130 lines
13 KiB
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130 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.803.28903" ID-GBIF-Dataset="6c6c0b59-636f-4502-8b48-642ba231eb16" ID-PMC="PMC6390078" ID-Pensoft-Pub="1313-2970-803-7" ID-PubMed="30814899" ID-ZBK="936CFD88F297440EA9BE4C258AE9BD09" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-803-7" ModsDocOrigin="ZooKeys 803" ModsDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1545315537716" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo & Balke, Michael" docDate="2018" docId="3967073F196D007E333101A799C3431E" docLanguage="en" docName="ZooKeys 803: 7-70" docOrigin="ZooKeys 803" docSource="http://dx.doi.org/10.3897/zookeys.803.28903" docTitle="Exocelina keki Shaverdo & Balke, sp. n." docType="treatment" docUuid="0C2652F3-452D-4826-918A-90D6C5C5ECA8" docUuidSource="ZooBank" docVersion="7" lastPageNumber="24" masterDocId="5133FFF6125A1065FF83FF96FFF3FFD4" masterDocTitle="Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="70" masterPageNumber="7" pageNumber="22" updateTime="1668166558239" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Introduction of the Exocelinacasuarina-group, with a key to its representatives and descriptions of 19 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2018</mods:date>
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<mods:detail type="volume">
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<mods:number>803</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>7</mods:start>
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<mods:end>70</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.803.28903</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.803.28903</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-803-7</mods:identifier>
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<mods:identifier type="ZBK">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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<mods:identifier type="ZooBank">936CFD88F297440EA9BE4C258AE9BD09</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="154125607" LSID="urn:lsid:zoobank.org:act:0C2652F3-452D-4826-918A-90D6C5C5ECA8" httpUri="http://treatment.plazi.org/id/3967073F196D007E333101A799C3431E" lastPageId="17" lastPageNumber="24" pageId="15" pageNumber="22">
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<subSubSection pageId="15" pageNumber="22" type="nomenclature">
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<paragraph pageId="15" pageNumber="22">
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10.
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<taxonomicName LSID="http://zoobank.org/0C2652F3-452D-4826-918A-90D6C5C5ECA8" authority="Shaverdo & Balke" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina keki" order="Coleoptera" pageId="15" pageNumber="22" phylum="Arthropoda" rank="species" species="keki">Exocelina keki Shaverdo & Balke</taxonomicName>
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<taxonomicNameLabel pageId="15" pageNumber="22">sp. n.</taxonomicNameLabel>
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Figs 6, 30
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="23" type="reference_group">
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<paragraph pageId="16" pageNumber="23">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina" order="Coleoptera" pageId="16" pageNumber="23" phylum="Arthropoda" rank="genus">
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<pageBreakToken pageId="16" pageNumber="23" start="start">Exocelina</pageBreakToken>
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</taxonomicName>
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undescribed sp. MB1530:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/ncomms5001" year="2014">Toussaint et al. 2014</bibRefCitation>
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: supplementary figs 1-4, table 2;
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary figs S1-S2, table S3.
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</paragraph>
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<paragraph pageId="16" pageNumber="23">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudokeki" order="Coleoptera" pageId="16" pageNumber="23" phylum="Arthropoda" rank="species" species="pseudokeki">Exocelina pseudokeki</taxonomicName>
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_New_Guinea_MB1530:
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<bibRefCitation author="Toussaint, EFA" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" url="https://doi.org/10.1038/srep16016" year="2015">Toussaint et al. 2015</bibRefCitation>
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: supplementary information S5-S6.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="23" type="type locality">
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<paragraph pageId="16" pageNumber="23">Type locality.</paragraph>
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<paragraph pageId="16" pageNumber="23">
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Papua New Guinea: Madang, Adelbert Mts, creek near Keki,
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<geoCoordinate direction="south" orientation="latitude" precision="9" value="-4.705">04°42.30'S</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="9" value="145.41817">145°25.09'E</geoCoordinate>
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, 790 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="23" type="type material">
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<paragraph pageId="16" pageNumber="23">Type material.</paragraph>
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<paragraph pageId="16" pageNumber="23">Holotype: male "Papua New Guinea: Madang, Keki, Adalbert [sic!] Mts., 500m, 29.xi.2006, nr 04.43.058S 145.24.437E, Balke & Kinibel (PNG 118)" (ZSM). Paratypes: Madang: 9 males, 4 females with the same labels as the holotype, one male with an additional green label "M.Balke 1530" (NHMW, ZSM). 1 male, 3 females "Papua New Guinea: Madang, Adalbert [sic!] Mts., creek nr Keki, 790m, 28.xi.1994, 04.42.300S 145.25.089E, Binatang Boys leg. (PNG 53a)" (ZSM). 1 male, 2 females "Papua New Guinea: Madang, Keki-Sewan, Adalbert [sic!] Mts., 700m, 30.xi.2006, nr 04.41.802S 145.25.460E, Binatang Boys (PNG 120)" (ZSM). Eastern Highlands: 1 male "Papua New Guinea: Eastern Highlands, Bena Bridge, 1400m, 8.xii.2007, 06.10.781S 145.26.034E, Balke & Sagata (PNG 164)" (ZSM).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="17" lastPageNumber="24" pageId="16" pageNumber="23" type="description">
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<paragraph pageId="16" pageNumber="23">Description.</paragraph>
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<paragraph pageId="16" pageNumber="23">Body size and form: Beetle small: TL-H 3.15-3.65 mm, TL 3.45-4.0 mm, MW 1.65-1.9 mm (holotype: TL-H 3.65 mm, TL 4 mm, MW 1.85 mm), with oblong habitus.</paragraph>
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<paragraph pageId="16" pageNumber="23">Coloration: Reddish to reddish brown, with head and pronotum paler. Head yellowish red to reddish brown, with small darker areas posterior to eyes. Pronotum yellowish red to reddish brown, with small brown to dark brown area on disc. Elytra reddish brown to brown, with narrow reddish sutural lines. Head appendages yellowish red, legs reddish, distally darker, especially metathoracic legs (Fig. 6). Teneral specimens paler.</paragraph>
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<paragraph lastPageId="17" lastPageNumber="24" pageId="16" pageNumber="23">
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Surface sculpture: Submatt dorsally. Head with rather dense punctation (spaces between punctures 1-2 times size of punctures), evidently finer and sparser anteriorly;
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<pageBreakToken pageId="17" pageNumber="24" start="start">diameter</pageBreakToken>
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of punctures smaller than diameter of cells of microreticulation. Pronotum and elytra with dense, distinct but fine punctation, sparser and finer than on head. Pronotum and elytra with strongly impressed microreticulation. Head with microreticulation stronger. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine sparse punctation.
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</paragraph>
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<paragraph pageId="17" pageNumber="24">Structures: Pronotum without lateral bead, in few specimens with its traces in posterior part. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively narrow, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 rounded.</paragraph>
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<paragraph pageId="17" pageNumber="24">
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Male: Antennae simple (Fig. 6). Protarsomere 4 with anterolateral seta thin, long, smaller than more laterally situated large setae, slightly curved downwards. Protarsomere 5 long and narrow, with anterior band of more than 40 and posterior row of 7 relatively long, thin setae (Fig. 30D). Median lobe in lateral view almost straight, its apex rounded and not curved downwards; in ventral view, distally distinctly narrowed before rounded, narrow apex. Paramere slightly concave on dorsal side and with dorsal setae distinctly divided to long, dense subdistal setae and sparser proximal ones (Fig. 30
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<normalizedToken originalValue="A–C">A-C</normalizedToken>
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). Abdominal ventrite 6 with 3-5 lateral striae on each side.
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</paragraph>
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<paragraph pageId="17" pageNumber="24">Female: Without evident differences in external morphology from males, except for not modified protarsi and abdominal ventrite 6 without striae.</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="24" type="distribution">
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<paragraph pageId="17" pageNumber="24">Distribution and variability.</paragraph>
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<paragraph pageId="17" pageNumber="24">
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Papua New Guinea: Madang and Eastern Highlands (Fig. 50). The species is known mainly from Keki area in Adelbert Mountains; only one beetle was collected in Bena, Eastern Highlands. It shows no morphological difference from the specimens of Keki populations, except for a small difference in the median lobe shape, which could be an expression of species variability. The species might have the same pattern of distribution as
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina brahminensis" order="Coleoptera" pageId="17" pageNumber="24" phylum="Arthropoda" rank="species" species="brahminensis">Exocelina brahminensis</taxonomicName>
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<bibRefCitation author="Shaverdo, HV" journalOrPublisher="ZooKeys" pageId="40" pageNumber="47" pagination="1 - 76" title="Introduction of the Exocelinaekari-group with descriptions of 22 new species from New Guinea (Coleoptera, Dytiscidae, Copelatinae)." url="https://doi.org/10.3897/zookeys.250.3715" volume="250" year="2012">Shaverdo et al., 2012</bibRefCitation>
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, which has a wide distribution in the Momase Region and is known from Adelbert Mountains and Bena.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="24" type="affinities">
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<paragraph pageId="17" pageNumber="24">Affinities.</paragraph>
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<paragraph pageId="17" pageNumber="24">
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina keki" order="Coleoptera" pageId="17" pageNumber="24" phylum="Arthropoda" rank="species" species="keki">Exocelina keki</taxonomicName>
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sp. n. is very similar to
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<taxonomicName class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina messeri" order="Coleoptera" pageId="17" pageNumber="24" phylum="Arthropoda" rank="species" species="messeri">Exocelina messeri</taxonomicName>
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(Balke, 1999) in body form and coloration, but has much more distinct dorsal punctation and stronger microreticulation, as well as median lobe more slender, with apex smaller and narrower in ventral view; the ventral setae of male protarsomere 5 are much less numerous and clearly divided into anterior band and posterior row.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="17" pageNumber="24" type="etymology">
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<paragraph pageId="17" pageNumber="24">Etymology. The</paragraph>
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<paragraph pageId="17" pageNumber="24">species is named after Keki Village. The name is a noun in the nominative singular standing in apposition.</paragraph>
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</subSubSection>
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</treatment>
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</document> |