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<document id="1D591B78226EA9BEB2F6AC9C4D2F88D2" ID-DOI="10.11646/zootaxa.3702.1.2" ID-GBIF-Dataset="70f89513-ee63-4a9a-8b31-0264324e0a28" ID-ISSN="1175-5326" ID-Zenodo-Dep="284544" ID-ZooBank="D7F22D44-604A-411E-8F1D-C312BB52212A" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460669573999" checkinUser="plazi" docAuthor="Schwendinger, Peter J." docDate="2013" docId="3E5A87D65920AD34C2F2A548FE0AFD17" docLanguage="en" docName="zt03702p060.pdf" docOrigin="Zootaxa 3702 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Liphistius laoticus Schwendinger, 2013, sp. n." docType="treatment" docVersion="5" lastPageNumber="59" masterDocId="C263FFAE5925AD3CC265A774FFB7FFE7" masterDocTitle="On two Liphistius species (Araneae: Liphistiidae) from Laos" masterLastPageNumber="60" masterPageNumber="51" pageNumber="56" updateTime="1698348034261" updateUser="plazi">
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<mods:title id="C98444959B6A4B8BE0704214F86ECF10">On two Liphistius species (Araneae: Liphistiidae) from Laos</mods:title>
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<mods:namePart id="68C87066A3BB62177DAB340C81296CC1">Schwendinger, Peter J.</mods:namePart>
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<mods:date id="90D879DF28CC51BE06ED64DED204DFC0">2013</mods:date>
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<mods:number id="F0331FBB721A7BB59B845F3C9B5D3D12">3702</mods:number>
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<treatment id="3E5A87D65920AD34C2F2A548FE0AFD17" ID-DOI="http://doi.org/10.5281/zenodo.6155044" ID-GBIF-Taxon="127665380" ID-Zenodo-Dep="6155044" LSID="urn:lsid:plazi:treatment:3E5A87D65920AD34C2F2A548FE0AFD17" httpUri="http://treatment.plazi.org/id/3E5A87D65920AD34C2F2A548FE0AFD17" lastPageId="8" lastPageNumber="59" pageId="5" pageNumber="56">
<subSubSection id="FEE9654B5920AD39C2F2A548FE74FD9F" pageId="5" pageNumber="56" type="nomenclature">
<paragraph id="B64C36C05920AD39C2F2A548FE08FDB0" blockId="5.[151,451,572,632]" box="[151,447,572,599]" pageId="5" pageNumber="56">
<heading id="ED0481AC5920AD39C2F2A548FE08FDB0" bold="true" box="[151,447,572,599]" fontSize="11" level="1" pageId="5" pageNumber="56" reason="1">
<emphasis id="8487EAD25920AD39C2F2A548FE08FDB0" bold="true" box="[151,447,572,599]" pageId="5" pageNumber="56">
<taxonomicName id="71F34D435920AD39C2F2A548FEC2FDB1" ID-CoL="6QJ5H" box="[151,373,572,598]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="laoticus" status="sp. nov.">
<emphasis id="8487EAD25920AD39C2F2A548FEC2FDB1" bold="true" box="[151,373,572,598]" italics="true" pageId="5" pageNumber="56">Liphistius laoticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FB457A95920AD39C319A549FE08FDB0" box="[380,447,573,599]" pageId="5" pageNumber="56" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="B64C36C05920AD39C2F2A52BFE74FD9F" blockId="5.[151,451,572,632]" box="[151,451,607,632]" pageId="5" pageNumber="56">
<figureCitation id="2EC82A455920AD39C2F2A52BFEB7FD9F" box="[151,256,607,632]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1896]" captionTargetBox="[161,1429,894,1844]" captionTargetId="figure@3.[151,1436,877,1852]" captionTargetPageId="3" captionText="FIGURE 2. Liphistius isan, male (A, F) and female (B) from Phou Asa, Laos; L. laoticus sp. n., female allotype (C), male holotype (D); L. thoranie, male holotype (E). A B, habitus. C F, sternum and labium, ventral view." httpUri="https://zenodo.org/record/284546/files/figure.png" pageId="5" pageNumber="56">Figures 2</figureCitation>
CD, 4AJ, NQ
</paragraph>
</subSubSection>
<subSubSection id="FEE9654B5920AD39C2F2A5D3FD77FC7F" pageId="5" pageNumber="56" type="materials_examined">
<paragraph id="B64C36C05920AD39C2F2A5D3FD77FC7F" blockId="5.[151,1436,679,2037]" pageId="5" pageNumber="56">
<emphasis id="8487EAD25920AD39C2F2A5D3FEB1FD27" bold="true" box="[151,262,679,704]" pageId="5" pageNumber="56">Material.</emphasis>
<collectingCountry id="CEE476505920AD39C377A5DCFED7FD27" box="[274,352,680,704]" name="Laos" pageId="5" pageNumber="56">LAOS</collectingCountry>
: Champasak Province, Bolaven Plateau, SW of Pakxong, near Tad Fane (= Fane Waterfall) (
<geoCoordinate id="D3C750075920AD39C2FAA5B8FE9BFD02" box="[159,300,716,741]" direction="north" orientation="latitude" pageId="5" pageNumber="56" precision="15" value="15.181667">15°1054”N</geoCoordinate>
,
<geoCoordinate id="D3C750075920AD39C35EA5B8FE65FD02" box="[315,466,716,741]" direction="east" orientation="longitude" pageId="5" pageNumber="56" precision="15" value="106.12723">106°0738”E</geoCoordinate>
),
<quantity id="710B9B255920AD39C38CA5B8FD80FD03" box="[489,567,716,740]" metricMagnitude="2" metricUnit="m" metricValue="9.3" pageId="5" pageNumber="56" unit="m" value="930.0">930 m</quantity>
; male
<typeStatus id="694888625920AD39C0E3A5B8FD5FFD02" box="[646,744,716,741]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
(matured
<date id="C24D10005920AD39C107A5B8FC58FD03" box="[866,1007,716,740]" pageId="5" pageNumber="56" value="1997-07-31">31.VII.1997</date>
),
<specimenCount id="A0F5FD495920AD39C663A5B8FBE3FD02" box="[1030,1108,716,741]" pageId="5" pageNumber="56" type="male">5 male</specimenCount>
<typeStatus id="694888625920AD39C63AA5B9FB7DFD02" box="[1119,1226,717,741]" pageId="5" pageNumber="56" type="paratype">paratypes</typeStatus>
(matured
<date id="C24D10005920AD39C721A5B8FA20FD03" box="[1348,1431,716,740]" pageId="5" pageNumber="56" value="1997-07">30.VII.</date>
,
<date id="C24D10005920AD39C2F2A584FF52FCE0" box="[151,229,752,775]" pageId="5" pageNumber="56" value="1997-08">7.VIII.</date>
,
<date id="C24D10005920AD39C290A584FEF3FCEF" box="[245,324,752,776]" pageId="5" pageNumber="56" value="1997-08">8.VIII.</date>
,
<date id="C24D10005920AD39C331A584FE2BFCEF" box="[340,412,752,776]" pageId="5" pageNumber="56" value="1997-08">9.VIII</date>
,
<date id="C24D10005920AD39C3C9A584FD88FCEF" box="[428,575,752,776]" pageId="5" pageNumber="56" value="1997-08-27">27.VIII.1997</date>
) and
<specimenCount id="A0F5FD495920AD39C0E2A584FD5AFCEF" box="[647,749,751,776]" pageId="5" pageNumber="56" type="female">3 female</specimenCount>
<typeStatus id="694888625920AD39C09DA584FCD4FCEF" box="[760,867,752,776]" pageId="5" pageNumber="56" type="paratype">paratypes</typeStatus>
(one of them the “allotype”, moulted
<date id="C24D10005920AD39C748A584FA39FCEF" box="[1325,1422,752,776]" pageId="5" pageNumber="56" value="1997-01-04">4.I.1997</date>
);
<date id="C24D10005920AD39C2F2A460FE94FCCA" box="[151,291,788,813]" pageId="5" pageNumber="56" value="1996-12-26">26.XII.1996</date>
. Same locality,
<quantity id="710B9B255920AD39C386A460FD85FCCB" box="[483,562,788,812]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="56" unit="m" value="900.0">900 m</quantity>
;
<specimenCount id="A0F5FD495920AD39C026A460FD1EFCCA" box="[579,681,788,813]" pageId="5" pageNumber="56" type="female">1 female</specimenCount>
<typeStatus id="694888625920AD39C0D3A461FCADFCCA" box="[694,794,789,813]" pageId="5" pageNumber="56" type="paratype">paratype</typeStatus>
;
<date id="C24D10005920AD39C14EA460FC25FCCB" box="[811,914,788,812]" pageId="5" pageNumber="56" value="2010-10-02">2.X.2010</date>
(sample LT-10/27). NW of Pakxong,
<taxonomicName id="71F34D435920AD39C73EA460FA2BFCCA" box="[1371,1436,788,813]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="58" phylum="Arthropoda" rank="species" species="tham">Tham</taxonomicName>
Champee (= Champee Cave) (
<geoCoordinate id="D3C750075920AD39C066A443FD38FCB7" box="[515,655,823,848]" direction="north" orientation="latitude" pageId="5" pageNumber="56" precision="15" value="15.201111">15°1204”N</geoCoordinate>
,
<geoCoordinate id="D3C750075920AD39C0C4A443FC8FFCB7" box="[673,824,823,848]" direction="east" orientation="longitude" pageId="5" pageNumber="56" precision="15" value="106.13528">106°0807”E</geoCoordinate>
),
<quantity id="710B9B255920AD39C137A44CFC15FCA8" box="[850,930,824,848]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="5" pageNumber="56" unit="m" value="980.0">980 m</quantity>
;
<specimenCount id="A0F5FD495920AD39C1D6A44CFBB3FCB7" box="[947,1028,823,848]" pageId="5" pageNumber="56" type="male">1 male</specimenCount>
<typeStatus id="694888625920AD39C677A44CFBC5FCB7" box="[1042,1138,824,848]" pageId="5" pageNumber="56" type="paratype">paratype</typeStatus>
and
<specimenCount id="A0F5FD495920AD39C6D0A44CFAABFCB7" box="[1205,1308,823,848]" pageId="5" pageNumber="56" type="female">3 female</specimenCount>
<typeStatus id="694888625920AD39C74CA44CFA2FFCB7" box="[1321,1432,824,848]" pageId="5" pageNumber="56" type="paratype">paratypes</typeStatus>
;
<date id="C24D10005920AD39C2F2A428FF48FC93" box="[151,255,860,884]" pageId="5" pageNumber="56" value="2010-10-02">2.X.2010</date>
(sample LT-10/26). All specimens collected by P.J. Schwendinger. One male and one female
<typeStatus id="694888625920AD39C759A429FA2BFC92" box="[1340,1436,861,885]" pageId="5" pageNumber="56" type="paratype">paratype</typeStatus>
deposited in SMF, all other specimens in MHNG.
</paragraph>
</subSubSection>
<subSubSection id="FEE9654B5920AD39C2A2A4D0FAE7FC5B" box="[199,1360,932,957]" pageId="5" pageNumber="56" type="etymology">
<paragraph id="B64C36C05920AD39C2A2A4D0FAE7FC5B" blockId="5.[151,1436,679,2037]" box="[199,1360,932,957]" pageId="5" pageNumber="56">
<emphasis id="8487EAD25920AD39C2A2A4D0FEF9FC5A" bold="true" box="[199,334,932,957]" pageId="5" pageNumber="56">Etymology.</emphasis>
The species epithet, a latinized adjective, refers to the country in which this species occurs.
</paragraph>
</subSubSection>
<subSubSection id="FEE9654B5920AD39C2A2A4B3FDBBFAAF" pageId="5" pageNumber="56" type="diagnosis">
<paragraph id="B64C36C05920AD39C2A2A4B3FDBBFAAF" blockId="5.[151,1436,679,2037]" pageId="5" pageNumber="56">
<emphasis id="8487EAD25920AD39C2A2A4B3FEF6FC07" bold="true" box="[199,321,967,992]" pageId="5" pageNumber="56">Diagnosis.</emphasis>
Males of the new species are similar to those of
<taxonomicName id="71F34D435920AD39C1E6A4BDFB41FC07" authority="Schwendinger, 1996" authorityName="Schwendinger" authorityYear="1996" box="[899,1270,967,992]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="thoranie">
<emphasis id="8487EAD25920AD39C1E6A4BDFBB5FC07" box="[899,1026,968,992]" italics="true" pageId="5" pageNumber="56">L. thoranie</emphasis>
Schwendinger, 1996
</taxonomicName>
, differing in a relatively wider ventral sternal surface with a heart-shaped posterior part (
<figureCitation id="2EC82A455920AD39C1BDA498FBABFBE3" box="[984,1052,1004,1029]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1896]" captionTargetBox="[161,1429,894,1844]" captionTargetId="figure@3.[151,1436,877,1852]" captionTargetPageId="3" captionText="FIGURE 2. Liphistius isan, male (A, F) and female (B) from Phou Asa, Laos; L. laoticus sp. n., female allotype (C), male holotype (D); L. thoranie, male holotype (E). A B, habitus. C F, sternum and labium, ventral view." httpUri="https://zenodo.org/record/284546/files/figure.png" pageId="5" pageNumber="56">Fig. 2</figureCitation>
D, cf.
<figureCitation id="2EC82A455920AD39C607A498FB12FBE3" box="[1122,1189,1004,1029]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1896]" captionTargetBox="[161,1429,894,1844]" captionTargetId="figure@3.[151,1436,877,1852]" captionTargetPageId="3" captionText="FIGURE 2. Liphistius isan, male (A, F) and female (B) from Phou Asa, Laos; L. laoticus sp. n., female allotype (C), male holotype (D); L. thoranie, male holotype (E). A B, habitus. C F, sternum and labium, ventral view." httpUri="https://zenodo.org/record/284546/files/figure.png" pageId="5" pageNumber="56">Fig. 2</figureCitation>
E), by palp (in ventral view) with a basally wider retrolateral tibial apophysis (
<figureCitation id="2EC82A455920AD39C17EA37BFCD6FBCF" box="[795,865,1039,1064]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">Fig. 4</figureCitation>
A, cf. Schwendinger 1996: fig. 48) carrying in its ventral part two long and one shorter apical spines that are continuously tapering towards the tip, and in its dorsal part a very short apical spine that is thick at base and abruptly tapering to a narrow apex in its distal third (
<figureCitation id="2EC82A455920AD39C723A323FA3DFB97" box="[1350,1418,1111,1136]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">
Fig.
<date id="C24D10005920AD39C71EA32CFA3DFB97" box="[1403,1418,1112,1136]" pageId="5" pageNumber="56">4</date>
</figureCitation>
I) (in
<taxonomicName id="71F34D435920AD39C2A0A309FEFDFB73" box="[197,330,1148,1172]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="thoranie">
<emphasis id="8487EAD25920AD39C2A0A309FEFDFB73" box="[197,330,1148,1172]" italics="true" pageId="5" pageNumber="56">L. thoranie</emphasis>
</taxonomicName>
all apical spines are abruptly tapering; two of them are broken off in the illustration by Schwendinger 1996: figs 4648); the cumulus is slightly more elevated, its thick bristles in a less compact group; the retrolateral heel of the paracymbium is less pronounced (
<figureCitation id="2EC82A455920AD39C15EA3B0FCC9FB3B" box="[827,894,1220,1245]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">Fig. 4</figureCitation>
J, cf.
<figureCitation id="2EC82A455920AD39C1DCA3B0FC4BFB3B" box="[953,1020,1220,1245]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">Fig. 4</figureCitation>
K); the paraembolic plate is narrower (
<figureCitation id="2EC82A455920AD39C2FAA393FF54FAE7" box="[159,227,1255,1280]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">Fig. 4</figureCitation>
D, cf.
<figureCitation id="2EC82A455920AD39C34DA393FEDCFAE7" box="[296,363,1255,1280]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">Fig. 4</figureCitation>
M). Females of the new species are very similar to those of
<taxonomicName id="71F34D435920AD39C66CA39DFF65FAC3" authority="Ono &amp; Schwendinger, 1990" authorityName="Ono &amp; Schwendinger" authorityYear="1990" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="ochraceus">
<emphasis id="8487EAD25920AD39C66CA39DFB2DFAE7" box="[1033,1178,1256,1280]" italics="true" pageId="5" pageNumber="56">L. ochraceus</emphasis>
Ono &amp; Schwendinger, 1990
</taxonomicName>
, differing in a poreplate with a slightly longer and posteriorly wider posterior stalk (
<figureCitation id="2EC82A455920AD39C68CA278FA84FAC3" box="[1257,1331,1292,1317]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="5" pageNumber="56">Fig. 4</figureCitation>
NQ, cf. Schwendinger 1996: figs 8082).
</paragraph>
</subSubSection>
<subSubSection id="FEE9654B5920AD3BC2A2A220FE7AF8AA" lastPageId="7" lastPageNumber="58" pageId="5" pageNumber="56" type="description">
<paragraph id="B64C36C05920AD39C2A2A220FB0EF9A2" blockId="5.[151,1436,679,2037]" pageId="5" pageNumber="56">
<emphasis id="8487EAD25920AD39C2A2A220FEEEFA8A" bold="true" box="[199,345,1364,1389]" pageId="5" pageNumber="56">Description.</emphasis>
MALE (
<typeStatus id="694888625920AD39C3ADA220FD87FA8A" box="[456,560,1364,1389]" pageId="5" pageNumber="56" type="holotype">holotype</typeStatus>
). Colour in alcohol: Most of body light brown (dark brown when alive). Carapace with light longitudinal area behind eye tubercle and indistinct light patches in posterior part of pars thoracica. Chelicerae light grey-brown distally, cream-coloured proximally. Femora and patellae of legs and palps speckled with grey-brown (speckles confluent on dorsal side of palpal femora); no dark annulations on tibiae to tarsi. Cymbium mostly reddish brown except for a light distal zone. Two anteriormost opisthosomal tergites uniformly medium brown; tergites IIIVI with increasingly larger, light brown, longitudinal areas separating three medium brown areas; on tergites VIIX medium brown areas becoming more extensive again.
</paragraph>
<paragraph id="B64C36C05920AD39C2A2A13BFD89F812" blockId="5.[151,1436,679,2037]" pageId="5" pageNumber="56">
Total length 14.1. Carapace 6.0 long, 5.2 wide, set with quite few short, blunt-tipped hairs scattered in peripheral areas (not in central area). Ocular tubercle 0.91 long, 0.99 wide. Eye sizes and interdistances: AME 0.04, ALE 0.61, PME 0.33, PLE 0.50; AMEAME 0.13, AMEALE 0.12, PMEPME 0.06, PMEPLE 0.05, ALEPLE 0.03. MOQ 0.38 long, front width 0.21, back width 0.58. Labium 0.7 long, 1.1 wide. Sternum 2.5 long, 2.0 wide (1.1 on ventral surface); posterior part of ventral surface heart-shaped, clearly delimited and set with longer and stronger setae than indistinctly delimited anterior part; a light central spot present between both parts (
<figureCitation id="2EC82A455920AD39C756A070FACFF8FB" box="[1331,1400,1796,1821]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1896]" captionTargetBox="[161,1429,894,1844]" captionTargetId="figure@3.[151,1436,877,1852]" captionTargetPageId="3" captionText="FIGURE 2. Liphistius isan, male (A, F) and female (B) from Phou Asa, Laos; L. laoticus sp. n., female allotype (C), male holotype (D); L. thoranie, male holotype (E). A B, habitus. C F, sternum and labium, ventral view." httpUri="https://zenodo.org/record/284546/files/figure.png" pageId="5" pageNumber="56">Fig. 2</figureCitation>
D). Border between labium and sternum indistinct (without a distinct suture as in females). Maxillae 2.0 long, 1.3 wide. Promargin of cheliceral groove with 10 small teeth on left side and 11 on right side. Paired tarsal claws of first leg with four teeth, those of other legs with 45 teeth; unpaired claw with two tiny denticles on legs II and IV, three on legs I and III. Lengths of limbs: palp 9.8 (2.9 + 1.8 + 3.4 + 1.7); leg I 16.9 (4.7 + 2.3 + 3.5 + 4.3 + 2.1); leg II 18.2 (4.9 + 2.4 + 3.7 + 4.8 + 2.4); leg III 20.3 (5.0 + 2.4 + 4.1 + 6.0 + 2.8); leg IV 26.1 (6.1 + 2.6 + 5.4 + 8.1 + 3.9). Opisthosoma 5.7 long, 3.6 wide.
</paragraph>
<caption id="E28C66485923AD3AC2F2A13FFA36F93B" httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="6" pageNumber="57" targetBox="[162,1418,206,1560]" targetPageId="6">
<paragraph id="B64C36C05923AD3AC2F2A13FFA36F93B" blockId="6.[151,1436,1611,1756]" pageId="6" pageNumber="57">
<emphasis id="8487EAD25923AD3AC2F2A13FFEA5F987" bold="true" box="[151,274,1611,1633]" pageId="6" pageNumber="57">FIGURE 4.</emphasis>
<taxonomicName id="71F34D435923AD3AC37DA13FFE65F986" box="[280,466,1611,1633]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="laoticus" status="sp. nov.">
<emphasis id="8487EAD25923AD3AC37DA13FFE65F986" box="[280,466,1611,1633]" italics="true" pageId="6" pageNumber="57">Liphistius laoticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FB457A95923AD3AC3BDA138FDA6F986" box="[472,529,1612,1633]" pageId="6" pageNumber="57" rank="species">
<emphasis id="8487EAD25923AD3AC3BDA138FDA6F986" bold="true" box="[472,529,1612,1633]" pageId="6" pageNumber="57">sp. n.</emphasis>
</taxonomicNameLabel>
, male holotype (AE, IJ), three male paratypes (FH), four female paratypes (NQ; N is the “allotype”);
<taxonomicName id="71F34D435923AD3AC35EA11FFE1BF967" box="[315,428,1642,1664]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="thoranie">
<emphasis id="8487EAD25923AD3AC35EA11FFE1BF967" box="[315,428,1642,1664]" italics="true" pageId="6" pageNumber="57">L. thoranie</emphasis>
</taxonomicName>
, male paratype (illustrated in original description; K), holotype (M);
<taxonomicName id="71F34D435923AD3AC60AA11FFB44F967" box="[1135,1267,1642,1664]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="57" phylum="Arthropoda" rank="species" species="ochraceus">
<emphasis id="8487EAD25923AD3AC60AA11FFB44F967" box="[1135,1267,1642,1664]" italics="true" pageId="6" pageNumber="57">L. ochraceus</emphasis>
</taxonomicName>
, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. EH, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. JL, paracymbium, dorsal view. NQ, vulva, ventral view. Scale lines 0.5 mm (AD, M; EH, JL; I; NQ).
</paragraph>
</caption>
<paragraph id="B64C36C05923AD3BC2A2A07CFB9EFEA7" blockId="6.[151,1437,1800,2006]" lastBlockId="7.[151,1437,151,2013]" lastPageId="7" lastPageNumber="58" pageId="6" pageNumber="57">
Pedipalps (
<figureCitation id="2EC82A455923AD3AC326A07CFE31F8C6" box="[323,390,1800,1825]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="6" pageNumber="57">Fig. 4</figureCitation>
AE, IJ) with retrolateral tibial apophysis carrying four strong apical spines, the ventral two of them long and continuously tapering to tip, the third one shorter and also continuously tapering, the dorsal one very short and abruptly tapering (
<figureCitation id="2EC82A455923AD3AC049A024FDC0F88E" box="[556,631,1872,1897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="6" pageNumber="57">
Fig.
<date id="C24D10005923AD3AC002A025FDC0F88E" box="[615,631,1873,1897]" pageId="6" pageNumber="57">4</date>
</figureCitation>
I); other spines on tibial apophysis subapical and distinctly weaker. Paracymbium medium-sized, with a distinctly projecting retrolateral-proximal heel (
<figureCitation id="2EC82A455923AD3AC631A000FB2EF86A" box="[1108,1177,1908,1933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="6" pageNumber="57">Fig. 4</figureCitation>
A, J); cumulus slightly elevated, carrying a relatively dispersed group of long thick bristles. Subtegulum without apophysis. Contrategulum with sharp dorsodistal edge rising to rounded angle; dorsal surface of contrategulum without proximal ledge; ventral contrategular process relatively large, with narrowly rounded apex (
<figureCitation id="2EC82A455922AD3BC6A3A7E3FABAFF57" box="[1222,1293,151,176]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
E). Tegulum relatively large, its proximal edge coarsely dentate and widely arched, its distal edge distinct, narrow, somewhat truncate, on its retrolateral side separated by a narrow unpigmented zone from contrategulum (
<figureCitation id="2EC82A455922AD3BC6B5A7ABFAA2FF1F" box="[1232,1301,223,248]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
BC). Paraembolic plate relatively narrow, with a narrowly rounded apex (
<figureCitation id="2EC82A455922AD3BC108A670FC05FEFB" box="[877,946,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
D); divided embolus fairly short, only little surpassing paraembolic plate and dorsodistal edge of contrategulum (
<figureCitation id="2EC82A455922AD3BC1FAA653FC54FEA7" box="[927,995,295,320]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
AD).
</paragraph>
<paragraph id="B64C36C05922AD3BC2A2A638FCB4FDFF" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">FEMALE (“allotype”). Colour as in male, but with more extensive light patches on carapace almost completely encircling a darker (as the periphery), indistinctly flower-shaped central area around fovea. Light brown areas on all opisthosomal tergites, these everywhere more extensive than in male. Indistinct, medium brown, proximal and subdistal annulations on leg tibiae and tarsi and on palpal tibiae, subproximal annulations on metatarsi of legs and palps; palpal tarsi uniformly medium brown. Coxae of legs III and of palps, and ventral side of chelicerae light reddish brown (darker than in male).</paragraph>
<paragraph id="B64C36C05922AD3BC2A2A550FC35FC42" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
Total length 17.0 mm. Carapace 6.6 long, 5.4 wide. Ocular tubercle 0.91 long, 0.99 wide. Eye sizes and interdistances: AME 0.04, ALE 0.60, PME 0.30, PLE 0.44; AMEAME 0.13, AMEALE 0.11, PMEPME 0.08, PMEPLE 0.08, ALEPLE 0.10. MOQ 0.46 long, front width 0.24, back width 0.56. Labium 0.9 long, 2.3 wide. Sternum 4.1 long, 2.8 wide (2.0 on ventral surface), not divided into two unequal parts as in male. Border between labium and sternum clearly marked by deep suture (
<figureCitation id="2EC82A455922AD3BC084A5C0FC93FD2B" box="[737,804,692,717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1874,1896]" captionTargetBox="[161,1429,894,1844]" captionTargetId="figure@3.[151,1436,877,1852]" captionTargetPageId="3" captionText="FIGURE 2. Liphistius isan, male (A, F) and female (B) from Phou Asa, Laos; L. laoticus sp. n., female allotype (C), male holotype (D); L. thoranie, male holotype (E). A B, habitus. C F, sternum and labium, ventral view." httpUri="https://zenodo.org/record/284546/files/figure.png" pageId="7" pageNumber="58">Fig. 2</figureCitation>
C). Maxillae 2.3 long, 1.5 wide. Promargin of cheliceral groove with 10 teeth on left side, 11 on right side. Paired tarsal claws of legs with 34 teeth; unpaired claw with four denticles on legs III, three/five denticles on leg III, and four/six denticles on leg IV; pedipalpal claws with three tiny denticles. Hairs spread over leg coxae and labium, and hairs in distal part of maxillae and sternum being distinctly longer and stronger than those in male. Lengths of limbs: palp 10.7 (3.4 + 2.0 + 2.6 + 2.7); leg I 13.1 (4.0 + 2.2 + 2.7 + 2.7 + 1.5); leg II 14.5 (4.3 + 2.4 + 2.9 + 3.1 + 1.8); leg III 15.6 (4.5 + 2.4 + 3.0 + 3.7 + 2.0); leg IV 21.6 (5.7 + 2.8 + 4.4 + 5.9 + 2.8). Opisthosoma 7.6 long, 5.4 wide.
</paragraph>
<paragraph id="B64C36C05922AD3BC2A2A4DBFE95FBBF" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
Vulva (
<figureCitation id="2EC82A455922AD3BC37AA4DBFEDFFC2F" box="[287,360,943,968]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
N). Poreplate simple, widely quadrangular, with moderately thick anterior and lateral rims without protuberances; a characteristic unpigmented area between each short posterior margin (without thickened rim) of poreplate and base of posterior stalk; receptacular cluster relatively simple, almost digitiform; posterior stalk medium-long, posteriorly wider than anteriorly, with widely rounded posterior margin. Genital atrium without lateral hairs.
</paragraph>
<paragraph id="B64C36C05922AD3BC2A2A310FE93FAB3" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
<emphasis id="8487EAD25922AD3BC2A2A310FE88FB9A" bold="true" box="[199,319,1124,1149]" pageId="7" pageNumber="58">Variation.</emphasis>
One male
<typeStatus id="694888625922AD3BC3D8A311FDAAFB9A" box="[445,541,1125,1149]" pageId="7" pageNumber="58" type="paratype">paratype</typeStatus>
from the
<typeStatus id="694888625922AD3BC0F5A311FD77FB9A" box="[656,704,1125,1149]" pageId="7" pageNumber="58">type</typeStatus>
locality lacks its right AME. The characteristic light spot in the centre of the sternum is difficult to see in the male most strongly bleached by alcohol preservation. One female lacks its right anterior median spinneret, having only a pigmented spot with bristles where the base of the spinneret should be. Variation in the shape of the contrategular processes of four males is shown in
<figureCitation id="2EC82A455922AD3BC6F3A3BBFB6FFB0F" box="[1174,1240,1231,1256]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
EH, variation in the shape of the poreplates of four females in
<figureCitation id="2EC82A455922AD3BC0F7A380FD64FAEB" box="[658,723,1268,1293]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
NQ. The “allotype” has no hairs in its genital atrium (
<figureCitation id="2EC82A455922AD3BC753A380FACFFAEB" box="[1334,1400,1268,1293]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
N), one female
<typeStatus id="694888625922AD3BC37EA26CFECBFAD7" box="[283,380,1304,1328]" pageId="7" pageNumber="58" type="paratype">paratype</typeStatus>
has two hairs on one side only (
<figureCitation id="2EC82A455922AD3BC08EA263FC98FAD7" box="[747,815,1303,1328]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
Q), the two others have up to seven hairs on both sides (
<figureCitation id="2EC82A455922AD3BC2FAA248FF54FAB3" box="[159,227,1340,1365]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
<collectingCountry id="CEE476505922AD3BC287A248FF40FAB3" box="[226,247,1340,1364]" name="Iceland" pageId="7" pageNumber="58">O</collectingCountry>
P).
</paragraph>
<paragraph id="B64C36C05922AD3BC2A2A215FBADFA7A" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">Measurements of males (n = 7) range: body length 11.514.0, carapace length 4.85.9, carapace width 4.15.2. The corresponding maximal measurements in females (n = 7) are: 18.0, 6.5, 5.5.</paragraph>
<paragraph id="B64C36C05922AD3BC2A2A2D3FE7AF8AA" blockId="7.[151,1437,151,2013]" pageId="7" pageNumber="58">
<emphasis id="8487EAD25922AD3BC2A2A2D3FEC7FA27" bold="true" box="[199,368,1447,1472]" pageId="7" pageNumber="58">Relationships.</emphasis>
<taxonomicName id="71F34D435922AD3BC313A2DCFDF3FA27" box="[374,580,1448,1472]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="laoticus" status="sp. nov.">
<emphasis id="8487EAD25922AD3BC313A2DCFDF3FA27" box="[374,580,1448,1472]" italics="true" pageId="7" pageNumber="58">Liphistius laoticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FB457A95922AD3BC02FA2D3FD3EFA27" box="[586,649,1447,1472]" pageId="7" pageNumber="58" rank="species">
<emphasis id="8487EAD25922AD3BC02FA2D3FD3EFA27" bold="true" box="[586,649,1447,1472]" pageId="7" pageNumber="58">sp. n.</emphasis>
</taxonomicNameLabel>
is a member of the
<taxonomicName id="71F34D435922AD3BC101A2DDFC17FA27" box="[868,928,1449,1472]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="trang">
<emphasis id="8487EAD25922AD3BC101A2DDFC17FA27" box="[868,928,1449,1472]" italics="true" pageId="7" pageNumber="58">trang</emphasis>
</taxonomicName>
-species group, as are all
<taxonomicName id="71F34D435922AD3BC6DFA2DCFA9EFA27" box="[1210,1321,1448,1472]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="8487EAD25922AD3BC6DFA2DCFA9EFA27" box="[1210,1321,1448,1472]" italics="true" pageId="7" pageNumber="58">Liphistius</emphasis>
</taxonomicName>
species in nearby northeastern
<collectingCountry id="CEE476505922AD3BC318A2B8FE53FA02" box="[381,484,1484,1509]" name="Thailand" pageId="7" pageNumber="58">Thailand</collectingCountry>
. Medium size, quite uniformly brown body colouration, and similarities in male and female copulatory organs (e.g. long and thick bristles on slightly or moderately elevated cumulus; distinct contrategular process; simple, more or less digitiform receptacular cluster) identify the new species as belonging to species complex B (Schwendinger 1996), later inappropriately called superspecies B (Schwendinger 1998), a subgroup which includes also
<taxonomicName id="71F34D435922AD3BC382A129FCA8F993" authority="Schwendinger, L." authorityName="Schwendinger, L." box="[487,799,1628,1653]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="erawan">
<emphasis id="8487EAD25922AD3BC382A129FDEFF993" box="[487,600,1629,1652]" italics="true" pageId="7" pageNumber="58">L. erawan</emphasis>
Schwendinger,
<emphasis id="8487EAD25922AD3BC16CA129FCA8F993" box="[777,799,1629,1652]" italics="true" pageId="7" pageNumber="58">L.</emphasis>
</taxonomicName>
tham Sedgwick &amp; Schwendinger,
<emphasis id="8487EAD25922AD3BC6FAA129FA2BF993" box="[1183,1436,1628,1652]" italics="true" pageId="7" pageNumber="58">
<taxonomicName id="71F34D435922AD3BC6FAA129FAA9F993" box="[1183,1310,1628,1652]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="thoranie">L. thoranie</taxonomicName>
,
<taxonomicName id="71F34D435922AD3BC74DA129FA2BF993" authority="Schwendinger, L." authorityName="Schwendinger, L." box="[1320,1436,1628,1652]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="pusohm">L. pusohm</taxonomicName>
</emphasis>
Schwendinger,
<taxonomicName id="71F34D435922AD3BC326A1F5FD8CF97F" authority="Schwendinger" authorityName="Schwendinger" box="[323,571,1663,1688]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="onoi">
<emphasis id="8487EAD25922AD3BC326A1F5FE24F970" box="[323,403,1664,1688]" italics="true" pageId="7" pageNumber="58">L. onoi</emphasis>
Schwendinger
</taxonomicName>
and
<taxonomicName id="71F34D435922AD3BC011A1F5FCB2F97F" box="[628,773,1664,1688]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="ochraceus">
<emphasis id="8487EAD25922AD3BC011A1F5FCB2F97F" box="[628,773,1664,1688]" italics="true" pageId="7" pageNumber="58">L. ochraceus</emphasis>
</taxonomicName>
.
<taxonomicName id="71F34D435922AD3BC171A1F4FC4AF97F" box="[788,1021,1664,1688]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="ochraceus">
<emphasis id="8487EAD25922AD3BC171A1F4FC4AF97F" box="[788,1021,1664,1688]" italics="true" pageId="7" pageNumber="58">Liphistius ochraceus</emphasis>
</taxonomicName>
(especially females which also have modifications at the posterior margins of the poreplate, Schwendinger 1996: figs 8082 cf.
<figureCitation id="2EC82A455922AD3BC6C2A1D0FB5EF95B" box="[1191,1257,1700,1725]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
NQ; males are without a retrolateral-proximal heel on the paracymbium,
<figureCitation id="2EC82A455922AD3BC157A1B3FCC2F907" box="[818,885,1735,1760]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
L) and
<taxonomicName id="71F34D435922AD3BC1AEA1BDFBFDF907" box="[971,1098,1736,1760]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="thoranie">
<emphasis id="8487EAD25922AD3BC1AEA1BDFBFDF907" box="[971,1098,1736,1760]" italics="true" pageId="7" pageNumber="58">L. thoranie</emphasis>
</taxonomicName>
(especially males which also have a distinct retrolateral-proximal heel on their paracymbium,
<figureCitation id="2EC82A455922AD3BC111A198FC01F8E3" box="[884,950,1772,1797]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
K cf.
<figureCitation id="2EC82A455922AD3BC19DA198FB8CF8E3" box="[1016,1083,1772,1797]" captionStart="FIGURE 4" captionStartId="6.[151,250,1611,1633]" captionTargetBox="[162,1418,206,1560]" captionTargetId="figure@6.[151,1436,193,1590]" captionTargetPageId="6" captionText="FIGURE 4. Liphistius laoticus sp. n., male holotype (A E, I J), three male paratypes (F H), four female paratypes (N Q; N is the “ allotype ”); L. thoranie, male paratype (illustrated in original description; K), holotype (M); L. ochraceus, male from type locality (L). A, distal part of left palp, ventral view. B, same, retrolateral view. C, same, retrodorsal view. D, M, distal part of palpal organ, slightly different ventral view. E H, palpal organ, distal view. I, tibial apophysis, retrolateral and slightly proximal view. J L, paracymbium, dorsal view. N Q, vulva, ventral view. Scale lines 0.5 mm (A D, M; E H, J L; I; N Q)." httpUri="https://zenodo.org/record/284548/files/figure.png" pageId="7" pageNumber="58">Fig. 4</figureCitation>
A, J) are morphologically most similar to the new species.
<taxonomicName id="71F34D435922AD3BC3BEA064FD03F8CF" box="[475,692,1808,1832]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="thoranie">
<emphasis id="8487EAD25922AD3BC3BEA064FD03F8CF" box="[475,692,1808,1832]" italics="true" pageId="7" pageNumber="58">Liphistius thoranie</emphasis>
</taxonomicName>
is geographically closer and thus regarded as the closest know relative of
<taxonomicName id="71F34D435922AD3BC375A041FE30F8AB" box="[272,391,1844,1868]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="laoticus" status="sp. nov.">
<emphasis id="8487EAD25922AD3BC375A041FE30F8AB" box="[272,391,1844,1868]" italics="true" pageId="7" pageNumber="58">L. laoticus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9FB457A95922AD3BC3E8A040FE7AF8AA" box="[397,461,1844,1869]" pageId="7" pageNumber="58" rank="species">
<emphasis id="8487EAD25922AD3BC3E8A040FE7AF8AA" bold="true" box="[397,461,1844,1869]" pageId="7" pageNumber="58">sp. n.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="FEE9654B5922AD34C2A2A023FF5AFF32" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58" type="distribution">
<paragraph id="B64C36C05922AD34C2A2A023FF5AFF32" blockId="7.[151,1437,151,2013]" lastBlockId="8.[151,1436,151,752]" lastPageId="8" lastPageNumber="59" pageId="7" pageNumber="58">
<emphasis id="8487EAD25922AD3BC2A2A023FE40F897" bold="true" box="[199,503,1879,1904]" pageId="7" pageNumber="58">Distribution and habitat.</emphasis>
The
<typeStatus id="694888625922AD3BC050A02DFDC7F896" box="[565,624,1881,1905]" pageId="7" pageNumber="58">types</typeStatus>
of the new species were collected from two localities on the Bolaven Plateau of southern
<collectingCountry id="CEE476505922AD3BC31DA009FE05F873" box="[376,434,1917,1940]" name="Laos" pageId="7" pageNumber="58">Laos</collectingCountry>
. Siegfried Huber (pers. comm.) has recently found
<taxonomicName id="71F34D435922AD3BC191A008FBD5F873" box="[1012,1122,1916,1940]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="8487EAD25922AD3BC191A008FBD5F873" box="[1012,1122,1916,1940]" italics="true" pageId="7" pageNumber="58">Liphistius</emphasis>
</taxonomicName>
(probably conspecific) also at the nearby Tad Itou. All spiders were found on earthbanks near waterfalls (
<taxonomicName id="71F34D435922AD3BC196A0D4FB81F85E" box="[1011,1078,1952,1977]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="58" phylum="Arthropoda" rank="species" species="tham">Tham</taxonomicName>
Champee is actually a waterfall and not a cave as the Laotian name indicates). These localities probably receive considerably more humidity during the dry season than other areas on the plateau, most of which is under some kind of cultivation (predominantly coffee).
</paragraph>
</subSubSection>
<subSubSection id="FEE9654B592DAD34C2A2A7ABFE0AFD17" pageId="8" pageNumber="59" type="biology_ecology">
<paragraph id="B64C36C0592DAD34C2A2A7ABFC31FE37" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="59">
<emphasis id="8487EAD2592DAD34C2A2A7ABFE90FF1F" bold="true" box="[199,295,223,248]" pageId="8" pageNumber="59">Biology.</emphasis>
The six males from the
<typeStatus id="69488862592DAD34C05AA794FDD8FF1F" box="[575,623,224,248]" pageId="8" pageNumber="59">type</typeStatus>
locality became mature between the end of July and the end of August (
<date id="C24D1000592DAD34C2FAA670FF43FEFB" box="[159,244,260,284]" pageId="8" pageNumber="59" value="1997-07">30.VII.</date>
,
<date id="C24D1000592DAD34C361A670FEEFFEFB" box="[260,344,260,284]" pageId="8" pageNumber="59" value="1997-07">31.VII.</date>
,
<date id="C24D1000592DAD34C30CA671FE0FFEFB" box="[361,440,261,284]" pageId="8" pageNumber="59" value="1997-08">7.VIII.</date>
,
<date id="C24D1000592DAD34C3ACA670FDA0FEFB" box="[457,535,260,284]" pageId="8" pageNumber="59" value="1997-08">8.VIII.</date>
,
<date id="C24D1000592DAD34C04CA670FDC0FEFB" box="[553,631,260,284]" pageId="8" pageNumber="59" value="1997-08">9.VIII.</date>
,
<date id="C24D1000592DAD34C0EDA670FCA9FEFB" box="[648,798,260,284]" pageId="8" pageNumber="59" value="1997-08-27">27.VIII.1997</date>
), after 78 months in captivity; the male from
<taxonomicName id="71F34D43592DAD34C73EA670FA2BFEFA" box="[1371,1436,260,285]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="58" phylum="Arthropoda" rank="species" species="tham">Tham</taxonomicName>
Champee matured in early October (
<date id="C24D1000592DAD34C054A65CFD29FEA7" box="[561,670,296,320]" pageId="8" pageNumber="59" value="2010-10-09">9.X.2010</date>
), only a week after being captured. Females with egg cases moulted between late December and early January, one of them again in May and August. Three egg cases were found in the field in late December; two others were constructed in captivity in late November. These were
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wide and
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high, and contained 4588 cream-coloured or light yellow eggs wrapped in a thin membrane and suspended above the bottom of the egg case with very fine threads.
</paragraph>
<paragraph id="B64C36C0592DAD34C2A2A6A9FAF6FD87" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="59">
Burrows of three juvenile males were T-shaped (with two trapdoors), all other burrows were unbranched and closed by a single trapdoor with its hinge on the upper side or slightly laterally. In penultimate males the doors were
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wide and 1.0
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long; in females up to
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wide and up to
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long. Six to eight signal lines, up to
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long, were spread over the soil surface or (rarely) stones. The largest burrow was
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long.
</paragraph>
<paragraph id="B64C36C0592DAD34C2A2A518FE0AFD17" blockId="8.[151,1436,151,752]" pageId="8" pageNumber="59">
One female (from
<taxonomicName id="71F34D43592DAD34C3FFA518FE6BFD62" box="[410,476,620,645]" class="Arachnida" family="Liphistiidae" genus="Liphistius" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="58" phylum="Arthropoda" rank="species" species="tham">Tham</taxonomicName>
Champee; with egg case) and one male (from Tad Fane; matured
<date id="C24D1000592DAD34C6AAA518FAE2FD63" box="[1231,1365,620,644]" pageId="8" pageNumber="59" value="1997-08-07">7.VIII.1997</date>
) were carrying ectoparasitic mites of the laelapid genus
<taxonomicName id="71F34D43592DAD34C08CA5E4FCF9FD4F" box="[745,846,656,680]" class="Arachnida" family="Laelapidae" genus="Ljunghia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="8487EAD2592DAD34C08CA5E4FCF9FD4F" box="[745,846,656,680]" italics="true" pageId="8" pageNumber="59">Ljunghia</emphasis>
</taxonomicName>
. The mites left bite marks (as illustrated for two
<taxonomicName id="71F34D43592DAD34C2F2A5C0FEA3FD2B" box="[151,276,692,716]" class="Arachnida" family="Liphistiidae" genus="Heptathela" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="59" phylum="Arthropoda" rank="genus">
<emphasis id="8487EAD2592DAD34C2F2A5C0FEA3FD2B" box="[151,276,692,716]" italics="true" pageId="8" pageNumber="59">Heptathela</emphasis>
</taxonomicName>
species in Schwendinger &amp; Ono 2011: figs 6768) on the carapace and on the proximal part of the chelicerae of both spiders.
</paragraph>
</subSubSection>
</treatment>
</document>