treatments-xml/data/3A/6C/87/3A6C87E6FF844C6EFC8BDF4D02320451.xml
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<document id="804CCF10C06D063E7481E9E277363970" ID-DOI="10.21248/contrib.entomol.71.1.029-085" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742999" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638270695310" checkinUser="felipe" docAuthor="Irmler, Ulrich" docDate="2021" docId="3A6C87E6FF844C6EFC8BDF4D02320451" docLanguage="en" docName="ContribEntomol.71.1.29-85.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 71 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lithocharodes somoleptoides Irmler 2021, spec. nov." docType="treatment" docVersion="3" lastPageNumber="58" masterDocId="C655FF9EFF984C73FFEEDB7E00260230" masterDocTitle="The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)" masterLastPageNumber="85" masterPageNumber="29" pageNumber="57" updateTime="1699437350205" updateUser="plazi">
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<mods:title id="CC7BD8B794B69FC5B3B736C9E09B8621">The Neotropical species of the genus Lithocharodes SHARP, 1876 (Coleoptera: Staphylinidae: Staphylininae: Xantholinini)</mods:title>
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<mods:namePart id="A96D4BF99A3E17CA02E2807C9708205E">Irmler, Ulrich</mods:namePart>
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<mods:title id="1C6396C4A35BD30625DEDF70CAAD5F75">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="658D5E7EF00D2960F21CF6221E4BE80B">
<mods:date id="1152886F41AC246E912FC826A35389C0">2021</mods:date>
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<mods:number id="3D9FB6B6A2B355F636ED419C492D559F">2021-06-30</mods:number>
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<mods:identifier id="C86F4CD6A714CEF9B83D97A34EC771FD" type="DOI">10.21248/contrib.entomol.71.1.029-085</mods:identifier>
<mods:identifier id="5DFB163A2F8DD7F379C77656BC5D6CB5" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="7E2C10F24CE7D593D47ABF7ED6638E54" type="Zenodo-Dep">5742999</mods:identifier>
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<treatment id="3A6C87E6FF844C6EFC8BDF4D02320451" LSID="urn:lsid:plazi:treatment:3A6C87E6FF844C6EFC8BDF4D02320451" httpUri="http://treatment.plazi.org/id/3A6C87E6FF844C6EFC8BDF4D02320451" lastPageId="29" lastPageNumber="58" pageId="28" pageNumber="57">
<subSubSection id="FADF657BFF844C6FFC8BDF4D054E065F" pageId="28" pageNumber="57" type="nomenclature">
<paragraph id="B27A36F0FF844C6FFC8BDF4D0526067E" blockId="28.[869,1384,1075,1167]" box="[869,1280,1075,1102]" pageId="28" pageNumber="57">
<heading id="E932819CFF844C6FFC8BDF4D0526067E" box="[869,1280,1075,1102]" fontSize="12" level="2" pageId="28" pageNumber="57" reason="2">
<taxonomicName id="75C54D73FF844C6FFC8BDF4D04B6067E" ID-CoL="8Y645" authority="Irmler, 2021" authorityName="Irmler" authorityYear="2021" box="[869,1168,1075,1102]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="somoleptoides" status="sp. nov.">
<emphasis id="80B1EAE2FF844C6FFC8BDF4D04B6067E" box="[869,1168,1075,1102]" italics="true" pageId="28" pageNumber="57">Lithocharodes somoleptoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="9B825799FF844C6FFB78DF4D0526067E" box="[1174,1280,1075,1102]" pageId="28" pageNumber="57" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="B27A36F0FF844C6FFC8BDF25054E065F" blockId="28.[869,1384,1075,1167]" box="[869,1384,1115,1135]" pageId="28" pageNumber="57">
<uri id="C6543AF2FF844C6FFC8BDF25054E065F" box="[869,1384,1115,1135]" pageId="28" pageNumber="57">
urn:lsid:zoobank.org:act:
<uuid id="C6630C25FF844C6FFBFADF25054E065F" box="[1044,1384,1115,1135]" pageId="28" pageNumber="57">0FBFC297-E285-4637-853C-2DC467FE2E1C</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF844C6FFC8BDF05042F06BF" box="[869,1033,1147,1167]" pageId="28" pageNumber="57" type="description">
<paragraph id="B27A36F0FF844C6FFC8BDF05042F06BF" blockId="28.[869,1384,1075,1167]" box="[869,1033,1147,1167]" pageId="28" pageNumber="57">
<figureCitation id="2AFE2A75FF844C6FFC8BDF0503BD06BF" box="[869,923,1147,1167]" captionStart="Fig" captionStartId="45.[151,184,2040,2062]" captionTargetBox="[167,1375,247,2020]" captionTargetId="figure-105@45.[156,1388,232,2032]" captionTargetPageId="45" captionText="Fig. 3742: L.-??-group: Lithocharodes compacta (37), L. elegans (38), L. elongata (39), L. fuscula (40), L. rufula (41); aedeagus (a)," figureDoi="http://doi.org/10.5281/zenodo.5743019" httpUri="https://zenodo.org/record/5743019/files/figure.png" pageId="28" pageNumber="57">Figs 42</figureCitation>
a-c, 45 L, 50 F
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF844C6FFCD1DFD705BF0771" pageId="28" pageNumber="57" type="materials_examined">
<paragraph id="B27A36F0FF844C6FFCD1DFD705BF0771" blockId="28.[831,1437,1193,1345]" pageId="28" pageNumber="57">
<emphasis id="80B1EAE2FF844C6FFCD1DFD703FE06F1" bold="true" box="[831,984,1193,1217]" pageId="28" pageNumber="57">Type material</emphasis>
:
<materialsCitation id="02AD3CADFF844C6FFC09DFD704820711" collectingDate="1992-09-23" collectionCode="NHM" collectorName="R. Anderson" country="Mexico" elevation="2070" location="Solistahuacan" municipality="Yerbabuena Reserve" pageId="28" pageNumber="57" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
male,
<typeStatus id="6D7E8852FF844C6FFBC7DFD704AC06F1" box="[1065,1162,1193,1217]" pageId="28" pageNumber="57" type="holotype">holotype</typeStatus>
:
<collectingCountry id="CAD27660FF844C6FFB7BDFD704C006F1" box="[1173,1254,1193,1217]" name="Mexico" pageId="28" pageNumber="57">Mexico</collectingCountry>
,
<collectingRegion id="7001F812FF844C6FFB1CDFD7056C06F1" box="[1266,1354,1193,1217]" country="Mexico" name="Chiapas" pageId="28" pageNumber="57">Chiapas</collectingRegion>
,
<collectingMunicipality id="521EAC8AFF844C6FFAB8DFD703FD06D1" pageId="28" pageNumber="57">Yerbabuena Reserve</collectingMunicipality>
,
<locationDeviation id="A3C82FD2FF844C6FFC0DDFB704DD06D1" box="[995,1275,1225,1249]" location="Pueblo Nuevo" pageId="28" pageNumber="57">
<quantity id="753D9B15FF844C6FFC0DDFB7040A06D1" box="[995,1068,1225,1249]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="28" pageNumber="57" unit="km" value="2.1">2.1 km</quantity>
NW Pueblo Nuevo
</locationDeviation>
,
<location id="B71A602BFF844C6FFAEDDFB705BE06D1" LSID="urn:lsid:plazi:treatment:3A6C87E6FF844C6EFC8BDF4D02320451:B71A602BFF844C6FFAEDDFB705BE06D1" box="[1283,1432,1225,1249]" country="Mexico" municipality="Yerbabuena Reserve" name="Solistahuacan" pageId="28" pageNumber="57" stateProvince="Chiapas">Solistahuacan</location>
,
<quantity id="753D9B15FF844C6FFCD1DF9703A90731" box="[831,911,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="28" pageNumber="57" unit="m" value="2070.0">
<elevation id="39E8D1C3FF844C6FFCD1DF9703A90731" box="[831,911,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="2.07" pageId="28" pageNumber="57" unit="m" value="2070.0">2070 m</elevation>
</quantity>
elev., Liquidambar litter forest,
<date id="C67B1030FF844C6FFB11DF9705410731" box="[1279,1383,1257,1281]" pageId="28" pageNumber="57" value="1992-09-23">
<collectingDate id="D63FE9D8FF844C6FFB11DF9705410731" box="[1279,1383,1257,1281]" pageId="28" pageNumber="57" value="1992-09-23">23.9.1992</collectingDate>
</date>
, leg.
<collectorName id="1F305326FF844C6FFCD1DE7403E10711" box="[831,967,1289,1313]" pageId="28" pageNumber="57">R. Anderson</collectorName>
#92-114 (
<collectionCode id="D4D4AE35FF844C6FFBB8DE7704BA0711" box="[1110,1180,1289,1313]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="28" pageNumber="57" type="Herbarium">NHM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="02AD3CADFF844C6FFB2EDE7405B30771" collectingDate="1992-09-23" collectionCode="KNHM, UIC" collectorName="R. Anderson" country="Mexico" elevation="2070" location="Solistahuacan" municipality="Yerbabuena Reserve" pageId="28" pageNumber="57" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Chiapas" typeStatus="paratype">
<typeStatus id="6D7E8852FF844C6FFB2EDE74050C0712" box="[1216,1322,1290,1314]" pageId="28" pageNumber="57" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="A4C3FD79FF844C6FFAA8DE7705BF0711" box="[1350,1433,1289,1313]" pageId="28" pageNumber="57" type="male">5 males</specimenCount>
,
<specimenCount id="A4C3FD79FF844C6FFCD1DE5703840771" box="[831,930,1321,1345]" pageId="28" pageNumber="57" type="female">6 females</specimenCount>
with same data as holotype (9
<collectionCode id="D4D4AE35FF844C6FFB04DE5405180771" box="[1258,1342,1322,1345]" country="Kuwait" httpUri="http://grbio.org/cool/k9cv-41mj" name="The Educational Science Museum [=Kuwait Natural History Museum?]" pageId="28" pageNumber="57">KNHM</collectionCode>
, 2
<collectionCode id="D4D4AE35FF844C6FFAB5DE5705AB0771" box="[1371,1421,1321,1345]" pageId="28" pageNumber="57">UIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF844C6FFCD1DE17054F0511" pageId="28" pageNumber="57" type="diagnosis">
<paragraph id="B27A36F0FF844C6FFCD1DE17054F0511" blockId="28.[831,1437,1385,1825]" pageId="28" pageNumber="57">
<emphasis id="80B1EAE2FF844C6FFCD1DE17038A07B1" bold="true" box="[831,940,1385,1409]" pageId="28" pageNumber="57">Diagnosis</emphasis>
: The species highly resembles species of the genus
<taxonomicName id="75C54D73FF844C6FFC68DEF703DB0791" authorityName="SHARP" authorityYear="1885" box="[902,1021,1417,1441]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="80B1EAE2FF844C6FFC68DEF703DB0791" box="[902,1021,1417,1441]" italics="true" pageId="28" pageNumber="57">Somoleptus</emphasis>
</taxonomicName>
by the slender shape. Even the shape of the paramere is similar to that of most
<taxonomicName id="75C54D73FF844C6FFACBDED703B707D1" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="80B1EAE2FF844C6FFACBDED705BA07F1" box="[1317,1436,1449,1473]" italics="true" pageId="28" pageNumber="57">Somoleptus</emphasis>
species.
</taxonomicName>
However, it has distinct interantennal furrows and the specific cone-like structure at the aedeagal orifice are absent. Within the genus
<taxonomicName id="75C54D73FF844C6FFB77DD77050C0411" authorityName="SHARP" authorityYear="1876" box="[1177,1322,1545,1569]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="genus">
<emphasis id="80B1EAE2FF844C6FFB77DD77050C0411" box="[1177,1322,1545,1569]" italics="true" pageId="28" pageNumber="57">Lithocharodes</emphasis>
</taxonomicName>
it mostly resembles
<taxonomicName id="75C54D73FF844C6FFC56DD54043B0471" authorityName="SHARP" authorityYear="1885" baseAuthorityName="SHARP" baseAuthorityYear="1885" box="[952,1053,1577,1601]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="elegans">
<emphasis id="80B1EAE2FF844C6FFC56DD54043B0471" box="[952,1053,1577,1601]" italics="true" pageId="28" pageNumber="57">L. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="75C54D73FF844C6FFB8ADD54052E0471" authorityName="Irmler" authorityYear="2021" box="[1124,1288,1578,1601]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="surinamensis">
<emphasis id="80B1EAE2FF844C6FFB8ADD54052E0471" box="[1124,1288,1578,1601]" italics="true" pageId="28" pageNumber="57">L. surinamensis</emphasis>
</taxonomicName>
in size and overall colouration.
<taxonomicName id="75C54D73FF844C6FFBF3DD37056F0451" authorityName="Irmler" authorityYear="2021" box="[1053,1353,1609,1633]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="somoleptoides">
<emphasis id="80B1EAE2FF844C6FFBF3DD37056F0451" box="[1053,1353,1609,1633]" italics="true" pageId="28" pageNumber="57">Lithocharodes somoleptoides</emphasis>
</taxonomicName>
can be separated from these two species by the short elytra and eyes. Similarly short elytra are also found in
<taxonomicName id="75C54D73FF844C6FFACCDDF4037B04F1" authorityName="Irmler" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="80B1EAE2FF844C6FFACCDDF4037B04F1" italics="true" pageId="28" pageNumber="57">L. curtipennis</emphasis>
</taxonomicName>
, but in
<taxonomicName id="75C54D73FF844C6FFC43DDD4041B04F1" authorityName="Irmler" authorityYear="2021" box="[941,1085,1706,1729]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="80B1EAE2FF844C6FFC43DDD4041B04F1" box="[941,1085,1706,1729]" italics="true" pageId="28" pageNumber="57">L. curtipennis</emphasis>
</taxonomicName>
, elytra are still shorter than wide, whereas they are approximately quadrate in
<taxonomicName id="75C54D73FF844C6FFAACDDB403BF0531" authorityName="Irmler" authorityYear="2021" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="somoleptoides">
<emphasis id="80B1EAE2FF844C6FFAACDDB403BF0531" italics="true" pageId="28" pageNumber="57">L. somoleptoides</emphasis>
</taxonomicName>
. Additionally, the endophallus with its several torsions distinctly differs from that of
<taxonomicName id="75C54D73FF844C6FFB3ADC7405420511" authorityName="Irmler" authorityYear="2021" box="[1236,1380,1802,1825]" class="Insecta" family="Staphylinidae" genus="Lithocharodes" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="57" phylum="Arthropoda" rank="species" species="curtipennis">
<emphasis id="80B1EAE2FF844C6FFB3ADC7405420511" box="[1236,1380,1802,1825]" italics="true" pageId="28" pageNumber="57">L. curtipennis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF844C6EFCD1DC3702D30431" lastPageId="29" lastPageNumber="58" pageId="28" pageNumber="57" type="description">
<paragraph id="B27A36F0FF844C6FFCD1DC37040F05D1" blockId="28.[831,1436,1865,2082]" pageId="28" pageNumber="57">
<emphasis id="80B1EAE2FF844C6FFCD1DC3703E70551" bold="true" box="[831,961,1865,1889]" pageId="28" pageNumber="57">Description</emphasis>
: Length:
<quantity id="753D9B15FF844C6FFBAADC3704BE0551" box="[1092,1176,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="28" pageNumber="57" unit="mm" value="4.7">4.7 mm</quantity>
. Colouration: Brown, pronotum lighter brown than head and elytra; abdominal segments lighter brown than elytra; darkened posteriad except segment VII, being again light brown; legs and antennae light brown.
</paragraph>
<paragraph id="B27A36F0FF844C6EFCD1DC9702D30431" blockId="28.[831,1436,1865,2082]" lastBlockId="29.[151,757,233,1537]" lastPageId="29" lastPageNumber="58" pageId="28" pageNumber="57">
Head:
<quantity id="753D9B15FF844C6FFC64DC9703CF0A31" box="[906,1001,2025,2049]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="28" pageNumber="57" unit="mm" value="0.81">0.81 mm</quantity>
long,
<quantity id="753D9B15FF844C6FFBDADC9704B50A31" box="[1076,1171,2025,2049]" metricMagnitude="-4" metricUnit="m" metricValue="6.1" pageId="28" pageNumber="57" unit="mm" value="0.61">0.61 mm</quantity>
wide; eyes small; postocular sides nearly 5 times as long as eyes; slightly divergent posteriad; posterior angles widely rounded; shape of posterior head nearly semi-circular; interantennal furrows reaching anterior third of eyes; setiferous punctation deep and moderately dense; on average, interstices at least as wide as diameter of punctures; narrow midline impunctate; surface with weak isodiametric microsculpture; moderately shiny. Antennae with first antennomere as long as half-length of head; second and third antennomere much longer than wide; both antennomeres combined approximately half as long as first antennomere; subsequent antennomeres much shorter, wider than long, and increasing in width; fourth antennomere twice as wide as long; tenth antennomere 2.5 times as wide as long; antennomeres four to eleven pubescent. Pronotum:
<quantity id="753D9B15FF854C6EFEFFD9D7015600F1" box="[273,368,681,705]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="29" pageNumber="58" unit="mm" value="0.94">0.94 mm</quantity>
long,
<quantity id="753D9B15FF854C6EFE5DD9D7023700F1" box="[435,529,681,705]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="29" pageNumber="58" unit="mm" value="0.59">0.59 mm</quantity>
wide; widest at anterior third; at widest width only weakly wider than at posterior margin; posterior angles obtuse; sub-rectangular; posterior margin slightly convexly curved; setiferous punctation on average deeper and distinctly denser than on head; wide midline impunctate; irregular line adjacent to midline with approximately 19 punctures; punctures of line partly coriaceous in posterior half; surface with weak isodiametric microsculpture; moderately shiny. Elytra:
<quantity id="753D9B15FF854C6EFF05D8B7016F01D1" box="[235,329,969,993]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="29" pageNumber="58" unit="mm" value="0.68">0.68 mm</quantity>
long,
<quantity id="753D9B15FF854C6EFE77D8B701D101D1" box="[409,503,969,993]" metricMagnitude="-4" metricUnit="m" metricValue="6.7" pageId="29" pageNumber="58" unit="mm" value="0.67">0.67 mm</quantity>
wide; humeral angles absent; sides distinctly divergent to posterior angles; posterior angles rectangular; posterior margin deeply retreated to suture forming triangular emargination; setiferous punctation deeper and denser than on head and pronotum; on average, interstices between punctures less than half as wide as diameter of punctures; isodiametric microsculpture even weaker than on head and pronotum; moderately shiny. Abdomen with weaker and sparser setiferous punctation than on fore-body; surface with transversely reticulate microsculpture; posterior margin of sternite VII of male semi-circularly prominent, posterior margin of tergite VII of male nearly straight. Meso- and metatibia with one subapical ctenidium each. Aedeagus oval; divergent posteriad; without anterior and posterior angles; endophallus as band with several torsions; covered by numerous medium-sized teeth; paramere slender; curved in apical third; with few short setae.
</paragraph>
</subSubSection>
<subSubSection id="FADF657BFF854C6EFF79DD5702320451" pageId="29" pageNumber="58" type="etymology">
<paragraph id="B27A36F0FF854C6EFF79DD5702320451" blockId="29.[151,756,1577,1634]" pageId="29" pageNumber="58">
<emphasis id="80B1EAE2FF854C6EFF79DD5701280471" bold="true" box="[151,270,1577,1601]" pageId="29" pageNumber="58">Etymology</emphasis>
: The species name refers to the high similarity in shape with the genus
<taxonomicName id="75C54D73FF854C6EFE79DD3702280451" authorityName="SHARP" authorityYear="1885" box="[407,526,1609,1633]" class="Insecta" family="Staphylinidae" genus="Somoleptus" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="80B1EAE2FF854C6EFE79DD3702280451" box="[407,526,1609,1633]" italics="true" pageId="29" pageNumber="58">Somoleptus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>