treatments-xml/data/3A/6C/87/3A6C87E6FF9E4C75FC8BDB8D055C0A11.xml
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<document id="C7C414C68AF180A82C867D4E1643F711" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF9E4C75FC8BDB8D055C0A11" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes bicolor Irmler 2021, new combination" docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="35" updateTime="1699437350205" updateUser="plazi">
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<mods:title id="EA4593543A4E7A5F7D301AD5F653F9A3">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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<mods:namePart id="B1C42FA3738D29B92AB84C37144D4543">Irmler, Ulrich</mods:namePart>
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<mods:title id="5DB2133E3A4390476E6F42A2B792B6FC">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="ADAC5B685F42FF0524D01D8A32755C80">2021</mods:date>
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<subSubSection id="FADF657BFF9E4C75FC8BDB8D05B4033E" box="[869,1426,243,270]" pageId="6" pageNumber="35" type="nomenclature">
<paragraph id="B27A36F0FF9E4C75FC8BDB8D05B4033E" blockId="6.[869,1426,243,303]" box="[869,1426,243,270]" pageId="6" pageNumber="35">
<heading id="E932819CFF9E4C75FC8BDB8D05B4033E" box="[869,1426,243,270]" centered="true" fontSize="12" level="2" pageId="6" pageNumber="35" reason="2">
<taxonomicName id="75C54D73FF9E4C75FC8BDB8D04F0033E" ID-CoL="8Y62Z" authority="(SHARP, 1885)" authorityName="Irmler" authorityYear="2021" baseAuthorityName="SHARP" baseAuthorityYear="1885" box="[869,1238,243,270]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="bicolor" status="comb. nov.">
<emphasis id="80B1EAE2FF9E4C75FC8BDB8D0467033E" box="[869,1089,243,270]" italics="true" pageId="6" pageNumber="35">Lithocharodes bicolor</emphasis>
(
<bibRefCitation id="D6544B01FF9E4C75FBA0DB8D04F6033E" author="SHARP, D." box="[1102,1232,243,270]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP, 1885</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="9B825799FF9E4C75FB32DB8D05B4033E" box="[1244,1426,243,270]" pageId="6" pageNumber="35" rank="species">new combination</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FC8BDA65042A031F" box="[869,1036,283,303]" pageId="6" pageNumber="35" type="description">
<paragraph id="B27A36F0FF9E4C75FC8BDA65042A031F" blockId="6.[869,1426,243,303]" box="[869,1036,283,303]" pageId="6" pageNumber="35">
<figureCitation id="2AFE2A75FF9E4C75FC8BDA6503BD031F" box="[869,923,283,303]" captionStart="Fig" captionStartId="44.[189,222,1959,1981]" captionTargetBox="[199,1417,242,1939]" captionTargetId="figure-111@44.[189,1436,232,1949]" captionTargetPageId="44" captionText="Fig. 3136: L.-??-group: Lithocharodes spinigera (31), L. curtipennis (32), L. fusciventris (33), L. nigerrima (34), L. puncticeps (35), L. bicolor (36); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: AD: 0.2 mm; B: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743017" httpUri="https://zenodo.org/record/5743017/files/figure.png" pageId="6" pageNumber="35">Figs 36</figureCitation>
a-c, 45 G, 50 E
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FCD1DA35051503B0" pageId="6" pageNumber="35" type="reference_group">
<paragraph id="B27A36F0FF9E4C75FCD1DA3504BE0351" blockId="6.[831,1331,331,385]" box="[831,1176,331,353]" pageId="6" pageNumber="35">
<treatmentCitationGroup id="92D511DEFF9E4C75FCD1DA3504BE0351" box="[831,1176,331,353]" pageId="6" pageNumber="35">
<treatmentCitation id="336410E1FF9E4C75FCD1DA3504BE0351" author="SHARP, D." box="[831,1176,331,353]" page="495" pageId="6" pageNumber="35" year="1885">
<taxonomicName id="75C54D73FF9E4C75FCD1DA3504BE0351" ID-CoL="4Y8Z9" authority="SHARP, 1885: 495" authorityName="SHARP" authorityPageNumber="495" authorityYear="1885" box="[831,1176,331,353]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="80B1EAE2FF9E4C75FCD1DA3503D60350" box="[831,1008,331,352]" italics="true" pageId="6" pageNumber="35">Somoleptus bicolor</emphasis>
<bibRefCitation id="D6544B01FF9E4C75FC1BDA3504BE0351" author="SHARP, D." box="[1013,1176,331,353]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP, 1885: 495</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="B27A36F0FF9E4C75FCD1DA15051503B0" blockId="6.[831,1331,331,385]" box="[831,1331,363,385]" pageId="6" pageNumber="35">
<treatmentCitationGroup id="92D511DEFF9E4C75FCD1DA15051503B0" box="[831,1331,363,385]" pageId="6" pageNumber="35">
<treatmentCitation id="336410E1FF9E4C75FCD1DA1504BD03B1" author="SHARP, D." box="[831,1179,363,385]" page="496" pageId="6" pageNumber="35" year="1885">
<taxonomicName id="75C54D73FF9E4C75FCD1DA1504BD03B1" ID-CoL="4Y8ZJ" authority="SHARP, 1885: 496" authorityName="SHARP" authorityPageNumber="496" authorityYear="1885" box="[831,1179,363,385]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="80B1EAE2FF9E4C75FCD1DA1503D403B0" box="[831,1010,363,384]" italics="true" pageId="6" pageNumber="35">Somoleptus gracilis</emphasis>
<bibRefCitation id="D6544B01FF9E4C75FC16DA1504BD03B1" author="SHARP, D." box="[1016,1179,363,385]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP, 1885: 496</bibRefCitation>
</taxonomicName>
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<emphasis id="80B1EAE2FF9E4C75FB4EDA15051503B0" bold="true" box="[1184,1331,363,384]" pageId="6" pageNumber="35">
<taxonomicNameLabel id="9B825799FF9E4C75FB4EDA15051503B0" box="[1184,1331,363,384]" pageId="6" pageNumber="35">new synonymy</taxonomicNameLabel>
</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FCD1DAD7038E0011" pageId="6" pageNumber="35" type="materials_examined">
<paragraph id="B27A36F0FF9E4C75FCD1DAD7038E0011" blockId="6.[831,1436,425,545]" pageId="6" pageNumber="35">
<emphasis id="80B1EAE2FF9E4C75FCD1DAD7047503F1" bold="true" box="[831,1107,425,449]" pageId="6" pageNumber="35">Type material examined</emphasis>
:
<materialsCitation id="02AD3CADFF9E4C75FB88DAD703830011" collectionCode="BMNH" collectorName="Champion &amp; Cordoba &amp; Salle" country="Guatemala" elevation="2377" location="Quetzaltenango" pageId="6" pageNumber="35" specimenCount="2" stateProvince="Quetzaltenango" typeStatus="syntype">
female,
<typeStatus id="6D7E8852FF9E4C75FB50DAD4053503F2" box="[1214,1299,426,450]" pageId="6" pageNumber="35" type="syntype">syntype</typeStatus>
:
<collectingCountry id="CAD27660FF9E4C75FACDDAD705BF03F1" box="[1315,1433,425,449]" name="Guatemala" pageId="6" pageNumber="35">Guatemala</collectingCountry>
,
<collectingRegion id="7001F812FF9E4C75FCD1DAB703CF03D1" box="[831,1001,457,481]" country="Guatemala" name="Quetzaltenango" pageId="6" pageNumber="35">Quetzaltenango</collectingRegion>
,
<quantity id="753D9B15FF9E4C75FBEADAB7047003D1" box="[1028,1110,457,481]" metricMagnitude="3" metricUnit="m" metricValue="2.37744" pageId="6" pageNumber="35" unit="ft" value="7800.0">
<elevation id="39E8D1C3FF9E4C75FBEADAB7047003D1" box="[1028,1110,457,481]" metricMagnitude="3" metricUnit="m" metricValue="2.37744" pageId="6" pageNumber="35" unit="ft" value="7800.0">7800 ft.</elevation>
</quantity>
, leg.
<collectorName id="1F305326FF9E4C75FB42DAB7053B03D1" box="[1196,1309,457,481]" pageId="6" pageNumber="35">Champion</collectorName>
(
<collectionCode id="D4D4AE35FF9E4C75FAD5DAB705B603D1" box="[1339,1424,457,481]" pageId="6" pageNumber="35">BMNH</collectionCode>
); male,
<typeStatus id="6D7E8852FF9E4C75FC6FDA9703F90031" box="[897,991,489,513]" pageId="6" pageNumber="35" type="holotype">holotype</typeStatus>
of
<taxonomicName id="75C54D73FF9E4C75FBEBDA9704400031" authorityName="SHARP" authorityYear="1885" box="[1029,1126,489,513]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="80B1EAE2FF9E4C75FBEBDA9704400031" box="[1029,1126,489,513]" italics="true" pageId="6" pageNumber="35">S. gracilis</emphasis>
</taxonomicName>
,
<collectingCountry id="CAD27660FF9E4C75FB9BDA9704E00031" box="[1141,1222,489,513]" name="Mexico" pageId="6" pageNumber="35">Mexico</collectingCountry>
,
<collectorName id="1F305326FF9E4C75FB3CDA9705140031" box="[1234,1330,489,513]" pageId="6" pageNumber="35">Cordoba</collectorName>
, leg.
<collectorName id="1F305326FF9E4C75FA85DA9705BA0031" box="[1387,1436,489,513]" pageId="6" pageNumber="35">Sallé</collectorName>
(
<collectionCode id="D4D4AE35FF9E4C75FCA9D97703BB0011" box="[839,925,521,545]" pageId="6" pageNumber="35">BMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FCD1D93705740191" pageId="6" pageNumber="35" type="diagnosis">
<paragraph id="B27A36F0FF9E4C75FCD1D93705740191" blockId="6.[831,1437,585,930]" pageId="6" pageNumber="35">
<emphasis id="80B1EAE2FF9E4C75FCD1D937038A0051" bold="true" box="[831,940,585,609]" pageId="6" pageNumber="35">Diagnosis</emphasis>
:
<bibRefCitation id="D6544B01FF9E4C75FC51D937047E0051" author="SHARP, D." box="[959,1112,585,609]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP (1885)</bibRefCitation>
already noted the similarity between
<taxonomicName id="75C54D73FF9E4C75FC4AD917044900B1" authorityName="SHARP" authorityYear="1885" box="[932,1135,617,641]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="bicolor">
<emphasis id="80B1EAE2FF9E4C75FC4AD917044900B1" box="[932,1135,617,641]" italics="true" pageId="6" pageNumber="35">Somoleptus bicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="75C54D73FF9E4C75FB40D917053600B1" authorityName="SHARP" authorityYear="1885" box="[1198,1296,617,641]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="80B1EAE2FF9E4C75FB40D917053600B1" box="[1198,1296,617,641]" italics="true" pageId="6" pageNumber="35">S. gracilis</emphasis>
</taxonomicName>
. The differences in size are still less than given by
<bibRefCitation id="D6544B01FF9E4C75FB38D9F7054C0091" author="SHARP, D." box="[1238,1386,649,673]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP (1885)</bibRefCitation>
. The minute differences in size may be within the variability of the species or in the different sexes of the
<specimenCount id="A4C3FD79FF9E4C75FAC3D9B4035E0131" pageId="6" pageNumber="35" type="generic">two specimens</specimenCount>
studied. The absence of the cone-like structure at the central orifice of the aedeagus indicates the species as a member of the genus
<taxonomicName id="75C54D73FF9E4C75FBAAD85704F30171" authorityName="SHARP" authorityYear="1876" box="[1092,1237,809,833]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="80B1EAE2FF9E4C75FBAAD85704F30171" box="[1092,1237,809,833]" italics="true" pageId="6" pageNumber="35">Lithocharodes</emphasis>
</taxonomicName>
. As
<bibRefCitation id="D6544B01FF9E4C75FAE6D85705BB0171" author="SHARP, D." box="[1288,1437,809,833]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">SHARP (1885)</bibRefCitation>
already described another species as
<emphasis id="80B1EAE2FF9E4C75FB27D837046601B1" italics="true" pageId="6" pageNumber="35">
<taxonomicName id="75C54D73FF9E4C75FB27D837037E01B1" authorityName="SHARP" authorityYear="1885" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="gracilis">Lithocharodes gracilis</taxonomicName>
,
<taxonomicName id="75C54D73FF9E4C75FC8CD817046601B1" authorityName="Irmler" authorityYear="2021" baseAuthorityName="SHARP" baseAuthorityYear="1885" box="[866,1088,873,897]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="bicolor">Lithocharodes bicolor</taxonomicName>
</emphasis>
is the valid name for the species, although only the female
<typeStatus id="6D7E8852FF9E4C75FBA0D8F4045A0192" box="[1102,1148,906,930]" pageId="6" pageNumber="35">type</typeStatus>
could be examined.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FCD1D8B7055C0A11" pageId="6" pageNumber="35" type="description">
<paragraph id="B27A36F0FF9E4C75FCD1D8B704F20611" blockId="6.[831,1437,969,2081]" pageId="6" pageNumber="35">
<emphasis id="80B1EAE2FF9E4C75FCD1D8B703E701D1" bold="true" box="[831,961,969,993]" pageId="6" pageNumber="35">Description</emphasis>
: Length: 5.0
<quantity id="753D9B15FF9E4C75FBBFD8B7048301D1" box="[1105,1189,969,993]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="35" unit="mm" value="5.2">5.2 mm</quantity>
. Colouration: Reddishbrown; pronotum and posterior margin of elytra light red to orange; legs and antennae light red.
</paragraph>
<paragraph id="B27A36F0FF9E4C75FCD1DF57055C0A11" blockId="6.[831,1437,969,2081]" pageId="6" pageNumber="35">
Head:
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long,
<quantity id="753D9B15FF9E4C75FBCADF5704A60671" box="[1060,1152,1065,1089]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="6" pageNumber="35" unit="mm" value="0.63">0.63 mm</quantity>
wide; eyes small; postocular space nearly 3.5 times as long as eyes; postocular sides slightly divergent; posterior angles rounded; interantennal furrows weak; setiferous punctation moderately deep; sparse; on average, interstices between punctures as wide as diameter of punctures; on posterior vertex interstices wider; nearly twice as wide as diameter of punctures; surface without microsculpture; polished. Antennae with second and third antennomere each longer than wide; second antennomere longer than third; subsequent antennomeres much wider than long and increasing in width; fourth antennomere twice as wide as long; tenth antennomere nearly three times as wide as long; all antennomeres pubescent. Pronotum:
<quantity id="753D9B15FF9E4C75FC1ADEB7047607D1" box="[1012,1104,1481,1505]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="6" pageNumber="35" unit="mm" value="0.94">0.94 mm</quantity>
long,
<quantity id="753D9B15FF9E4C75FB7BDEB704D707D1" box="[1173,1265,1481,1505]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" pageId="6" pageNumber="35" unit="mm" value="0.54">0.54 mm</quantity>
wide; widest in anterior third, anteriad smoothly narrowed toward neck in wide curve; posterior third nearly parallel; posterior angles widely rounded; setiferous punctation as deep and sparse as on posterior head; wide midline impunctate; adjacent to midline with irregular line of about 22-25 punctures; surface without microsculpture; polished. Elytra:
<quantity id="753D9B15FF9E4C75FAAEDDF705BB0491" box="[1344,1437,1673,1697]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="6" pageNumber="35" unit="mm" value="0.89">0.89 mm</quantity>
long,
<quantity id="753D9B15FF9E4C75FC93DDD703FF04F1" box="[893,985,1705,1729]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="6" pageNumber="35" unit="mm" value="0.78">0.78 mm</quantity>
wide; sides slightly divergent to posterior angles; posterior and humeral angles sub-rectangular; setiferous punctation slightly deeper and denser than on head and pronotum; without microsculpture; less shiny than pronotum. Abdomen with similar setiferous punctation as on elytra; surface without microsculpture; polished; sternite VII of male with short central incision; tergite VII of male with straight posterior margin. Aedeagus longoval; thick endophallus with one torsion in anterior half; endophallus covered by minute teeth; paramere straight with wide base; conically narrowed toward apex; inner face with long setae; setae at base much longer than at apex.
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