treatments-xml/data/3A/6C/87/3A6C87E6FF9D4C75FCD1DC2D021F05F1.xml
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<document id="FC5DA81DE2EFDA0DDF978764054C58AA" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF9D4C75FCD1DC2D021F05F1" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes bierigi Irmler 2021, spec. nov." docType="treatment" docVersion="3" lastPageNumber="35" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="34" updateTime="1699437350205" updateUser="plazi">
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<mods:title id="C72880010134D3F2E12F23A0BB5E727F">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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<mods:namePart id="98BFD0905C3DB0ACF3D8E2E50A405316">Irmler, Ulrich</mods:namePart>
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<mods:title id="0CCDC8F3E409F4F260BC7BF499C8614E">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:date id="3B094C267374499373782D6CD7C78AFE">2021</mods:date>
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<mods:identifier id="08FE5689A5341EF595AB669B02980F67" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="9224686B8B91FD265F0D23ECB7D4CF0B" type="Zenodo-Dep">5742999</mods:identifier>
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<treatment id="3A6C87E6FF9D4C75FCD1DC2D021F05F1" ID-DOI="http://doi.org/10.5281/zenodo.5743051" ID-Zenodo-Dep="5743051" LSID="urn:lsid:plazi:treatment:3A6C87E6FF9D4C75FCD1DC2D021F05F1" httpUri="http://treatment.plazi.org/id/3A6C87E6FF9D4C75FCD1DC2D021F05F1" lastPageId="6" lastPageNumber="35" pageId="5" pageNumber="34">
<subSubSection id="FADF657BFF9D4C76FCD1DC2D056505BF" pageId="5" pageNumber="34" type="nomenclature">
<paragraph id="B27A36F0FF9D4C76FCD1DC2D04A3055E" blockId="5.[831,1347,1875,1967]" box="[831,1157,1875,1902]" pageId="5" pageNumber="34">
<heading id="E932819CFF9D4C76FCD1DC2D04A3055E" box="[831,1157,1875,1902]" fontSize="12" level="2" pageId="5" pageNumber="34" reason="2">
<taxonomicName id="75C54D73FF9D4C76FCD1DC2D0430055E" ID-CoL="8Y633" authority="Irmler, 2021" authorityName="Irmler" authorityYear="2021" box="[831,1046,1875,1902]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="34" phylum="Arthropoda" rank="species" species="bierigi" status="sp. nov.">
<emphasis id="80B1EAE2FF9D4C76FCD1DC2D0430055E" box="[831,1046,1875,1902]" italics="true" pageId="5" pageNumber="34">Lithocharodes bierigi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B825799FF9D4C76FBF2DC2D04A3055E" box="[1052,1157,1875,1902]" pageId="5" pageNumber="34" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="B27A36F0FF9D4C76FCD1DC05056505BF" blockId="5.[831,1347,1875,1967]" box="[831,1347,1915,1935]" pageId="5" pageNumber="34">
<uri id="C6543AF2FF9D4C76FCD1DC05056505BF" box="[831,1347,1915,1935]" pageId="5" pageNumber="34">
urn:lsid:zoobank.org:act:
<uuid id="C6630C25FF9D4C76FC02DC05056505BF" box="[1004,1347,1915,1935]" pageId="5" pageNumber="34">40819238-8854-4AA3-8080-FEA8778C77CE</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9D4C76FCD1DCE503CF059F" box="[831,1001,1947,1967]" pageId="5" pageNumber="34" type="description">
<paragraph id="B27A36F0FF9D4C76FCD1DCE503CF059F" blockId="5.[831,1347,1875,1967]" box="[831,1001,1947,1967]" pageId="5" pageNumber="34">
<figureCitation id="2AFE2A75FF9D4C76FCD1DCE50353059F" box="[831,885,1947,1967]" captionStart="Fig" captionStartId="42.[189,222,2010,2032]" captionTargetBox="[217,1431,249,1995]" captionTargetId="figure-111@42.[189,1439,232,2005]" captionTargetPageId="42" captionText="Fig. 1924: L.-fuscipennis-group:Lithocharodesfuscipennis(19),L. bierigi(20),L. nigripennis(21),L. clavicornis(22),L. karinae (23), L. katharinae (24); aedeagus (a), paramere (b), sternite VII of male (c), tergite VII of male (d); scale bare: AD: 0.2 mm; B: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743013" httpUri="https://zenodo.org/record/5743013/files/figure.png" pageId="5" pageNumber="34">Figs 20</figureCitation>
a-d, 45 E, 49 H
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9D4C76FCF4DCB7057A0A11" pageId="5" pageNumber="34" type="materials_examined">
<paragraph id="B27A36F0FF9D4C76FCF4DCB7057A0A11" blockId="5.[794,1399,1993,2081]" pageId="5" pageNumber="34">
<emphasis id="80B1EAE2FF9D4C76FCF4DCB7040805D1" bold="true" box="[794,1070,1993,2017]" pageId="5" pageNumber="34">Type material examined</emphasis>
:
<materialsCitation id="02AD3CADFF9D4C76FBAFDCB705270A31" collectingDate="1939-11-08" collectionCode="FMNH" collectorName="A. Bierig" country="Costa Rica" location="Puntarenas" pageId="5" pageNumber="34" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
male,
<typeStatus id="6D7E8852FF9D4C76FB69DCB704CE05D1" box="[1159,1256,1993,2017]" pageId="5" pageNumber="34" type="holotype">holotype</typeStatus>
:
<collectingCountry id="CAD27660FF9D4C76FB16DCB7055405D1" box="[1272,1394,1993,2017]" name="Costa Rica" pageId="5" pageNumber="34">Costa Rica</collectingCountry>
,
<collectingRegion id="7001F812FF9D4C76FCF4DC9403B20A31" box="[794,916,2026,2049]" country="Costa Rica" name="Puntarenas" pageId="5" pageNumber="34">Puntarenas</collectingRegion>
,
<date id="C67B1030FF9D4C76FC73DC9704230A31" box="[925,1029,2025,2049]" pageId="5" pageNumber="34" value="1939-11-08">
<collectingDate id="D63FE9D8FF9D4C76FC73DC9704230A31" box="[925,1029,2025,2049]" pageId="5" pageNumber="34" value="1939-11-08">11.8.1939</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF9D4C76FBD7DC9704BE0A31" box="[1081,1176,2025,2049]" pageId="5" pageNumber="34">A. Bierig</collectorName>
(
<collectionCode id="D4D4AE35FF9D4C76FB4BDC9704DF0A31" box="[1189,1273,2025,2049]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="34" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="02AD3CADFF9D4C76FAE4DC94057E0A11" collectingDate="1939-11-08" collectionCode="FMNH, UIC" collectorName="A. Bierig" country="Costa Rica" location="Puntarenas" pageId="5" pageNumber="34" specimenCount="5" specimenCount-male="5" stateProvince="Puntarenas" typeStatus="paratype">
<typeStatus id="6D7E8852FF9D4C76FAE4DC9405520A32" box="[1290,1396,2026,2050]" pageId="5" pageNumber="34" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A4C3FD79FF9D4C76FCF4D377034C0A11" box="[794,874,2057,2081]" pageId="5" pageNumber="34" type="male">5 males</specimenCount>
with same data as holotype (4
<collectionCode id="D4D4AE35FF9D4C76FB5FD37405270A11" box="[1201,1281,2058,2081]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="34" type="Museum">FMNH</collectionCode>
, 1
<collectionCode id="D4D4AE35FF9D4C76FAF0D37705760A11" box="[1310,1360,2057,2081]" pageId="5" pageNumber="34">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FF53DB97023903D1" pageId="6" pageNumber="35" type="diagnosis">
<paragraph id="B27A36F0FF9E4C75FF53DB97023903D1" blockId="6.[189,794,233,481]" pageId="6" pageNumber="35">
<emphasis id="80B1EAE2FF9E4C75FF53DB97010A0331" bold="true" box="[189,300,233,257]" pageId="6" pageNumber="35">Diagnosis</emphasis>
: The species most closely resembles
<taxonomicName id="75C54D73FF9E4C75FD2ADB9401840311" authority="(SHARP, 1876)" baseAuthorityName="SHARP" baseAuthorityYear="1876" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="80B1EAE2FF9E4C75FD2ADB9401240311" italics="true" pageId="6" pageNumber="35">L. fuscipennis</emphasis>
(
<bibRefCitation id="D6544B01FF9E4C75FEFDDA7701BD0311" author="SHARP, D." box="[275,411,265,289]" pageId="6" pageNumber="35" refId="ref41426" refString="SHARP, D. 1876: Contribution to an insect fauna of the Amazon Valley. Coleoptera, Staphylinidae. - Transactions of the Entomological Society of London: 27 - 424." type="book" year="1876">SHARP, 1876</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="75C54D73FF9E4C75FE34DA7402DA0311" authority="SHARP, 1885" authorityName="SHARP" authorityYear="1885" box="[474,764,265,289]" class="Insecta" family="Staphylinidae" genus="Leptacinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="fusciventris">
<emphasis id="80B1EAE2FF9E4C75FE34DA7402480311" box="[474,622,265,289]" italics="true" pageId="6" pageNumber="35">L. fusciventris</emphasis>
<bibRefCitation id="D6544B01FF9E4C75FD98DA7702DA0311" author="SHARP, D." box="[630,764,265,289]" pageId="6" pageNumber="35" pagination="393 - 536" refId="ref41459" refString="SHARP, D. 1885: Fam. Staphylinidae. - In: Biologia Centrali-Americana. Insecta, Coleoptera. Vol. 1 (2): pp. 393 - 536. - London, Taylor &amp; Francis." type="journal article" year="1885">
SHARP,
<quantity id="753D9B15FF9E4C75FD26DA7702DA0311" box="[712,764,265,289]" metricMagnitude="1" metricUnit="m" metricValue="4.7879" pageId="6" pageNumber="35" unit="in" value="1885.0">1885</quantity>
</bibRefCitation>
</taxonomicName>
in size, colouration and punctation of the head. Females cannot be certainly separated from both species because the main separating characters are found in the shape of sternite VII of male. In
<taxonomicName id="75C54D73FF9E4C75FE37DAF4021E0391" authorityName="Irmler" authorityYear="2021" box="[473,568,393,417]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="80B1EAE2FF9E4C75FE37DAF4021E0391" box="[473,568,393,417]" italics="true" pageId="6" pageNumber="35">L. bierigi</emphasis>
</taxonomicName>
, sternite VII of male has a large process, whereas it is simply rounded in
<taxonomicName id="75C54D73FF9E4C75FF53DAB4016803D1" baseAuthorityName="SHARP" baseAuthorityYear="1876" box="[189,334,457,481]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="80B1EAE2FF9E4C75FF53DAB4016803D1" box="[189,334,457,481]" italics="true" pageId="6" pageNumber="35">L. fuscipennis</emphasis>
</taxonomicName>
and
<taxonomicName id="75C54D73FF9E4C75FE6ADAB4023E03D1" authorityName="SHARP" authorityYear="1885" box="[388,536,457,481]" class="Insecta" family="Staphylinidae" genus="Leptacinus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="species" species="fusciventris">
<emphasis id="80B1EAE2FF9E4C75FE6ADAB4023E03D1" box="[388,536,457,481]" italics="true" pageId="6" pageNumber="35">L. fusciventris</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FF53D97701280571" pageId="6" pageNumber="35" type="description">
<paragraph id="B27A36F0FF9E4C75FF53D97702BB0051" blockId="6.[189,794,521,1857]" pageId="6" pageNumber="35">
<emphasis id="80B1EAE2FF9E4C75FF53D977011B0011" bold="true" box="[189,317,521,545]" pageId="6" pageNumber="35">Description</emphasis>
: Length:
<quantity id="753D9B15FF9E4C75FE72D97701C90011" box="[412,495,521,545]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="6" pageNumber="35" unit="mm" value="5.2">5.2 mm</quantity>
. Colouration: Yellow-brown; base of elytra dark, nearly black; only narrow posterior margin yellow; legs and antennae dark yellow.
</paragraph>
<paragraph id="B27A36F0FF9E4C75FF53D91701280571" blockId="6.[189,794,521,1857]" pageId="6" pageNumber="35">
Head:
<quantity id="753D9B15FF9E4C75FEEBD917014300B1" box="[261,357,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" pageId="6" pageNumber="35" unit="mm" value="0.88">0.88 mm</quantity>
long,
<quantity id="753D9B15FF9E4C75FE44D917022C00B1" box="[426,522,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="6" pageNumber="35" unit="mm" value="0.71">0.71 mm</quantity>
wide; eyes large; slightly prominent; postocular sides slightly divergent posteriad; nearly 3 times as long as eyes; posterior angles rounded; posterior margin slightly emarginate in front of neck; interantennal furrows weak; setiferous punctation moderately dense and deep; on average, interstices between punctures twice as wide as diameter of punctures; on clypeus denser; on posterior vertex sparser; moderately narrow midline impunctate; surface without microsculpture; polished. Antennae with first antennomere as long as half-length of head; second and third antennomere triangular; slightly longer than apical width; subsequent antennomeres wider than long and increasing in width; both fourth and tenth antennomere approximately twice as wide as long; antennomeres four to eleven pubescent; all antennomeres with setae. Pronotum:
<quantity id="753D9B15FF9E4C75FDE3DF17024B06B1" box="[525,621,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="6" pageNumber="35" unit="mm" value="0.95">0.95 mm</quantity>
long,
<quantity id="753D9B15FF9E4C75FD54DF17033C06B1" box="[698,794,1129,1153]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="6" pageNumber="35" unit="mm" value="0.64">0.64 mm</quantity>
wide; widest in anterior third; shortly narrowed toward neck; slightly narrowed in middle third; nearly parallel in posterior third; posterior angles shortly rounded; posterior margin 0.8 times as wide as widest width; Elytra:
<quantity id="753D9B15FF9E4C75FEFEDE7701560711" box="[272,368,1289,1313]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="6" pageNumber="35" unit="mm" value="0.98">0.98 mm</quantity>
long,
<quantity id="753D9B15FF9E4C75FE50DE7702380711" box="[446,542,1289,1313]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="6" pageNumber="35" unit="mm" value="0.77">0.77 mm</quantity>
wide; humeral angles sub-rectangular; sides parallel; posterior angles nearly rectangular; posterior margin widely reflexed to suture; setiferous punctation irregularly dense and moderately deep; on average, interstices between punctures as wide as diameter of punctures but partly coriaceous; surface with weak irregular ground sculpture; moderately matt. Abdomen with weaker and sparser setiferous punctation than pronotum and elytra; without microsculpture; surface shiny; sternite VII of male with long and wide central process at posterior margin; on each side of central process with deep emargination; tergite VII of male straight at posterior margin. Aedeagus long oval; anteriorly sub-rectangular; endophallus with stripe covered by large lobes on one side and minute teeth on other side; paramere slender; slightly curved at apex; on all faces with numerous setae; setae at apex shorter than at base.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF9E4C75FF53DC17021F05F1" pageId="6" pageNumber="35" type="etymology">
<paragraph id="B27A36F0FF9E4C75FF53DC17021F05F1" blockId="6.[189,794,1897,1985]" pageId="6" pageNumber="35">
<emphasis id="80B1EAE2FF9E4C75FF53DC17011005B1" bold="true" box="[189,310,1897,1921]" pageId="6" pageNumber="35">Etymology</emphasis>
: The species name honours the collector A. Bierig, who collected and described many
<taxonomicName id="75C54D73FF9E4C75FD57DCF700DF05F1" authorityName=", HERMAN" authorityYear="2001" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="35" phylum="Arthropoda" rank="family">Staphylinidae</taxonomicName>
from the Neotropical region.
</paragraph>
</subSubSection>
</treatment>
</document>