">Outer margin of femora with 2-3 rows of long bristles with obtuse to bluntly pointed tips proximally and centrally (Figs 13, 20, 21), and one row of shorter and stouter obtuse bristles distally; central part of outer margin of femora occasionally with long bristles arranged in 1-2 rows. Long marginal bristles alternating with submarginal STSm-bp setae and elongated Hr setae. Inner margin with 2-6 STSs-bp setae near to proximal end. Surface of femora with STSs-bp and STSs-ov setae and tiny setae [Hr and more abundant FT setae]. Outer and inner margins of tibiae with STSm-p and STSm-bp setae and short Hr setae; surface of tibia with STSs-bp to nearly STSs-ov setae; a group of long Hr setae near distal end of outer margin of tibia. Tarsi with 6-10 middle to elongated STS-p setae along the inner margin, and several STSm-p and/or STSm-bp setae on outer margin; both margins of tarsi covered with tiny Hr setae; surface of tarsi with a few FT and more abundant Hr setae, and small STS-bp setae. Tarsal claws not elongated, moderately hooked; with 10-11 teeth arranged in single row and two subapical tiny Hr setae (Figs 13, 14).</paragraph>
|
||
<paragraph lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
|
||
Abdomen. Posterior margin of terga with broad triangular spines of different size, bluntly pointed or occasionally pointed apically; broader spines along posterior margin of terga
|
||
<normalizedToken originalValue="III–VIII">III-VIII</normalizedToken>
|
||
; spines alternating with 1-3 tiny B and a single Hr setae. Irregular
|
||
<pageBreakToken pageId="10" pageNumber="11" start="start">row</pageBreakToken>
|
||
of smaller submarginal spines on terga
|
||
<normalizedToken originalValue="III–VIII">III-VIII</normalizedToken>
|
||
(IX) (Fig. 22). Surface of terga with few, not elongated, tongue-shaped [SC-tg] to triangular [SC-it; bluntly pointed to rounded apically] scales, and their few sockets (mainly lacking on tergum X), concentrated on central part of segment (Figs 23, 24); solitary Hr and more abundant FT setae stretched over the whole surface of terga
|
||
<normalizedToken originalValue="I–X">I-X</normalizedToken>
|
||
. Posterior margin and surface of sterna without spines, stout setae or scales, with B and Hr setae only.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">Paraproct plate as in Figs 16-18. Inner margin of paraproct with 8-12 spines of different size along apical half, alternating with tiny setae [solitary FT and more abundant B setae], and 2-8 (mainly 4-7) submarginal STSm-bp setae (Figs 16, 17); a single row of relatively small and stout spines along inner margin of cercotractor (for definition of cercotractor see Kluge 2004) (Fig. 18). Surface of paraproct covered with sparse FT, B and Hr setae and their bases only.</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">
|
||
Tracheal gills whitish yellow to light brown, not elongated, broadly rounded apically (Fig. 15,
|
||
<normalizedToken originalValue="I–VII">I-VII</normalizedToken>
|
||
); gills I and VII nearly symmetrical; gills
|
||
<normalizedToken originalValue="II–VI">II-VI</normalizedToken>
|
||
asymmetrical; serrated margins of gills more or less well marked, with tiny Hr setae inserting in small, articulated bases; tracheation poorly visible.
|
||
</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">Cerci as long as 1.20-1.32 of body length. Paracercus reduced to 2-16 segments (Fig. 24). Posterior margin of cercal and paracercal segments with row of broad, triangular spines, and uneven submarginal row of smaller spines. Length of paracercus of mature larvae apparently variable in different populations, as well as in specimens within each population. Paracercus in larvae from Cryos River (type locality) vestigial (evidently shorter than abdominal tergum X, consisting of approximately up to 5-7 segments, some segments at least partially fused, Fig. 1); paracercus in paratype larvae from Diplos River strongly reduced (but evidently longer than abdominal tergum X), only consisting of about 10 or more apparently separated or distinguishable segments; Fig. 4).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="11" type="male and female adults">
|
||
<paragraph pageId="10" pageNumber="11">Male and female adults.</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph pageId="10" pageNumber="11">Etymology.</paragraph>
|
||
<paragraph pageId="10" pageNumber="11">
|
||
The specific epithet is a combination of the name of Cyprus, where the new species was found, and the specific epithet of the closely related species
|
||
<taxonomicName class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis melanonyx" order="Ephemeroptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="melanonyx">Baetis melanonyx</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |