145 lines
16 KiB
XML
145 lines
16 KiB
XML
<document id="D94F5216684B2BD3F69CC27E4189479E" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF9F4C74FF53DB8D04790011" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes bicornis Irmler 2021, spec. nov." docType="treatment" docVersion="3" lastPageNumber="36" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="36" updateTime="1699437350205" updateUser="plazi">
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<mods:titleInfo id="5E972C19FBFA1409714CE842B97FBB69">
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<mods:title id="49B04A9995ED1CFEC6C3CC31500EDFE7">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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</mods:titleInfo>
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<mods:name id="55E94EE734D59FB43F5D12307FD8A3A7" type="personal">
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<mods:role id="864F119C6A4951A9549C0E9A61450170">
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<mods:roleTerm id="C8C7180053A4545568AC638ECA711094">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A2B0F055062766B57C04B2FD624224E7">Irmler, Ulrich</mods:namePart>
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<mods:typeOfResource id="836C16AD0859891A36CAD4A788AF6F72">text</mods:typeOfResource>
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<mods:title id="12E6D95D3001B40180889DC707BDFDB5">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="034AE9F0C8BD05171FC6572C0B82E2D2">
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<mods:date id="5142292A479E18325A97AD57249EA890">2021</mods:date>
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<mods:detail id="4C217449FC5F3599F60008ACCE46BBE2" type="pubDate">
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<mods:number id="AD6C6D764C494A3F4D02EE3B780F94C4">2021-06-30</mods:number>
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<mods:detail id="20E2019FF89BAE9D1AC9D46A2BD013D7" type="volume">
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<mods:number id="B5B9CCC442CBC0C77F01061B01739745">71</mods:number>
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</mods:detail>
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<mods:detail id="519D98120FE82DA11B02EB450531377C" type="issue">
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<mods:number id="0C257E6B44B6987398B6333F90E0638B">1</mods:number>
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</mods:detail>
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<mods:extent id="AFC87D3074554B2C664EEA2EBD50841E" unit="page">
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<mods:start id="7B965864A37ABD13F43A785B7CE5637C">29</mods:start>
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<mods:end id="A72EC28E52A408C84CF5E75F24D9D7F0">85</mods:end>
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</mods:extent>
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<mods:classification id="18BDA9E639BEDDB670928E6BA6E29587">journal article</mods:classification>
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<mods:identifier id="330465B320241476782A7420812C1B19" type="DOI">10.21248/contrib.entomol.71.1.029-085</mods:identifier>
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<mods:identifier id="9F9E3FE303FDD0B1278649A876D665ED" type="ISSN">0005-805X</mods:identifier>
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<mods:identifier id="801F80555451068B3DB3E024775E7683" type="Zenodo-Dep">5742999</mods:identifier>
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<treatment id="3A6C87E6FF9F4C74FF53DB8D04790011" ID-DOI="http://doi.org/10.5281/zenodo.5743053" ID-Zenodo-Dep="5743053" LSID="urn:lsid:plazi:treatment:3A6C87E6FF9F4C74FF53DB8D04790011" httpUri="http://treatment.plazi.org/id/3A6C87E6FF9F4C74FF53DB8D04790011" lastPageNumber="36" pageId="7" pageNumber="36">
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<subSubSection id="FADF657BFF9F4C74FF53DB8D0235033E" box="[189,531,243,270]" pageId="7" pageNumber="36" type="nomenclature">
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<paragraph id="B27A36F0FF9F4C74FF53DB8D0235033E" blockId="7.[189,531,243,303]" box="[189,531,243,270]" pageId="7" pageNumber="36">
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<heading id="E932819CFF9F4C74FF53DB8D0235033E" box="[189,531,243,270]" centered="true" fontSize="12" level="2" pageId="7" pageNumber="36" reason="2">
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<taxonomicName id="75C54D73FF9F4C74FF53DB8D0185033E" ID-CoL="8Y632" authority="Irmler, 2021" authorityName="Irmler" authorityYear="2021" box="[189,419,243,270]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="bicornis" status="sp. nov.">
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<emphasis id="80B1EAE2FF9F4C74FF53DB8D0185033E" box="[189,419,243,270]" italics="true" pageId="7" pageNumber="36">Lithocharodes bicornis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="9B825799FF9F4C74FE44DB8D0235033E" box="[426,531,243,270]" pageId="7" pageNumber="36" rank="species">spec. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FADF657BFF9F4C74FF53DA650145031F" box="[189,355,283,303]" pageId="7" pageNumber="36" type="description">
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<paragraph id="B27A36F0FF9F4C74FF53DA650145031F" blockId="7.[189,531,243,303]" box="[189,355,283,303]" pageId="7" pageNumber="36">
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<figureCitation id="2AFE2A75FF9F4C74FF53DA6500D7031F" box="[189,241,283,303]" captionStart="Fig" captionStartId="40.[189,222,1871,1893]" captionTargetBox="[197,1420,250,1840]" captionTargetId="figure-109@40.[189,1436,232,1848]" captionTargetPageId="40" captionText="Fig. 8–12: L.-hanagarthi-group and L.-sordida-group:Lithocharodes boliviana (8), L. hanagarthi (9), L. obscura (10), L. bicornis (11), L. sordida (12); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: A – D: 0.2 mm; B: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743009" httpUri="https://zenodo.org/record/5743009/files/figure.png" pageId="7" pageNumber="36">Figs 11</figureCitation>
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a-d, 44 O, 49 C
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</paragraph>
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</subSubSection>
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<subSubSection id="FADF657BFF9F4C74FF79DA3700F803F1" pageId="7" pageNumber="36" type="materials_examined">
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<paragraph id="B27A36F0FF9F4C74FF79DA3700F803F1" blockId="7.[151,756,329,449]" pageId="7" pageNumber="36">
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<emphasis id="80B1EAE2FF9F4C74FF79DA3701130351" bold="true" box="[151,309,329,353]" pageId="7" pageNumber="36">Type material</emphasis>
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:
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<materialsCitation id="02AD3CADFF9F4C74FEA6DA3700FD03F1" collectingDate="1975-03-25" collectionCode="UIC" collectorName="W. Hanagarth" country="Peru" latitude="-9.616667" location="Panguana Station" longLatPrecision="1299" longitude="-74.933334" municipality="Llullapichis" pageId="7" pageNumber="36" specimenCount="1" stateProvince="Huanuco" typeStatus="holotype">
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male,
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<typeStatus id="6D7E8852FF9F4C74FE61DA3701D60351" box="[399,496,329,353]" pageId="7" pageNumber="36" type="holotype">holotype</typeStatus>
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:
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<collectingCountry id="CAD27660FF9F4C74FDEEDA3402120351" box="[512,564,330,353]" name="Peru" pageId="7" pageNumber="36">Peru</collectingCountry>
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,
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<collectingRegion id="7001F812FF9F4C74FDABDA34028F0351" box="[581,681,330,353]" country="Peru" name="Huanuco" pageId="7" pageNumber="36">Huanuco</collectingRegion>
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,
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<collectingMunicipality id="521EAC8AFF9F4C74FD57DA3700C503B1" pageId="7" pageNumber="36">Llullapichis</collectingMunicipality>
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,
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<location id="B71A602BFF9F4C74FF1CDA14019603B1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF9F4C74FF53DB8D04790011:B71A602BFF9F4C74FF1CDA14019603B1" box="[242,432,361,386]" country="Peru" latitude="-9.616667" longLatPrecision="1299" longitude="-74.933334" municipality="Llullapichis" name="Panguana Station" pageId="7" pageNumber="36" stateProvince="Huanuco">Panguana Station</location>
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(
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<geoCoordinate id="D7F15037FF9F4C74FE2CDA17023A03B1" box="[450,540,361,385]" degrees="74" direction="west" minutes="56" orientation="longitude" pageId="7" pageNumber="36" precision="925" value="-74.933334">74°56'W</geoCoordinate>
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,
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<geoCoordinate id="D7F15037FF9F4C74FDC5DA17024903B1" box="[555,623,361,385]" degrees="9" direction="south" minutes="37" orientation="latitude" pageId="7" pageNumber="36" precision="925" value="-9.616667">9°37'S</geoCoordinate>
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), floodplain forest, sifted litter, male,
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<date id="C67B1030FF9F4C74FE58DAF702380391" box="[438,542,393,417]" pageId="7" pageNumber="36" value="1975-03-25">
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<collectingDate id="D63FE9D8FF9F4C74FE58DAF702380391" box="[438,542,393,417]" pageId="7" pageNumber="36" value="1975-03-25">25.3.1975</collectingDate>
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</date>
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, leg.
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<collectorName id="1F305326FF9F4C74FD8EDAF402D20391" box="[608,756,393,417]" pageId="7" pageNumber="36">W. Hanagarth</collectorName>
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(
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<collectionCode id="D4D4AE35FF9F4C74FF71DAD700F403F1" box="[159,210,425,449]" pageId="7" pageNumber="36">UIC</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="FADF657BFF9F4C74FF79DA9701250131" pageId="7" pageNumber="36" type="diagnosis">
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<paragraph id="B27A36F0FF9F4C74FF79DA9701250131" blockId="7.[151,756,489,769]" pageId="7" pageNumber="36">
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<emphasis id="80B1EAE2FF9F4C74FF79DA9701220031" bold="true" box="[151,260,489,513]" pageId="7" pageNumber="36">Diagnosis</emphasis>
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:
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<taxonomicName id="75C54D73FF9F4C74FECEDA97023E0031" authorityName="Irmler" authorityYear="2021" box="[288,536,489,513]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="bicornis">
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<emphasis id="80B1EAE2FF9F4C74FECEDA97023E0031" box="[288,536,489,513]" italics="true" pageId="7" pageNumber="36">Lithocharodes bicornis</emphasis>
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</taxonomicName>
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closely resembles
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<taxonomicName id="75C54D73FF9F4C74FF79D97400D90011" authorityName="SHARP" authorityYear="1885" box="[151,255,521,545]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="sordida">
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<emphasis id="80B1EAE2FF9F4C74FF79D97400D90011" box="[151,255,521,545]" italics="true" pageId="7" pageNumber="36">L. sordida</emphasis>
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</taxonomicName>
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in size, colouration, punctation of the head and even in the structure of the aedeagus and the shape of the paramere. Males are easily to separate by the structure of sternite and tergite VII. The pair of spines at the posterior margin of sternite VII of
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<taxonomicName id="75C54D73FF9F4C74FE39D9F402650091" authorityName="Irmler" authorityYear="2021" box="[471,579,649,673]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="bicornis">
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<emphasis id="80B1EAE2FF9F4C74FE39D9F402650091" box="[471,579,649,673]" italics="true" pageId="7" pageNumber="36">L. bicornis</emphasis>
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</taxonomicName>
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is unique in the genus, although many
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<taxonomicName id="75C54D73FF9F4C74FE62D9D7025600F1" box="[396,624,681,705]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="36" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="80B1EAE2FF9F4C74FE62D9D7023B00F1" box="[396,541,681,705]" italics="true" pageId="7" pageNumber="36">Lithocharodes</emphasis>
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species
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</taxonomicName>
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have paired processes at sternite VII but these are broader and more teeth-like.
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</paragraph>
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</subSubSection>
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<subSubSection id="FADF657BFF9F4C74FF79D857051503B1" pageId="7" pageNumber="36" type="description">
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<paragraph id="B27A36F0FF9F4C74FF79D85702610151" blockId="7.[151,756,809,2081]" pageId="7" pageNumber="36">
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<emphasis id="80B1EAE2FF9F4C74FF79D857013F0171" bold="true" box="[151,281,809,833]" pageId="7" pageNumber="36">Description</emphasis>
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: Length:
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<quantity id="753D9B15FF9F4C74FE91D85701F40171" box="[383,466,809,833]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="7" pageNumber="36" unit="mm" value="4.5">4.5 mm</quantity>
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. Colouration: Brown; head and elytra darker brown than pronotum.
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</paragraph>
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<paragraph id="B27A36F0FF9F4C74FF79D817051503B1" blockId="7.[151,756,809,2081]" lastBlockId="7.[794,1399,233,385]" pageId="7" pageNumber="36">
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Head:
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<quantity id="753D9B15FF9F4C74FF04D817016F01B1" box="[234,329,873,897]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="7" pageNumber="36" unit="mm" value="0.77">0.77 mm</quantity>
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long,
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<quantity id="753D9B15FF9F4C74FE4BD817022501B1" box="[421,515,873,897]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="7" pageNumber="36" unit="mm" value="0.67">0.67 mm</quantity>
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wide; eyes moderately large; not prominent; postocular sides divergent; 2.5 times as long as eyes; posterior angles widely rounded to neck; posterior margin slightly emarginate in front of neck; interantennal furrows extremely weak; setiferous punctation deep and dense; on average, interstices half as wide as diameter of punctures; on anterior vertex denser than on posterior vertex; surface without microsculpture; glossy. Antennae with first antennomere reaching half-length of head; second and third antennomere each conical and approximately twice as long as wide; subsequent antennomeres wider than long and increasing in width; fourth antennomere 1.5 times as wide as long; tenth antennomere twice as wide as long; all antennomeres pubescent. Pronotum:
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<quantity id="753D9B15FF9F4C74FE75DE5701DC0771" box="[411,506,1321,1345]" metricMagnitude="-4" metricUnit="m" metricValue="8.7" pageId="7" pageNumber="36" unit="mm" value="0.87">0.87 mm</quantity>
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long,
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<quantity id="753D9B15FF9F4C74FDA3DE57028A0771" box="[589,684,1321,1345]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="36" unit="mm" value="0.6">0.60 mm</quantity>
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wide; widest in anterior third; anteriad, conically narrowed toward neck; posteriad, slightly narrowed in middle third; nearly parallel in posterior third; posterior angles shortly rounded; posterior margin nearly straight; setiferous punctation deep and dense; on average, interstices between punctures half as wide as diameter of punctures; wide midline impunctate; midline wider in anterior half than in posterior half; irregular row of punctures adjacent to impunctate midline with approximately 20 punctures; surface without microsculpture; glossy. Elytra:
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<quantity id="753D9B15FF9F4C74FD78DD1702D204B1" box="[662,756,1641,1665]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="7" pageNumber="36" unit="mm" value="0.83">0.83 mm</quantity>
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long,
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<quantity id="753D9B15FF9F4C74FF34DDF7011F0491" box="[218,313,1673,1697]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="7" pageNumber="36" unit="mm" value="0.69">0.69 mm</quantity>
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wide; humeral angles sub-rectangular; sides slightly divergent posteriad; posterior angles subrectangular; posterior margin nearly straight; setiferous punctation as dense and deep as on pronotum but partly coriaceous; surface with weak irregular microsculpture; less shiny than pronotum. Abdomen with dense setiferous punctation; punctures distinctly finer than on fore-body; sternite VII of male with concavely emarginate posterior margin; posterior margin with two fine spines and several thick setae; tergite VII of male with straight posterior margin; short rectangular process with pair of teeth at each angle in middle of posterior margin. Meso- and metatibia with one subapical ctenidium each. Aedeagus oval; in anterior half sub-rectangular; endophallus apically with three long and thick teeth; posterior part sack-like; covered by minute teeth; paramere broad; spoon-like; at apex with transparent inner lobe; several long, fine setae and several short and curved setae.
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</paragraph>
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</subSubSection>
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<subSubSection id="FADF657BFF9F4C74FCF4DAD704790011" pageId="7" pageNumber="36" type="etymology">
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<paragraph id="B27A36F0FF9F4C74FCF4DAD704790011" blockId="7.[794,1398,425,545]" pageId="7" pageNumber="36">
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<emphasis id="80B1EAE2FF9F4C74FCF4DAD703B703F1" bold="true" box="[794,913,425,449]" pageId="7" pageNumber="36">Etymology</emphasis>
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: The species epithet is a combination of the Latin words
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<emphasis id="80B1EAE2FF9F4C74FC3EDAB703C203D1" box="[976,996,457,481]" italics="true" pageId="7" pageNumber="36">bi</emphasis>
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(meaning two) and
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<emphasis id="80B1EAE2FF9F4C74FB26DAB4052003D1" box="[1224,1286,458,481]" italics="true" pageId="7" pageNumber="36">cornis</emphasis>
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(meaning horn) and refers to the spectacular spines at the posterior margin of sternite VII of male.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |