plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/3A/4F/67/3A4F67A717BD5ACEB47D4E7C567A01EC.xml
maintenance 5e4b328909 added 3A
2024-06-21 12:33:50 +02:00

734 lines
119 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1132.94579" ID-Pensoft-Pub="1313-2970-1132-1" ID-Pensoft-UUID="A682E06DC8B3510EBB1DB1F926E99F45" ID-ZooBank="D0144E9FA4854450B279A813F3EF0AEF" ModsDocID="1313-2970-1132-1" checkinTime="1669467072147" checkinUser="pensoft" docAuthor="Rohacek, Jindrich &amp; Przhiboro, Andrey A." docDate="2022" docId="3A4F67A717BD5ACEB47D4E7C567A01EC" docLanguage="en" docName="ZooKeys 1132: 1-49" docOrigin="ZooKeys 1132" docPubDate="2022-11-25" docSource="http://dx.doi.org/10.3897/zookeys.1132.94579" docTitle="Pullimosina (Pullimosina) turfosa Roháček &amp; Przhiboro 2022, sp. nov." docType="treatment" docUuid="3D55B25A-F2A7-4FAF-ACC2-D189349E7AFC" docUuidSource="ZooBank" docVersion="1" id="A682E06DC8B3510EBB1DB1F926E99F45" lastPageNumber="1" masterDocId="A682E06DC8B3510EBB1DB1F926E99F45" masterDocTitle="Pullimosina (Pullimosina) turfosa sp. nov. and other Sphaeroceridae (Diptera) from peat bogs in the North Caucasus (Russia)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="1" updateTime="1669467072147" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Pullimosina (Pullimosina) turfosa sp. nov. and other Sphaeroceridae (Diptera) from peat bogs in the North Caucasus (Russia)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rohacek, Jindrich</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3311-2087</mods:nameIdentifier>
<mods:affiliation>Silesian Museum, Nadrazni okruh 31, CZ- 746 01 Opava, Czech Republic</mods:affiliation>
<mods:nameIdentifier type="email">rohacek@szm.cz</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Przhiboro, Andrey A.</mods:namePart>
<mods:affiliation>Zoological Institute, Russian Academy of Sciences, St Petersburg, 199034 Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-11-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1132</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>49</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1132.94579</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1132.94579</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1132-1</mods:identifier>
<mods:identifier type="ZooBank">D0144E9FA4854450B279A813F3EF0AEF</mods:identifier>
<mods:identifier type="Pensoft-UUID">A682E06DC8B3510EBB1DB1F926E99F45</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:zoobank.org:act:3D55B25A-F2A7-4FAF-ACC2-D189349E7AFC" httpUri="http://treatment.plazi.org/id/3A4F67A717BD5ACEB47D4E7C567A01EC" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/3D55B25A-F2A7-4FAF-ACC2-D189349E7AFC" authority="Roháček &amp; Przhiboro, 2022" authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" status="sp. nov." subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">Figs 12-13</figureCitation>
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">, 14-17</figureCitation>
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">, 18-22</figureCitation>
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">, 23-26</figureCitation>
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">, 27-30</figureCitation>
<figureCitation captionStart="Figures 31–32" captionStartId="F12" captionText="Figures 31 - 32. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 31 whole body, dorsally 32 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures31-32" httpUri="https://binary.pensoft.net/fig/774337" pageId="0" pageNumber="1">, 31-32</figureCitation>
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">, 33-39</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
♂ labelled: &quot;Russia: N Ossetia, W Digoria, Chifandzar mire in Kharesidon River valley,
<geoCoordinate degrees="42.91867" direction="north" orientation="latitude" precision="1" value="42.91867">42.91867°N</geoCoordinate>
,
<geoCoordinate degrees="43.51493" direction="east" orientation="longitude" precision="1" value="43.51493">43.51493°E</geoCoordinate>
, 2289 m, sifting from
<taxonomicName authorityName="Angstrom in C. J. Hartman" authorityYear="1861" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum teres" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="teres">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum teres</emphasis>
</taxonomicName>
hummocks, 18.ix.2018, A. Przhiboro leg.&quot;, &quot;Holotypus ♂
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. n., J.
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
det. 2022&quot; (red label). The specimen is dried from ethanol and mounted on pinned triangular card, intact (deposited in ZISP, Figs
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">12</figureCitation>
,
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">13</figureCitation>
).
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: 6♂ 5♀ with same locality labels but with &quot;Paratypus [♂ or ♀],
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. n., J.
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
det. 2022&quot; yellow labels; 3♂ 1♀ paratypes preserved in pinned microvial in glycerin, with abdomen detached, and terminalia dissected; others dry-mounted from ethanol and pinned as is the holotype; 1♂ 1♀ with wing removed for photography and also preserved in glycerin in pinned plastic tube below the specimen (4♂ 3♀ in ZISP, 2♂ 2♀ in SMOC). Other paratypes: 1♀, same locality data, but with &quot;Sample
<normalizedToken originalValue="Ч">CH</normalizedToken>
14 (
<taxonomicName authorityName="Angstrom in C. J. Hartman" authorityYear="1861" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum teres" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="teres">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum teres</emphasis>
</taxonomicName>
), 17.ix.2018&quot;; 1♀, same locality data, but with &quot;Sample
<normalizedToken originalValue="Ч">CH</normalizedToken>
9 (
<taxonomicName authorityName="Nees in Sturm" authorityYear="1819" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum subsecundum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subsecundum">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum subsecundum</emphasis>
</taxonomicName>
), 17.ix.2018&quot;, both A. Przhiboro leg. (ZISP).
</paragraph>
<caption doi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1" start="Figures 12, 13" startId="F7">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 12, 13.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (male holotype)
<emphasis bold="true" pageId="0" pageNumber="1">12</emphasis>
whole body, dorsally
<emphasis bold="true" pageId="0" pageNumber="1">13</emphasis>
ditto, laterally. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
This strongly brachypterous species is named
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina turfosa</emphasis>
</taxonomicName>
(= peaty, Latin adjective) owing to its strict association with
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum</emphasis>
</taxonomicName>
hummocks in the type locality.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Male</emphasis>
(Figs
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">12</figureCitation>
,
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">13</figureCitation>
). Total body length 1.27-1.64 (holotype 1.64) mm; general color brown to dark brown with greyish brown microtomentum, subshining dorsally (thorax, abdomen) and ventrally (abdomen), dull on thoracic pleuron.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Head</emphasis>
(Figs
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">13</figureCitation>
,
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">14</figureCitation>
) higher than long (ca. 5:4), bicolorous, dorsally and posteriorly brown to pale brown, anteriorly and ventrally yellow to pale ochreous. Frons brown to pale brown, with anterior margin and orbits pale ochreous (Figs
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">14</figureCitation>
,
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">16</figureCitation>
), sparsely microtomentose and partly (mainly medially) shining; occiput dark brown with brownish grey microtomentum. Orbits, interfrontalia and ocellar triangle with paler greyish microtomentum; orbit separated from interfrontalia by dark brown dull stripe (shortened anteriorly, never reaching anterior margin of frons); frontal triangle indistinctly delimited but long, almost reaching anterior margin of frons, finely longitudinally microsculptured (Fig.
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">14</figureCitation>
), and more shining than rest of frons. Cephalic chaetotaxy (cf. Figs
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">12</figureCitation>
-
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">14</figureCitation>
,
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">16</figureCitation>
): pvt present but reduced, hair-like but convergent and with apices almost meeting medially; occe and occi subequal (or occi slightly longer) and ca. two-thirds to three-fourths length of vti; vti normally thickest and longest of frontal setae; vte and oc only slightly shorter than vti; 2 ors, posterior almost as long as vte (or oc) and only slightly longer than anterior ors; 4 ifr, none markedly enlarged, middle 2 pairs usually longer than posterior pair, foremost pair small, about ca. half the length of the previous pair; 1 microseta in front of anterior margin of frons, lateral to foremost ifr; 4-6 minute ads inside and below ors; g small, ca. as long as foremost peristomal setula and 1 or 2 short setae behind it; vi robust, ca. as long as vti; peristomal setulae (5-6) slightly longer than those in single postocular row; 3 postgenal setae, all relatively strong and curved. Frontal lunule of moderate length, well-developed, yellow and sparsely whitish microtomentose, slightly paler than anterior margin of frons. Face yellow, sparsely whitish microtomentose but facial cavities below antennae relatively shining; medial carina small, most distinct dorsally, below frontal lunule. Parafacialia darker than face, ochreous brown. Gena yellow, somewhat darkened only at vibrissal angle and very narrowly on ventral margin, all sparsely whitish microtomentose and rather dull. Postgena brown, sharply delimited from gena. Mouthparts ochreous to brownish including clypeus. Palpus yellowish, slender but distinctly clavate (Fig.
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">14</figureCitation>
), with ca. 5 dark setae (subapical longest) along ventral margin. Eye broadly suboval (9:8), of moderate size, with longest diameter ~ 6.0
<normalizedToken originalValue="×">x</normalizedToken>
as long as smallest genal height. Antenna brown (1st flagellomere) to dark brown (scape and pedicel); 1st flagellomere ca. as long as scape + pedicel, ellipsoid, with short greyish ciliation on apex (not longer than cilia on arista). Arista ~ 3.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as antenna, shortly but densely ciliate.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Thorax</emphasis>
brown to pale brown (pleuron paler) and greyish brown microtomentose; mesonotum subshining, pleuron and scutellum more densely microtomentose and duller (Figs
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">12</figureCitation>
,
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">13</figureCitation>
). Mesonotum laterally (notopleural area) and posteriorly (in front of scutellum) paler, usually ochreous; scutellum also somewhat paler posteromedially. Thoracic pleuron with propleuron and sternopleuron largely pale brown to ochreous, other sclerites more or less ochreous margined. Scutellum large, transversely (8:5) rounded, trapezoidal, flat on disc. Thoracic chaetotaxy: mesonotal macrosetae relatively short and weak; 1 hu and 2 microsetae on humeral callus; 3 postsutural dc but the foremost very small (less than twice as long as dc microseta in front of it), the middle dc weak, ca. half the length of posterior, the latter long, ca. as long as scutellum; 6 rows of ac microsetae on suture; medial prescutellar ac pair distinctly prolonged, only slightly shorter than middle dc; 2 long sc, laterobasal ~ 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as scutellum, apical (longest thoracic seta) ~ 1.4
<normalizedToken originalValue="×">x</normalizedToken>
as long as laterobasal; 2 stpl but anterior reduced to very small setula, sometimes indistinct.
</paragraph>
<caption doi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1" start="Figures 14–17" startId="F8">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 14-17.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (holotype, paratypes) heads and wings
<emphasis bold="true" pageId="0" pageNumber="1">14</emphasis>
male head, frontally (holotype)
<emphasis bold="true" pageId="0" pageNumber="1">15</emphasis>
male right wing, dorsally (paratype)
<emphasis bold="true" pageId="0" pageNumber="1">16</emphasis>
female head, frontally (paratype)
<emphasis bold="true" pageId="0" pageNumber="1">17</emphasis>
female right wing, dorsally (paratype). Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">14, 16</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">15, 17</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Legs</emphasis>
brown to pale brown, coxae, trochanters and knees ochreous to yellow; fore coxa and all trochanters lightest, dirty yellow. Chaetotaxy: f1 with a posterodorsal row of 6 or 7 shorter setae and a posteroventral row of 7 or 8 longer setae in addition to ventrobasal fine seta (Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">18</figureCitation>
). f2 ventrally uniformly setulose but with 3 anterodorsal setae in distal third, including longest subapical seta (Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">20</figureCitation>
). t2 (as in most European congeners) ventrally with 1 short and weak seta below middle (in distal two-fifths), 1 longer (but also relatively short) va seta and 1 small anteroapical seta (see Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">20</figureCitation>
); dorsally with only 4 setae, viz. 1 anterodorsal seta in proximal third, 1 anterodorsal seta in distal third, 1 long dorsal (most robust) seta in distal sixth and 1 small posterodorsal seta in distal fifth (Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">19</figureCitation>
). Hind leg, including f3, uniformly setulose. Ratio t2: mt2 = 2.17-2.30 (holotype 2.17).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Wing</emphasis>
(Figs
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">15</figureCitation>
,
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">22</figureCitation>
) strongly reduced, only ca. twice as long as scutellum, racket-shaped, with brownish membrane, most darkened around R2+3 and M; veins brown- to pale-pigmented. Distal radial and anal part of wing strongly reduced, thus R4+5 and A1 entirely absent. Basal part of C (= Cs1) well developed, including both breaks; distal part of C abbreviated (only Cs2 developed) so that C only slightly produced beyond apex of R2+3. Subcosta absent but presence of humeral (h) cross-vein indicated by darkened stump in front of humeral break (Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">22</figureCitation>
). Basal stem of radial veins robust but R1 short, pale pigmented and poorly visible (Fig.
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">15</figureCitation>
); R2+3 dark brown, very slightly to distinctly upcurved to C. M present, dark brown, forming anterior remnant of discal cell (Figs
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">15</figureCitation>
,
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">22</figureCitation>
); CuA1 strongly reduced, only indicated by a darkening near base of M. Anal lobe and hence also A1 absent; alula distinct but very narrow. Wing measurements: length 0.36-0.52 (holotype 0.52) mm, width 0.18-0.26 (holotype 0.26) mm, Cs1: Cs2 = 1.80-2.27 (holotype 1.80). Haltere present but strongly reduced (see Fig.
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">12</figureCitation>
), with knob entirely absent and stem shortened (length of haltere remnant 0.09-0.11 mm), dirty yellow.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Abdomen</emphasis>
(Figs
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">12</figureCitation>
,
<figureCitation captionStart="Figures 12, 13" captionStartId="F7" captionText="Figures 12, 13. Pullimosina (Pullimosina) turfosa sp. nov. (male holotype) 12 whole body, dorsally 13 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures12-13" httpUri="https://binary.pensoft.net/fig/774332" pageId="0" pageNumber="1">13</figureCitation>
) darker brown dorsally, paler (mainly anteriorly) brown ventrally. Preabdominal terga broad, transversely suboblong, and relatively shining because of sparse greyish brown microtomentum. T2-T5 sparsely but relatively long-setose, with longest setae in posterior corners and margins. T1+2 largest tergum, ~ 1.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as T3, simply sclerotized (without medial depression) but original T1 pale brown to ochreous and distinctly delimited from original T2 (being dark brown) by a transverse wrinkle. T3-T5 subequal in length but becoming slightly narrower posteriorly, T5 smallest. Preabdominal sterna: S1+2 small, reduced to pale and bare poorly delimited sclerite; S3 and S4 subequal in length, relatively large and broad (becoming wider posteriorly), brown and well-sclerotized; both S3 and S4 transversely trapezoidal, narrower anteriorly, but S3 distinctly smaller than S4, the latter smaller and narrower than S5. S3 and S4 with shorter and finer setae than adjacent terga. S5 (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">23</figureCitation>
) darker brown than S3 or S4, more transverse, slightly asymmetrical (longer on left), with short posterior submembranous, unpigmented and finely haired margin and with a transverse group of robust setae, those in the middle particularly thickened, spine-like. Postabdominal sclerites S6+7 and S8 forming a relatively long complex synsclerite situated left ventrolaterally to dorsolaterally (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">23</figureCitation>
). S6+7 strongly asymmetrical, with various projections and placed ventrolaterally to laterally; S8 less asymmetrical and situated more dorsally. Synsclerite S6+7 with original S6 attenuated right ventrally and bearing a distinctive subtriangular posteromedial (in medial axis of abdomen) process (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">23</figureCitation>
), left ventrally dilated, without setae; original S7 ventrolaterally incised and with unusual slender T-shaped projection arising near this incision and directed right medially/internally (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">23</figureCitation>
); left compact part of S7 with 2 pairs of relatively long and stout setae. S8 relatively simple, saddle-shaped, with only a few (3-5) shorter setae, mainly situated at posterior margin.
</paragraph>
<caption doi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1" start="Figures 18–22" startId="F9">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 18-22.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (paratypes)
<emphasis bold="true" pageId="0" pageNumber="1">18</emphasis>
male fore leg without tarsus, posteriorly
<emphasis bold="true" pageId="0" pageNumber="1">19</emphasis>
male mid tibia, dorsally
<emphasis bold="true" pageId="0" pageNumber="1">20</emphasis>
male mid femur, tibia and basitarsus, anteriorly
<emphasis bold="true" pageId="0" pageNumber="1">21</emphasis>
female mid tibia and basitarsus, anteriorly
<emphasis bold="true" pageId="0" pageNumber="1">22</emphasis>
male right wing, dorsally. Scale bars: 0.2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">18-21</emphasis>
); 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">22</emphasis>
).
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1" start="Figures 23–26" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 23-26.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (male paratype)
<emphasis bold="true" pageId="0" pageNumber="1">23</emphasis>
S5 and postabdominal sclerites, ventrally
<emphasis bold="true" pageId="0" pageNumber="1">24</emphasis>
gonostylus, laterally
<emphasis bold="true" pageId="0" pageNumber="1">25</emphasis>
external genitalia, caudally
<emphasis bold="true" pageId="0" pageNumber="1">26</emphasis>
ditto, laterally. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">23, 25, 26</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="1">24</emphasis>
). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genitalia</emphasis>
: Epandrium (Figs
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">25</figureCitation>
,
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">26</figureCitation>
) of medium length and width, very slightly asymmetrical in caudal view (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">25</figureCitation>
), rather uniformly setose (longest setae postero-ventrally but sometimes also 1 dorsolateral seta enlarged). Anal fissure not large, roughly hexagonal, higher than wide (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">25</figureCitation>
). Cerci short, fused with epandrium and medially forming subanal plate being ventromedially deeply narrowly incised (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">25</figureCitation>
); each cercus with 1 longer and 2 or 3 short setae, micropubescent. Medandrium subquadrate in caudal view but its posterior part Y-shaped, hence ventrally narrowed (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">25</figureCitation>
), posteromedially fused with cerci and posteroventrally movably connected with gonostyli. Hypandrium roughly Y-shaped in dorsal view, with simple anteromedial rod-like apodeme, relatively robust paired lateral sclerites, and more medially with small sclerites connecting hypandrium with postgonites via remnants of pregonites. Gonostylus (Figs
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">24-26</figureCitation>
) very distinctive, of unusual (in
<taxonomicName class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina</emphasis>
</taxonomicName>
) shape: dorsally with small and low lateral part overgrown with a tuft of long sinuous setae and some micropubescence; anteroventrally (and more medially) protruding into a slender and long, slightly bent, apically blunt and shortly setulose projection. Aedeagal complex (Figs
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">27-30</figureCitation>
). Phallapodeme distinctly longer and more robust than hypandrial apodeme, with well-developed dorsal keel. Aedeagus composed of compact, laterally flattened phallophore (Figs
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">27</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">28</figureCitation>
) and relatively short distiphallus. Distiphallus basally with slender arcuate sclerite bent on lateral sides (Figs
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">27</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">28</figureCitation>
) and dilated ventrally; the latter dorsally connected with slender sclerite projecting anteriorly where bearing small wing-like processes and longer medial projection almost reaching apex of distiphallus (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">27</figureCitation>
); distal part of distiphallus formed by large trough-like lateroventral sclerite and by a pair of apical dorsal sclerites, each of which having a group of 4 or 5 short dark spines attached laterally (Figs
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">27</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">28</figureCitation>
). Postgonite (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">29</figureCitation>
) relatively large (somewhat longer than distiphallus) but simple, wider proximally and gradually tapered distally, slightly bent and with acute apex, with only 2 or 3 microsetae anteriorly and posteriorly in distal half and fourth, respectively. Remnant of pregonite (Figs
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">29</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">30</figureCitation>
) forming small but distinct and separate sclerite situated in anterodorsal emargination of postgonite, possessing distally 2 short blunt spines and 1 setula (see Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">30</figureCitation>
). Ejacapodeme reduced, represented by small and very slender, rod-like but proximally somewhat dilated, sclerite (Figs
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">27</figureCitation>
,
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">30</figureCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Female</emphasis>
(Figs
<figureCitation captionStart="Figures 31–32" captionStartId="F12" captionText="Figures 31 - 32. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 31 whole body, dorsally 32 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures31-32" httpUri="https://binary.pensoft.net/fig/774337" pageId="0" pageNumber="1">31</figureCitation>
,
<figureCitation captionStart="Figures 31–32" captionStartId="F12" captionText="Figures 31 - 32. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 31 whole body, dorsally 32 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures31-32" httpUri="https://binary.pensoft.net/fig/774337" pageId="0" pageNumber="1">32</figureCitation>
). Similar to male unless mentioned otherwise below. Total body length 1.27-1.67 mm. Foremost ifr more robust, often almost as long as other ifr setae. t2 with all macrosetae relatively longer, both ventrally (cf. ventroapical seta on Figs
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">20</figureCitation>
and
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">21</figureCitation>
) and dorsally. mt2 relatively (compared to t2) longer (Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">21</figureCitation>
). Ratio t2: mt2 = 1.92-2.09. Wing (Fig.
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">17</figureCitation>
) slightly shorter on the average and often with more cut apex. Remnant of haltere also shorter, only 0.06-0.08 mm long. Wing measurements: length 0.36-0.43 mm, width 0.19-0.25 mm, Cs1: Cs2 = 2.14-3.00. Preabdominal terga somewhat shorter and more transverse (Fig.
<figureCitation captionStart="Figures 31–32" captionStartId="F12" captionText="Figures 31 - 32. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 31 whole body, dorsally 32 ditto, laterally. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1132.94579.figures31-32" httpUri="https://binary.pensoft.net/fig/774337" pageId="0" pageNumber="1">32</figureCitation>
); T1+2 only slightly shorter than T3; T3-T5 becoming distinctly narrower posteriorly but similarly setose as in male. Preabdominal sterna S3-S5 sparsely and shortly setose, subequal in length and width. S5 unmodified, transversely suboblong, subequal to S4; preabdominal sterna S3-S5 brown, well sclerotized but paler than adjacent terga.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Postabdomen</emphasis>
(Figs
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">34-36</figureCitation>
) relatively short and broad, with sparsely setose sclerites, narrower than preabdomen at 5th segment. T6 markedly narrower and only ca. half the length of T5, transverse, only slightly wider than S6, with both lateral and posterior margins pale and setose in posterior half (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">34</figureCitation>
), setae at posterior margin long; T7 transversely suboblong, slightly shorter and seemingly narrower than T6 because bent farther onto lateral side (see Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">36</figureCitation>
), with pale posterior margin and 8 setae in single row of setae in front of it. T8 dorsomedially narrowly interrupted to form two lateral sclerites (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">34</figureCitation>
), each dorsally shortened but ventrally expanded and longer than other postabdominal sclerites (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">36</figureCitation>
) and bearing 1 long and a few short to small setae. T10 transversely pentagonal, distinctly wider than long, pale-pigmented, finely sparsely micropubescent and with a pair of relatively distant setae (see Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">34</figureCitation>
). S6 slightly narrower but distinctly (0.7
<normalizedToken originalValue="×">x</normalizedToken>
as long as) shorter than S5, and only slightly wider and more setulose than S7 (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">35</figureCitation>
). S7 simple, transversely suboblong (as is S6), slightly wider than T6, with setae only at pale posterior margin. S8 (Figs
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">35</figureCitation>
,
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">37</figureCitation>
) transversely subellipsoid, much larger than S10 (in largest extension view, see Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">37</figureCitation>
), somewhat convex in the middle, posteriorly more rounded than anteriorly, with only 4 or 6 short setae centrally but with distinctive micropubescence. Additional sclerite unusual, situated behind and partly under S8 (its anterior part overlapped by S8, cf. Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">36</figureCitation>
, asc), narrowly trapezoidal but anteriorly membranous and hence its anterior margin undefined, largely bare, with only 4 setulae at posterior margin (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">37</figureCitation>
). S10 slightly more than half length of S8, transversely pentagonal, pale pigmented, micropubescent and setulose only in posterior third, posteromedially with a pair of longer setae (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">35</figureCitation>
). Spectacles-shaped sclerite (= sclerotization of female genital chamber) oriented rather vertically (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">38</figureCitation>
, see in situ, Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">36</figureCitation>
), with rings of moderate size and its medial anterior sclerotization relatively complex (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">39</figureCitation>
). Spermathecae 2+1 (Figs
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">33</figureCitation>
,
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">39</figureCitation>
), blackish brown; body of single spermatheca distinctly larger than those of paired ones; each spermatheca of relatively robust tyre-shaped form, most resembling those of
<taxonomicName lsidName="P. moesta" pageId="0" pageNumber="1" rank="species" species="moesta">
<emphasis italics="true" pageId="0" pageNumber="1">P. moesta</emphasis>
</taxonomicName>
(Villeneuve, 1918), with plain surface, terminal invagination somewhat widened internally and terminal parts of ducts well-sclerotized, slightly conically dilated towards insertion and ca. as long as body of spermatheca. Cerci (Figs
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">34-36</figureCitation>
) short but not robust, tapered both towards base and terminal seta, micropubescent, each with 4 or 5 setae, apical one longest (slightly longer than cercus) and sinuate as also is the shorter dorsopreapical seta.
</paragraph>
<caption doi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1" start="Figures 27–30" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 27-30.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (male paratype)
<emphasis bold="true" pageId="0" pageNumber="1">27</emphasis>
aedeagus (phallus) dorsally
<emphasis bold="true" pageId="0" pageNumber="1">28</emphasis>
ditto, laterally
<emphasis bold="true" pageId="0" pageNumber="1">29</emphasis>
aedeagal complex, laterally
<emphasis bold="true" pageId="0" pageNumber="1">30</emphasis>
pregonite (enlarged), laterally. Scale bars 0.05 mm (27-29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1132.94579.figures31-32" httpUri="https://binary.pensoft.net/fig/774337" pageId="0" pageNumber="1" start="Figures 31–32" startId="F12">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 31-32.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (female paratype)
<emphasis bold="true" pageId="0" pageNumber="1">31</emphasis>
whole body, dorsally
<emphasis bold="true" pageId="0" pageNumber="1">32</emphasis>
ditto, laterally. Scale bar: 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
Despite a number of peculiarities in the male and female terminalia and unusual reduction of wing venation,
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina turfosa</emphasis>
</taxonomicName>
sp. nov. clearly is a representative of the subgenus
<taxonomicName authority="Pullimosina" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina)" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="subGenus" subGenus="Pullimosina">Pullimosina Pullimosina</taxonomicName>
s. str. (
<bibRefCitation author="Rohacek, J" journalOrPublisher="Beitraege zur Entomologie, Berlin" pageId="0" pageNumber="1" pagination="3 - 195" publicationUrl="https://www.contributions-to-entomology.org/article/view/1184/1183" refId="B56" refString="Rohacek, J, 1983. A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II. Beitraege zur Entomologie, Berlin 33: 3 - 195, https://www.contributions-to-entomology.org/article/view/1184/1183" title="A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II." url="https://www.contributions-to-entomology.org/article/view/1184/1183" volume="33" year="1983">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
1983
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
). However, it proved not to be closely related to any other described European (or Palaearctic) species of this subgenus (cf.
<bibRefCitation author="Papp, L" journalOrPublisher="Acta Zoologica Academiae Scientiarum Hungaricae" pageId="0" pageNumber="1" pagination="369 - 425" refId="B38" refString="Papp, L, 1973. Sphaeroceridae (Diptera) from Mongolia. Acta Zoologica Academiae Scientiarum Hungaricae 19: 369 - 425" title="Sphaeroceridae (Diptera) from Mongolia." volume="19" year="1973">Papp 1973</bibRefCitation>
;
<bibRefCitation author="Rohacek, J" journalOrPublisher="Beitraege zur Entomologie, Berlin" pageId="0" pageNumber="1" pagination="3 - 195" publicationUrl="https://www.contributions-to-entomology.org/article/view/1184/1183" refId="B56" refString="Rohacek, J, 1983. A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II. Beitraege zur Entomologie, Berlin 33: 3 - 195, https://www.contributions-to-entomology.org/article/view/1184/1183" title="A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part II." url="https://www.contributions-to-entomology.org/article/view/1184/1183" volume="33" year="1983">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
1983
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.7601/mez.57.265" author="Hayashi, T" journalOrPublisher="Japanese Journal of Sanitary Zoology" pageId="0" pageNumber="1" pagination="265 - 272" refId="B22" refString="Hayashi, T, 2006. The genus Pullimosina Rohacek (Diptera, Sphaeroceridae) from Japan. Japanese Journal of Sanitary Zoology 57 (4): 265 - 272, DOI: https://doi.org/10.7601/mez.57.265" title="The genus Pullimosina Rohacek (Diptera, Sphaeroceridae) from Japan." url="https://doi.org/10.7601/mez.57.265" volume="57" year="2006">Hayashi 2006</bibRefCitation>
;
<bibRefCitation author="Su, L" journalOrPublisher="Liaoning University Press, Shenyang, Liaoning, China" pageId="0" pageNumber="1" refId="B76" refString="Su, L, 2011. Lesser Dung Flies. Liaoning University Press, Shenyang, Liaoning, China" title="Lesser Dung Flies." year="2011">Su 2011</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.33338/ef.84594" author="Su, L-X" journalOrPublisher="Entomologica Fennica" pageId="0" pageNumber="1" pagination="1 - 8" refId="B77" refString="Su, L-X, Liu, G-C, Xu, J, 2013. Genus Pullimosina (Diptera: Sphaeroceridae) in China with description of a new species. Entomologica Fennica 24 (1): 1 - 8, DOI: https://doi.org/10.33338/ef.84594" title="Genus Pullimosina (Diptera: Sphaeroceridae) in China with description of a new species." url="https://doi.org/10.33338/ef.84594" volume="24" year="2013">Su et al. 2013</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.2478/aemnp-2019-0009" author="Rohacek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="1" pagination="107 - 124" refId="B65" refString="Rohacek, J, 2019. First Sphaeroceridae (Diptera) endemic to Madeira - three new terricolous species of Spelobia and Pullimosina. Acta Entomologica Musei Nationalis Pragae 59 (1): 107 - 124, DOI: https://doi.org/10.2478/aemnp-2019-0009" title="First Sphaeroceridae (Diptera) endemic to Madeira - three new terricolous species of Spelobia and Pullimosina." url="https://doi.org/10.2478/aemnp-2019-0009" volume="59" year="2019">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
2019
</bibRefCitation>
). Based on structures of its male and female terminalia it surely belongs to the
<taxonomicName baseAuthorityName="Duda" baseAuthorityYear="1918" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina antennata" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="antennata">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina antennata</emphasis>
</taxonomicName>
group (as defined by
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
). Note: this group should be re-named to
<taxonomicName lsidName="P. moesta" pageId="0" pageNumber="1" rank="species" species="moesta">
<emphasis italics="true" pageId="0" pageNumber="1">P. moesta</emphasis>
</taxonomicName>
group because
<taxonomicName lsidName="P. antennata" pageId="0" pageNumber="1" rank="species" species="antennata">
<emphasis italics="true" pageId="0" pageNumber="1">P. antennata</emphasis>
</taxonomicName>
(Duda, 1918) is a junior synonym of
<taxonomicName lsidName="P. moesta" pageId="0" pageNumber="1" rank="species" species="moesta">
<emphasis italics="true" pageId="0" pageNumber="1">P. moesta</emphasis>
</taxonomicName>
(Villeneuve, 1918), see
<bibRefCitation DOI="https://doi.org/10.3406/bsef.2000.16706" author="Rohacek, J" journalOrPublisher="Higher Brachycera. Science Herald, Budapest" pageId="0" pageNumber="1" refId="B62" refString="Rohacek, J, 2001. The type material of Sphaeroceridae described by J. Villeneuve with lectotype designations and nomenclatural and taxonomic notes (Diptera). Bulletin de la Societe entomologique de France 105(5) (2000): 467-478. https://doi.org/10.3406/bsef.2000.16706" title="The type material of Sphaeroceridae described by J. Villeneuve with lectotype designations and nomenclatural and taxonomic notes (Diptera). Bulletin de la Societe entomologique de France 105 (5) (2000): 467 - 478." url="https://doi.org/10.3406/bsef.2000.16706" year="2001">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
(2001)
</bibRefCitation>
and
<bibRefCitation author="Rohacek, J" journalOrPublisher="Slezske zemske muzeum, Opava" pageId="0" pageNumber="1" refId="B70" refString="Rohacek, J, Marshall, SA, Norrbom, AL, Buck, M, Quiros, DI, Smith, I, 2001. World catalog of Sphaeroceridae (Diptera). Slezske zemske muzeum, Opava" title="World catalog of Sphaeroceridae (Diptera)." year="2001">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
et al. (2001)
</bibRefCitation>
.
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina turfosa</emphasis>
</taxonomicName>
shares all synapomorphic characters defining this group (cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 100), viz. the densely and long setose gonostylus, the distiphallus with spinose or toothed distal sclerites and a well-developed additional sclerite between female S8 and S10, except for his character 11 (middle interfrontal setae cruciate). Surprisingly,
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
appears to have the male terminalia most similar to those of the macropterous Nearctic species
<taxonomicName lsidName="P. vockerothi" pageId="0" pageNumber="1" rank="species" species="vockerothi">
<emphasis italics="true" pageId="0" pageNumber="1">P. vockerothi</emphasis>
</taxonomicName>
Marshall, 1986. The shared characters include (1) male S6 with a ventromedial process (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">23</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 79), (2) the gonostylus with long and slender anteroventral projection (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">24</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 77), and (3) similar shape of postgonite (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">30</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 78). The former two features (1, 2) could be considered synapomorphic and demonstrating a closer relationship of these species. Additionally, the female T8 and the spectacles-shaped sclerite seem to be similarly formed in
<taxonomicName lsidName="P. vockerothi" pageId="0" pageNumber="1" rank="species" species="vockerothi">
<emphasis italics="true" pageId="0" pageNumber="1">P. vockerothi</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">34</figureCitation>
,
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">39</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: figs 38, 40) but T8 in
<taxonomicName lsidName="P. vockerothi" pageId="0" pageNumber="1" rank="species" species="vockerothi">
<emphasis italics="true" pageId="0" pageNumber="1">P. vockerothi</emphasis>
</taxonomicName>
has a small medial strip-like sclerite in addition to large lateral sclerites and the female S8 and additional (acs) sclerite are markedly different in the shape and chaetotaxy (Fig.
<figureCitation captionStart="Figures 33–39" captionStartId="F13" captionText="Figures 33 - 39. Pullimosina (Pullimosina) turfosa sp. nov. (female paratype) 33 spermathecae 34 postabdomen, dorsally 35 ditto, ventrally 36 ditto, laterally 37 S 8 and additional sclerite, ventrally 38 spectacles-shaped sclerite, laterally 39 ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (34 - 36); 0.05 mm (33, 37 - 39). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum." figureDoi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1">37</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 39). There are also distinct differences in the armature of the male S5 (Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">23</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 79), shape of the gonostylus (having basal part very small and anteroventral projection simple in
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
: Fig.
<figureCitation captionStart="Figures 23–26" captionStartId="F10" captionText="Figures 23 - 26. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 23 S 5 and postabdominal sclerites, ventrally 24 gonostylus, laterally 25 external genitalia, caudally 26 ditto, laterally. Scale bars: 0.1 mm (23, 25, 26); 0.05 mm (24). Abbreviations: ce - cercus, dp - distiphallus, ep - epandrium, gs - gonostylus, hy - hypandrium, ma - medandrium, pg - postgonite, pha - phallapodeme, S - sternum." figureDoi="10.3897/zookeys.1132.94579.figures23-26" httpUri="https://binary.pensoft.net/fig/774335" pageId="0" pageNumber="1">24</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 77) and detailed structure of the distiphallus (Fig.
<figureCitation captionStart="Figures 27–30" captionStartId="F11" captionText="Figures 27 - 30. Pullimosina (Pullimosina) turfosa sp. nov. (male paratype) 27 aedeagus (phallus) dorsally 28 ditto, laterally 29 aedeagal complex, laterally 30 pregonite (enlarged), laterally. Scale bars 0.05 mm (27 - 29), 0.02 mm (30). Abbreviations: dp - distiphallus, ea - ejacapodeme, pg - postgonite, pha - phallapodeme, pp - phallophore, prg - pregonite." figureDoi="10.3897/zookeys.1132.94579.figures27-30" httpUri="https://binary.pensoft.net/fig/774336" pageId="0" pageNumber="1">28</figureCitation>
, cf.
<bibRefCitation DOI="https://doi.org/10.1139/z86-077" author="Marshall, SA" journalOrPublisher="Canadian Journal of Zoology" pageId="0" pageNumber="1" pagination="522 - 536" refId="B28" refString="Marshall, SA, 1986. A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae). Canadian Journal of Zoology 64 (2): 522 - 536, DOI: https://doi.org/10.1139/z86-077" title="A revision of the Nearctic species of the genus Pullimosina (Diptera, Sphaeroceridae)." url="https://doi.org/10.1139/z86-077" volume="64" year="1986">Marshall 1986</bibRefCitation>
: fig. 78).
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina turfosa</emphasis>
</taxonomicName>
can be most easily recognized from all Holarctic
<taxonomicName class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Pullimosina</emphasis>
</taxonomicName>
species by its strongly abbreviated wings with very characteristic venation (Figs
<figureCitation captionStart="Figures 14–17" captionStartId="F8" captionText="Figures 14 - 17. Pullimosina (Pullimosina) turfosa sp. nov. (holotype, paratypes) heads and wings 14 male head, frontally (holotype) 15 male right wing, dorsally (paratype) 16 female head, frontally (paratype) 17 female right wing, dorsally (paratype). Scale bars: 0.2 mm (14, 16); 0.1 mm (15, 17)." figureDoi="10.3897/zookeys.1132.94579.figures14-17" httpUri="https://binary.pensoft.net/fig/774333" pageId="0" pageNumber="1">15</figureCitation>
,
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">22</figureCitation>
). As for European species, the brachypterous form of the wing-polymorphic
<taxonomicName lsidName="P. meijerei" pageId="0" pageNumber="1" rank="species" species="meijerei">
<emphasis italics="true" pageId="0" pageNumber="1">P. meijerei</emphasis>
</taxonomicName>
(Duda, 1918) externally most resembles this new species (cf.
<bibRefCitation author="Rohacek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="1" pagination="535 - 558" publicationUrl="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" refId="B64" refString="Rohacek, J, 2012. Wing polymorphism in European species of Sphaeroceridae (Diptera). Acta Entomologica Musei Nationalis Pragae 52 (2): 535 - 558, https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" title="Wing polymorphism in European species of Sphaeroceridae (Diptera)." url="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" volume="52" year="2012">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
2012
</bibRefCitation>
: figs 3, 4) including coloration of the head but the wings of
<taxonomicName lsidName="P. meijerei" pageId="0" pageNumber="1" rank="species" species="meijerei">
<emphasis italics="true" pageId="0" pageNumber="1">P. meijerei</emphasis>
</taxonomicName>
are less shortened, more elongate and with more complete venation (
<bibRefCitation author="Rohacek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="1" pagination="535 - 558" publicationUrl="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" refId="B64" refString="Rohacek, J, 2012. Wing polymorphism in European species of Sphaeroceridae (Diptera). Acta Entomologica Musei Nationalis Pragae 52 (2): 535 - 558, https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" title="Wing polymorphism in European species of Sphaeroceridae (Diptera)." url="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" volume="52" year="2012">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
2012
</bibRefCitation>
: figs 24-27) not to mention very dissimilar structures of the male and female postabdomen (cf.
<bibRefCitation author="Rohacek, J" journalOrPublisher="Beitraege zur Entomologie, Berlin" pageId="0" pageNumber="1" pagination="101 - 179" publicationUrl="https://www.contributions-to-entomology.org/article/view/1209/1208" refId="B58" refString="Rohacek, J, 1985. A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part IV. Beitraege zur Entomologie, Berlin 35: 101 - 179, https://www.contributions-to-entomology.org/article/view/1209/1208" title="A monograph and re-classification of the previous genus Limosina Macquart (Diptera, Sphaeroceridae) of Europe. Part IV." url="https://www.contributions-to-entomology.org/article/view/1209/1208" volume="35" year="1985">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
1985
</bibRefCitation>
: figs 792-802).
</paragraph>
<caption doi="10.3897/zookeys.1132.94579.figures33-39" httpUri="https://binary.pensoft.net/fig/774338" pageId="0" pageNumber="1" start="Figures 33–39" startId="F13">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figures 33-39.</emphasis>
<taxonomicName authorityName="Roháček &amp; Przhiboro" authorityYear="2022" class="Insecta" family="Sphaeroceridae" genus="Pullimosina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pullimosina (Pullimosina) turfosa" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="turfosa" subGenus="Pullimosina">Pullimosina (Pullimosina) turfosa</taxonomicName>
sp. nov. (female paratype)
<emphasis bold="true" pageId="0" pageNumber="1">33</emphasis>
spermathecae
<emphasis bold="true" pageId="0" pageNumber="1">34</emphasis>
postabdomen, dorsally
<emphasis bold="true" pageId="0" pageNumber="1">35</emphasis>
ditto, ventrally
<emphasis bold="true" pageId="0" pageNumber="1">36</emphasis>
ditto, laterally
<emphasis bold="true" pageId="0" pageNumber="1">37</emphasis>
S8 and additional sclerite, ventrally
<emphasis bold="true" pageId="0" pageNumber="1">38</emphasis>
spectacles-shaped sclerite, laterally
<emphasis bold="true" pageId="0" pageNumber="1">39</emphasis>
ditto and single spermatheca, ventrally. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">34-36</emphasis>
); 0.05 mm (
<emphasis bold="true" pageId="0" pageNumber="1">33, 37-39</emphasis>
). Abbreviations: asc - additional sclerite, ce - cercus, S - sternum, T - tergum.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
The peculiar reduction of the wing and its veins in
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
(Fig.
<figureCitation captionStart="Figures 18–22" captionStartId="F9" captionText="Figures 18 - 22. Pullimosina (Pullimosina) turfosa sp. nov. (paratypes) 18 male fore leg without tarsus, posteriorly 19 male mid tibia, dorsally 20 male mid femur, tibia and basitarsus, anteriorly 21 female mid tibia and basitarsus, anteriorly 22 male right wing, dorsally. Scale bars: 0.2 mm (18 - 21); 0.1 mm (22)." figureDoi="10.3897/zookeys.1132.94579.figures18-22" httpUri="https://binary.pensoft.net/fig/774334" pageId="0" pageNumber="1">22</figureCitation>
) needs a special comment. It differs from all other cases of brachyptery known in West Palaearctic
<taxonomicName authorityName="Macquart" authorityYear="1835" class="Insecta" family="Sphaeroceridae" kingdom="Animalia" lsidName="" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="family">Sphaeroceridae</taxonomicName>
(
<bibRefCitation author="Rohacek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="1" pagination="535 - 558" publicationUrl="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" refId="B64" refString="Rohacek, J, 2012. Wing polymorphism in European species of Sphaeroceridae (Diptera). Acta Entomologica Musei Nationalis Pragae 52 (2): 535 - 558, https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" title="Wing polymorphism in European species of Sphaeroceridae (Diptera)." url="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" volume="52" year="2012">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
2012
</bibRefCitation>
) in having the distal part of wing strongly abbreviated while its basal part (up to subcostal break) is almost normal, R4+5 is completely absent (in this somewhat resembling the wing venation of
<taxonomicName baseAuthorityName="Papp &amp; Rohacek" baseAuthorityYear="1981" class="Insecta" family="Sphaeroceridae" genus="Aptilotus" kingdom="Animalia" lsidName="Aptilotus anapterus" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="anapterus">
<emphasis italics="true" pageId="0" pageNumber="1">Aptilotus anapterus</emphasis>
</taxonomicName>
(Papp &amp;
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
, 1981) from La Palma, Canary Is, which, however, has a small basal remnant of this vein retained) but simultaneously with M present. Thus, the reduction of veins in
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
is somewhat intermediate between stages 5 and 6 as recognized by
<bibRefCitation author="Rohacek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="1" pagination="535 - 558" publicationUrl="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" refId="B64" refString="Rohacek, J, 2012. Wing polymorphism in European species of Sphaeroceridae (Diptera). Acta Entomologica Musei Nationalis Pragae 52 (2): 535 - 558, https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" title="Wing polymorphism in European species of Sphaeroceridae (Diptera)." url="https://www.aemnp.eu/data/article-1389/1370-52_2_535.pdf" volume="52" year="2012">
<normalizedToken originalValue="Roháček">Rohacek</normalizedToken>
(2012
</bibRefCitation>
: figs 39, 40).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">The species is known only from its type locality in Russia, North Ossetia (Caucasus Mts).</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
<paragraph pageId="0" pageNumber="1">
All specimens of the new species were collected on 17 and 18 August 2018 in a high-montane Chifandzar mire (Fig.
<figureCitation captionStart="Figures 6, 7" captionStartId="F4" captionText="Figures 6, 7. Mires under study 6 Ushtulu, 21 September 2018 7 Chifandzar, 2 June 2018 (arrow indicates an area in which large Sphagnum hummocks are located)." figureDoi="10.3897/zookeys.1132.94579.figures6-7" httpUri="https://binary.pensoft.net/fig/774329" pageId="0" pageNumber="1">7</figureCitation>
), which is the highest (2289 m) and the largest (ca. 0.5 km2) of the mires under study. This mire is much more open and windier compared to the others.
</paragraph>
<paragraph pageId="0" pageNumber="1">
All type specimens but one were collected from large
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum</emphasis>
</taxonomicName>
hummocks (Fig.
<figureCitation captionStart="Figures 8, 9" captionStartId="F5" captionText="Figures 8, 9. Mires under study 8 Chifandzar, 18 September 2018, a large Sphagnum hummock 9 Tarskoe, 11 September 2018." figureDoi="10.3897/zookeys.1132.94579.figures8-9" httpUri="https://binary.pensoft.net/fig/774330" pageId="0" pageNumber="1">8</figureCitation>
). This habitat is distinctive and represented only by nearly 15 hummocks all of which are located in the eastern part of the mire (Fig.
<figureCitation captionStart="Figures 6, 7" captionStartId="F4" captionText="Figures 6, 7. Mires under study 6 Ushtulu, 21 September 2018 7 Chifandzar, 2 June 2018 (arrow indicates an area in which large Sphagnum hummocks are located)." figureDoi="10.3897/zookeys.1132.94579.figures6-7" httpUri="https://binary.pensoft.net/fig/774329" pageId="0" pageNumber="1">7</figureCitation>
: arrow). The hummocks are scattered over an area of ca. 100
<normalizedToken originalValue="×">x</normalizedToken>
50 m. Each hummock is 0.15-0.3 m high and 0.5-2 m wide. It consists mostly of loose thick cushion of
<taxonomicName authorityName="C.Linnaeus" authorityYear="1753" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum</emphasis>
</taxonomicName>
(
<taxonomicName lsidName="S. teres" pageId="0" pageNumber="1" rank="species" species="teres">
<emphasis italics="true" pageId="0" pageNumber="1">S. teres</emphasis>
</taxonomicName>
is predominant;
<taxonomicName lsidName="S. centrale" pageId="0" pageNumber="1" rank="species" species="centrale">
<emphasis italics="true" pageId="0" pageNumber="1">S. centrale</emphasis>
</taxonomicName>
and
<taxonomicName lsidName="S. squarrosum" pageId="0" pageNumber="1" rank="species" species="squarrosum">
<emphasis italics="true" pageId="0" pageNumber="1">S. squarrosum</emphasis>
</taxonomicName>
are common), with sparse shoots of
<taxonomicName class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" lsidName="Carex" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Carex</emphasis>
</taxonomicName>
spp. and
<taxonomicName class="Liliopsida" family="Poaceae" genus="Nardus" kingdom="Plantae" lsidName="Nardus stricta" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis italics="true" pageId="0" pageNumber="1">Nardus stricta</emphasis>
</taxonomicName>
and abundant remains of monocotyledons. The substrate of hummocks is dry to slightly wet, as distinct from moist or water-logged substrate on flat areas surrounding the hummocks and in other parts of the mire.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Most specimens were collected by means of sifting substrata of hummocks. Two females were collected by washing and subsequent flotation of substrate in NaCl solution: one specimen was sampled from the same habitat, and another one, from moist substrate beyond the hummocks, with predominating
<taxonomicName authorityName="Nees in Sturm" authorityYear="1819" class="Insecta" family="Sphagnaceae" genus="Sphagnum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sphagnum subsecundum" order="Diptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="subsecundum">
<emphasis italics="true" pageId="0" pageNumber="1">Sphagnum subsecundum</emphasis>
</taxonomicName>
and
<taxonomicName class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" lsidName="Carex rostrata" order="Poales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="rostrata">
<emphasis italics="true" pageId="0" pageNumber="1">Carex rostrata</emphasis>
</taxonomicName>
. Hence, most individuals of
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
concentrate in hummocks but some flies may also occur at some distance from them. No specimens were collected in early summer (2-3 June), suggesting that the adults of
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
appear later.
</paragraph>
<paragraph pageId="0" pageNumber="1">
Due to exclusive association of
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
with the sphagnetum habitat, particularly with hummocks, we consider it a tyrphobiont (= eucoenic to peat-bog habitat) sphagnicolous species. Interestingly, no specimens of
<taxonomicName lsidName="P. turfosa" pageId="0" pageNumber="1" rank="species" species="turfosa">
<emphasis italics="true" pageId="0" pageNumber="1">P. turfosa</emphasis>
</taxonomicName>
were collected from similar substrata in other bogs using the same techniques (sifting and washing/flotation). It is possible that the new peculiar species is confined to high montane bogs or even endemic to Chifandzar, considering that the montane bogs of the North Caucasus are rare and isolated island ecosystems.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink