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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="3A7F611E2AAD5328931F180198BE83A1" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria davidcauseyi Causey 1950" docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
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<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Marek, Paul E.</mods:namePart>
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<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
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<mods:number>1030</mods:number>
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<mods:start>1</mods:start>
<mods:end>180</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
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<treatment ID-GBIF-Taxon="181004749" LSID="urn:lsid:plazi:treatment:3A7F611E2AAD5328931F180198BE83A1" httpUri="http://treatment.plazi.org/id/3A7F611E2AAD5328931F180198BE83A1" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="45M4J" LSID="3A7F611E-2AAD-5328-931F-180198BE83A1" authority="Causey, 1950" authorityName="Causey" authorityYear="1950" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria davidcauseyi" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="davidcauseyi">Nannaria davidcauseyi Causey, 1950</taxonomicName>
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Nannaria davidcauseyi Causey, 1950 ♂ (VMNH, NAN 0537) left gonopod A anterior view; red triangle indicates small acropodite medial flange, red arrow indicates medial flange at apex B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure34" httpUri="https://binary.pensoft.net/fig/531137" pageId="0" pageNumber="1">
Fig. 34 Vernacular name: &quot;David
<normalizedToken originalValue="Causeys">Causey's</normalizedToken>
Twisted-Claw Millipede&quot;
</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Causey" authorityYear="1950" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria davidcauseyi" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="davidcauseyi">Nannaria davidcauseyi</taxonomicName>
Causey, 1950b: 194, figs 3, 4.
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Virginia Museum of Natural History Special Publication Number" pageId="0" pageNumber="1" pagination="1 - 584" refId="B33" refString="Hoffman, RL, 1999. Checklist of the millipeds of North and Middle America. Virginia Museum of Natural History Special Publication Number 8: 1 - 584" title="Checklist of the millipeds of North and Middle America." volume="8" year="1999">Hoffman 1999</bibRefCitation>
: 366.
<bibRefCitation author="Marek, PE" journalOrPublisher="Virginia Museum of Natural History Publications" pageId="0" pageNumber="1" pagination="1 - 117" refId="B46" refString="Marek, PE, Tanabe, T, Sierwald, P, 2014. A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida). Virginia Museum of Natural History Publications 17: 1 - 117" title="A species catalog of the millipede family Xystodesmidae (Diplopoda: Polydesmida)." volume="17" year="2014">Marek et al. 2014</bibRefCitation>
: 36.
<bibRefCitation DOI="https://doi.org/10.1093/isd/ixab003" author="Means, JC" journalOrPublisher="Insect Systematics and Diversity" pageId="0" pageNumber="1" pagination="1 - 26" refId="B52" refString="Means, JC, Hennen, DA, Tanabe, T, Marek, PE, 2021. Phylogenetic systematics of the millipede family Xystodesmidae. Insect Systematics and Diversity 5: 1 - 26, DOI: https://doi.org/10.1093/isd/ixab003" title="Phylogenetic systematics of the millipede family Xystodesmidae." url="https://doi.org/10.1093/isd/ixab003" volume="5" year="2021">Means et al. 2021</bibRefCitation>
: S69.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Diplopoda" family="Xystodesmidae" genus="Mimuloria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mimuloria davidcauseyi" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="davidcauseyi">Mimuloria davidcauseyi</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.5479/si.03629236.212" author="Chamberlin, RV" journalOrPublisher="United States National Museum Bulletin" pageId="0" pageNumber="1" pagination="1 - 236" refId="B22" refString="Chamberlin, RV, Hoffman, RL, 1958. Checklist of the millipeds of North America. United States National Museum Bulletin 212: 1 - 236, DOI: https://doi.org/10.5479/si.03629236.212" title="Checklist of the millipeds of North America." url="https://doi.org/10.5479/si.03629236.212" volume="212" year="1958">Chamberlin and Hoffman 1958</bibRefCitation>
: 37.
<bibRefCitation author="Hennen, DA" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 21" publicationUrl="http://digitalcommons.unl.edu/insectamundi/924" refId="B28" refString="Hennen, DA, Shelley, RM, 2015. A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964. Insecta Mundi 0418: 1 - 21, http://digitalcommons.unl.edu/insectamundi/924" title="A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964." url="http://digitalcommons.unl.edu/insectamundi/924" volume="0418" year="2015">Hennen and Shelley 2015</bibRefCitation>
: 10, figs 4-8, 16, 17.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355446" collectingDate="2013-11-02" collectingMethod="pitfall trap" collectorName="M. Skvarla" country="United States" county="Newton County" latitude="35.8506" location="Dismal Hollow" longLatPrecision="6" longitude="-93.2701" specimenCount="1" specimenCount-female="1" stateProvince="Arkansas">
<collectingCountry name="United States of America">United States</collectingCountry>
-
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Arkansas">Arkansas</collectingRegion>
</emphasis>
<specimenCount type="female">1 ♀</specimenCount>
;
<collectingCounty>Newton County</collectingCounty>
,
<location LSID="urn:lsid:plazi:treatment:3A7F611E2AAD5328931F180198BE83A1:C393AA282FD5482A1CF5C63455EE4F75" country="United States" county="Newton County" latitude="35.8506" longLatPrecision="6" longitude="-93.2701" name="Dismal Hollow" stateProvince="Arkansas">Dismal Hollow</location>
;
<geoCoordinate degrees="35.8506" direction="north" orientation="latitude" precision="5" value="35.8506">35.8506°N</geoCoordinate>
, -
<geoCoordinate degrees="93.2701" direction="west" orientation="longitude" precision="5" value="-93.2701">93.2701°W</geoCoordinate>
;
<collectingDate value="2013-11-02">2 Nov. 2013</collectingDate>
;
<collectingMethod>pitfall trap</collectingMethod>
;
<collectorName>M. Skvarla</collectorName>
leg.; NCSM NAN0539
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355325" collectingDate="2013-11-07" collectingMethod="pitfall trap" collectorName="M. Skvarla, For, Suppl." country="United States" county="Newton County" latitude="36.038" location="Buffalo Nt River" longLatPrecision="6" longitude="-93.3402" municipality="Steel Creek" specimenCount="2" specimenCount-male="2" stateProvince="Arkansas">
<specimenCount type="male">2 ♂♂</specimenCount>
;
<collectingCounty>Newton County</collectingCounty>
,
<collectingMunicipality>Steel Creek</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:3A7F611E2AAD5328931F180198BE83A1:6FC299583A43F1C872EC851CAC2A290E" country="United States" county="Newton County" latitude="36.038" longLatPrecision="6" longitude="-93.3402" municipality="Steel Creek" name="Buffalo Nt River" stateProvince="Arkansas">Buffalo Nt River</location>
;
<geoCoordinate degrees="36.0380" direction="north" orientation="latitude" precision="5" value="36.038">36.0380°N</geoCoordinate>
, -
<geoCoordinate degrees="93.3402" direction="west" orientation="longitude" precision="5" value="-93.3402">93.3402°W</geoCoordinate>
;
<collectingDate value="2013-11-07">7 Nov. 2013</collectingDate>
;
<collectingMethod>pitfall trap</collectingMethod>
;
<collectorName>M. Skvarla</collectorName>
leg.; NCSM NAN0537.
<collectorName>For</collectorName>
detailed collection data see
<collectorName>Suppl.</collectorName>
material 7
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. davidcauseyi" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="davidcauseyi">
<emphasis italics="true" pageId="0" pageNumber="1">N. davidcauseyi</emphasis>
</taxonomicName>
are distinct from other
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
and the nearby
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
based on the following combination of characters:
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite gently curving medially, and distal zone with medial and lateral flanges at 90° angle to solenomere, not with medial flange at 135° angle to solenomere as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Nannaria davidcauseyi Causey, 1950 ♂ (VMNH, NAN 0537) left gonopod A anterior view; red triangle indicates small acropodite medial flange, red arrow indicates medial flange at apex B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure34" httpUri="https://binary.pensoft.net/fig/531137" pageId="0" pageNumber="1">34A</figureCitation>
, red arrow). Acropodite with small medial flange near apex, not simple lacking flange as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figure 34" captionStartId="F34" captionText="Figure 34. Nannaria davidcauseyi Causey, 1950 ♂ (VMNH, NAN 0537) left gonopod A anterior view; red triangle indicates small acropodite medial flange, red arrow indicates medial flange at apex B medial view C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure34" httpUri="https://binary.pensoft.net/fig/531137" pageId="0" pageNumber="1">34A</figureCitation>
, red triangle). Telopodite basal zone height 1/2 length of acropodite. Prefemoral process pronounced, acicular, arising distantly from small blunt prefemoral spine, not from top of projected prefemoral spine as in
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. castanea" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="castanea">
<emphasis italics="true" pageId="0" pageNumber="1">N. castanea</emphasis>
</taxonomicName>
.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
<bibRefCitation author="Hennen, DA" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 21" publicationUrl="http://digitalcommons.unl.edu/insectamundi/924" refId="B28" refString="Hennen, DA, Shelley, RM, 2015. A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964. Insecta Mundi 0418: 1 - 21, http://digitalcommons.unl.edu/insectamundi/924" title="A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964." url="http://digitalcommons.unl.edu/insectamundi/924" volume="0418" year="2015">Hennen and Shelley (2015)</bibRefCitation>
described specimens of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. davidcauseyi" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="davidcauseyi">
<emphasis italics="true" pageId="0" pageNumber="1">N. davidcauseyi</emphasis>
</taxonomicName>
as having clear paranota with irregular orange spots internally, and orange/pink collum margins.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure34" httpUri="https://binary.pensoft.net/fig/531137" pageId="0" pageNumber="1" start="Figure 34" startId="F34">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 34.</emphasis>
<taxonomicName authorityName="Causey" authorityYear="1950" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria davidcauseyi" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="davidcauseyi">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria davidcauseyi</emphasis>
</taxonomicName>
Causey, 1950 ♂ (VMNH, NAN0537) left gonopod
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view; red triangle indicates small acropodite medial flange, red arrow indicates medial flange at apex
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ (VMNH, NAN0537): BL = 23.3, CW = 3.0, IW = 1.5, ISW = 0.6, B11W = 3.8, B11H = 2.2.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Causey, NB" journalOrPublisher="Entomological News" pageId="0" pageNumber="1" pagination="193 - 198" refId="B11" refString="Causey, NB, 1950b. On four new polydesmoid millipeds. Entomological News 61: 193 - 198" title="On four new polydesmoid millipeds." volume="61" year="1950 b">Causey (1950b</bibRefCitation>
: 194) mentions that the male paratype of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. davidcauseyi" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="davidcauseyi">
<emphasis italics="true" pageId="0" pageNumber="1">N. davidcauseyi</emphasis>
</taxonomicName>
has variation in the shape of both the prefemoral spine and the &quot;keel at the end of the main blade&quot; referring to the caudally-directed acropodite tip.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Known only from the northwest corner of Arkansas (Arkansas: Johnson and Newton counties, Fig.
<figureCitation captionStart="Figure 129" captionStartId="F129" captionText="Figure 129. Distribution map of Nannaria minor species group collection localities west of the Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure129" httpUri="https://binary.pensoft.net/fig/531232" pageId="0" pageNumber="1">129</figureCitation>
). Distribution area: 131 km2; status: MRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Causey, NB" journalOrPublisher="Entomological News" pageId="0" pageNumber="1" pagination="193 - 198" refId="B11" refString="Causey, NB, 1950b. On four new polydesmoid millipeds. Entomological News 61: 193 - 198" title="On four new polydesmoid millipeds." volume="61" year="1950 b">Causey (1950b)</bibRefCitation>
describes the habitat in which she and her husband found
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. davidcauseyi" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="davidcauseyi">
<emphasis italics="true" pageId="0" pageNumber="1">N. davidcauseyi</emphasis>
</taxonomicName>
as an eastern-facing hillside in an oak-hickory woodland. Two individuals of
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. davidcauseyi" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="davidcauseyi">
<emphasis italics="true" pageId="0" pageNumber="1">N. davidcauseyi</emphasis>
</taxonomicName>
were collected from Steel Creek Campground in Arkansas by M. Skvarla (NAN0537) who listed oak, hickory, American beech, and eastern red cedar as the primary tree species in the area (
<bibRefCitation DOI="https://doi.org/10.3897/BDJ.3.e6832" author="Skvarla, M" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" refId="B64" refString="Skvarla, M, Fisher, D, Schnepp, K, Dowling, A, 2015. Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. I. Select beetles (Coleoptera: Buprestidae, Carabidae, Cerambycidae, Curculionoidea excluding Scolytinae). Biodiversity Data Journal 3: e6832. https://doi.org/10.3897/BDJ.3.e6832" title="Terrestrial arthropods of Steel Creek, Buffalo National River, Arkansas. I. Select beetles (Coleoptera: Buprestidae, Carabidae, Cerambycidae, Curculionoidea excluding Scolytinae). Biodiversity Data Journal 3: e 6832." url="https://doi.org/10.3897/BDJ.3.e6832" year="2015">Skvarla et al. 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
<bibRefCitation author="Causey, NB" journalOrPublisher="Entomological News" pageId="0" pageNumber="1" pagination="193 - 198" refId="B11" refString="Causey, NB, 1950b. On four new polydesmoid millipeds. Entomological News 61: 193 - 198" title="On four new polydesmoid millipeds." volume="61" year="1950 b">Causey (1950b)</bibRefCitation>
gave no explanation of her choice in name for this species, however it is presumed that this species is named after the
<normalizedToken originalValue="authors">author's</normalizedToken>
husband and its collector, Dr. David Causey.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">United States, Arkansas, Newton County, about three miles northwest of Jasper.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="notes">
<paragraph pageId="0" pageNumber="1">Notes.</paragraph>
<paragraph pageId="0" pageNumber="1">
In the original description
<bibRefCitation author="Causey, NB" journalOrPublisher="Entomological News" pageId="0" pageNumber="1" pagination="193 - 198" refId="B11" refString="Causey, NB, 1950b. On four new polydesmoid millipeds. Entomological News 61: 193 - 198" title="On four new polydesmoid millipeds." volume="61" year="1950 b">Causey (1950b</bibRefCitation>
: 194) designates a male holotype and one male and two immature paratypes collected by Dr. David Causey on 25 August 1950.
<bibRefCitation author="Hennen, DA" journalOrPublisher="Insecta Mundi" pageId="0" pageNumber="1" pagination="1 - 21" publicationUrl="http://digitalcommons.unl.edu/insectamundi/924" refId="B28" refString="Hennen, DA, Shelley, RM, 2015. A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964. Insecta Mundi 0418: 1 - 21, http://digitalcommons.unl.edu/insectamundi/924" title="A contribution on the milliped tribe Nannariini (Polydesmida: Xystodesmidae): Revalidation of Mimuloria Chamberlin 1928; identities of Fontaria oblonga C. L. Koch 1847, and Nannaria minor Chamberlin 1918; elucidation of the tribal range; and commentaries on Nannaria Chamberlin 1918, and Oenomaea Hoffman 1964." url="http://digitalcommons.unl.edu/insectamundi/924" volume="0418" year="2015">Hennen and Shelley (2015</bibRefCitation>
: 10) refer to a female paratype and one immature paratype, which would suggest that
<normalizedToken originalValue="Causeys">Causey's</normalizedToken>
original identification of the female specimen as an immature was incorrect.
</paragraph>
</subSubSection>
</treatment>
</document>