1188 lines
143 KiB
XML
1188 lines
143 KiB
XML
<document id="0EF4069390F2134E6E6A1F98E2FDD29B" ID-DOI="10.11646/zootaxa.4911.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="4448898" ID-ZooBank="361E198B-5F04-4E5F-824A-73F488AB4A8B" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="diego" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1611043012182" checkinUser="plazi" docAuthor="David, Patrick, Vogel, Gernot, Nguyen, Truong Quang, Orlov, Nikolai L., Pauwels, Olivier S. G., Teynié, Alexandre & Ziegler, Thomas" docDate="2021" docId="383AEC41FF93FFC8FF6AFDD5FDB4CE5E" docLanguage="en" docName="zootaxa.4911.1.1.pdf" docOrigin="Zootaxa 4911 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Hebius annamensis" docType="treatment" docVersion="2" lastPageNumber="35" masterDocId="C4039439FF8FFFEAFFFDFFF6FFF3C80F" masterDocTitle="A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand" masterLastPageNumber="61" masterPageNumber="1" pageNumber="29" updateTime="1730213606799" updateUser="diego">
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<mods:title id="A7C6C8D7A24F42FF22E09F1428347A57">A revision of the dark-bellied, stream-dwelling snakes of the genus Hebius (Reptilia: Squamata: Natricidae) with the description of a new species from China, Vietnam and Thailand</mods:title>
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<mods:name id="DA5EB7751E1D2FA0C65FC025CC209BE9" type="personal">
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<mods:namePart id="937D8B358B1E583CD9649199042615F4">David, Patrick</mods:namePart>
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<mods:affiliation id="B022A678A89BBBC92C655291E7C1F531">Reptiles & Amphibiens, UMR 7205 OSEB, Département Évolution et Systématique, CP 30, Muséum National d’Histoire Naturelle. 57 rue Cuvier, F- 75231 Paris Cedex 05, France. patrick. david @ mnhn. fr; https: // orcid. org / 0000 - 0001 - 7562 - 684 X</mods:affiliation>
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<mods:nameIdentifier id="6890995CA8A9B52A7D45272EB37D962C" type="email">patrick.david@mnhn.fr</mods:nameIdentifier>
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<mods:roleTerm id="E87BF5A92211579530AD15CFCC975EDD">Author</mods:roleTerm>
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<mods:namePart id="AD22232F79EA4BA0CD780DA99B3400B4">Vogel, Gernot</mods:namePart>
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<mods:affiliation id="01C5D3A3B22A2446E09F3E027DBBB98C">Society for Southeast Asian Herpetology, Im Sand 3, D- 69115 Heidelberg, Germany. Gernot. Vogel @ t-online. de; https: // orcid. org / 0000 - 0002 - 4542 - 518 X</mods:affiliation>
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<mods:nameIdentifier id="8E56C37EDBF6E5407759E971105CC2A7" type="email">Vogel@t-online.de</mods:nameIdentifier>
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<mods:namePart id="2A0DB7617AE3997AB51FF406A47C60A9">Nguyen, Truong Quang</mods:namePart>
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<mods:affiliation id="6CABCB3692FA0691ADF6772AA57ED86C">Institute of Ecology and Biological Resources and Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet Road, Hanoi, Vietnam. nqt 2 @ yahoo. com; https: // orcid. org / 0000 - 0002 - 6601 - 0880</mods:affiliation>
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<mods:nameIdentifier id="4811D4A6ECCF613067D2F7D95E290697" type="email">nqt2@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="E4C1F3C0DECF646AA28D440CDD720948">Orlov, Nikolai L.</mods:namePart>
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<mods:affiliation id="09DC9C8855EFED8A3CE1E07F88906574">Laboratory of Herpetology, Zoological Institute, Academy of Sciences, 199034 St. Petersburg, Russia. azemiops @ zin. ru; https: // orcid. org / 0000 - 0003 - 4401 - 348 X</mods:affiliation>
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<mods:nameIdentifier id="1516BC3D7BA4C3ED4EF484F4863CD7C4" type="email">azemiops@zin.ru</mods:nameIdentifier>
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<mods:namePart id="A34729730B6060E37474ADE2887A6B9D">Pauwels, Olivier S. G.</mods:namePart>
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<mods:affiliation id="73550DA9DC3D963F2BC12F74118B1C52">Département des Vertébrés Récents, Institut Royal des Sciences Naturelles de Belgique, Rue Vautier 29, B- 1000 Brussels, Belgium. osgpauwels @ yahoo. fr; https: // orcid. org / 0000 - 0002 - 3265 - 5496</mods:affiliation>
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<mods:namePart id="1D1140CEA36FD04193F35CCFA94D3001">Teynié, Alexandre</mods:namePart>
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<mods:affiliation id="4BFF815AE95B96E0A1BF3424CA19BEB6">Société d’Histoire Naturelle Alcide d’Orbigny, 57 rue de Gergovie, F- 63170 Aubière, France. alexandre. teynie @ inrae. fr; https: // orcid. org / 0000 - 0002 - 6588 - 229 X</mods:affiliation>
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<mods:nameIdentifier id="25C3592793639F47302538B13F081D41" type="email">alexandre.teynie@inrae.fr</mods:nameIdentifier>
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<mods:roleTerm id="E3D2EBD53F764B46B0ED7DF2A4A65E16">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B568864C87E978819C58C8DD714E72A4">Ziegler, Thomas</mods:namePart>
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<mods:affiliation id="FAA6947AEA3872E3B1FB4CD1736D0DAA">Reptiles & Amphibiens, UMR 7205 OSEB, Département Évolution et Systématique, CP 30, Muséum National d’Histoire Naturelle. 57 rue Cuvier, F- 75231 Paris Cedex 05, France. patrick. david @ mnhn. fr; https: // orcid. org / 0000 - 0001 - 7562 - 684 X & AG Zoologischer Garten K ̂ ln, Riehler Str. 173, D- 50735 K ̂ ln, Germany. ziegler @ koelnerzoo. de; https: // orcid. org / 0000 - 0002 - 4797 - 609 X Corresponding author & Reptiles & Amphibiens, UMR 7205 OSEB, Département Évolution et Systématique, CP 30, Muséum National d’Histoire Naturelle. 57 rue Cuvier, F- 75231 Paris Cedex 05, France. patrick. david @ mnhn. fr; https: // orcid. org / 0000 - 0001 - 7562 - 684 X</mods:affiliation>
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<mods:nameIdentifier id="A245860C05A5ADFCE61105456BEB3442" type="email">patrick.david@mnhn.fr</mods:nameIdentifier>
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<mods:typeOfResource id="BFCC1FA2CBD9C462E3D95267CD8B086E">text</mods:typeOfResource>
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<mods:title id="917A6D933070AB9F03BB5AC04ACDE610">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="1F17EDFFD68E8F4486153018A8230152">
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<mods:date id="0AFB84DA239A3ED9A4F4FBCDC3FFD686">2021</mods:date>
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<mods:number id="5A8F6FA38A36C645C3FC3EF6B72DA840">2021-01-19</mods:number>
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</mods:detail>
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<mods:detail id="201F68774D36196B201F9352591B43DA" type="volume">
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<mods:number id="F0F3A5791AAFB1951B64ECF8324B5E79">4911</mods:number>
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</mods:detail>
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<mods:detail id="884AFC72CB82B9E9870278F1169E220E" type="issue">
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<mods:number id="5D02F8BD05408089E61D03C2E1D2A0E1">1</mods:number>
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<mods:start id="251BD2D8B5F23E74504C8D7483C1E0CE">1</mods:start>
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<mods:end id="020BA53E2D9959B200CAC854FE454FC2">61</mods:end>
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<mods:classification id="F02B180A4F662801FF0975444CA38908">journal article</mods:classification>
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<mods:identifier id="B2999BEB3E603735FB3C9F7222D525FB" type="DOI">10.11646/zootaxa.4911.1.1</mods:identifier>
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<mods:identifier id="5376E3E5BB809445541F9B45EBB6E107" type="ISSN">1175-5326</mods:identifier>
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<treatment id="383AEC41FF93FFC8FF6AFDD5FDB4CE5E" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E" httpUri="http://treatment.plazi.org/id/383AEC41FF93FFC8FF6AFDD5FDB4CE5E" lastPageId="34" lastPageNumber="35" pageId="28" pageNumber="29">
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<subSubSection id="F8890EDCFF93FFF6FF6AFDD5FD99CA33" box="[151,618,546,573]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="B02C5D57FF93FFF6FF6AFDD5FD99CA33" blockId="28.[151,618,546,609]" box="[151,618,546,573]" pageId="28" pageNumber="29">
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<emphasis id="82E78145FF93FFF6FF6AFDD5FD99CA33" bold="true" box="[151,618,546,573]" pageId="28" pageNumber="29">
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5.
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<taxonomicName id="779326D4FF93FFF6FF48FDD4FD99CA33" ID-CoL="8H5HW" authority="(Bourret, 1934)" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[181,618,546,573]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
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<emphasis id="82E78145FF93FFF6FF48FDD4FE50CA32" bold="true" box="[181,419,546,573]" italics="true" pageId="28" pageNumber="29">Hebius annamensis</emphasis>
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(Bourret, 1934)
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</taxonomicName>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="F8890EDCFF93FFF6FF6AFDB1FEDECA6E" box="[151,301,583,609]" pageId="28" pageNumber="29" type="description">
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<paragraph id="B02C5D57FF93FFF6FF6AFDB1FEDECA6E" blockId="28.[151,618,546,609]" box="[151,301,583,609]" pageId="28" pageNumber="29">
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(
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<figureCitation id="28A841D2FF93FFF6FF62FDB1FED7CA6E" box="[159,292,583,609]" captionStart-0="FIGURE 10" captionStart-1="FIGURE 11" captionStartId-0="30.[151,260,1942,1968]" captionStartId-1="31.[151,260,1941,1967]" captionTargetBox-0="[151,1436,179,1917]" captionTargetBox-1="[330,1257,180,1917]" captionTargetId-0="figure-28@30.[151,1436,179,1917]" captionTargetId-1="figure-19@31.[330,1257,179,1917]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 10. Hebius annamensis Bourret, holotype (MNHN 1938.0117)—A: dorsal view; B: lateral view of the head, right side. Photographs: Patrick David." captionText-1="FIGURE 11. Hebius annamensis Bourret, alive—A: specimen from Quang Tri Province, Vietnam. Photograph: Nikolai Orlov. B: specimen from Quang Binh Province. Photograph: Nikolai Orlov. C: specimen from Kon Tum Province. Photograph: Truong Quang Nguyen." figureDoi-0="http://doi.org/10.5281/zenodo.4463766" figureDoi-1="http://doi.org/10.5281/zenodo.4463768" httpUri-0="https://zenodo.org/record/4463766/files/figure.png" httpUri-1="https://zenodo.org/record/4463768/files/figure.png" pageId="28" pageNumber="29">Figs. 10–11</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="F8890EDCFF93FFF6FF6AFD7DFB82CAAC" box="[151,1137,651,675]" pageId="28" pageNumber="29" type="reference_group">
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<paragraph id="B02C5D57FF93FFF6FF6AFD7DFB82CAAC" blockId="28.[151,1437,651,1105]" box="[151,1137,651,675]" pageId="28" pageNumber="29">
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<taxonomicName id="779326D4FF93FFF6FF6AFD7DFD80CAAC" authority="Bourret, 1934 c: 167" authorityName="Bourret" authorityPageNumber="167" authorityYear="1934" box="[151,627,651,675]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
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<emphasis id="82E78145FF93FFF6FF6AFD7DFE6DCAAC" box="[151,414,651,675]" italics="true" pageId="28" pageNumber="29">Parahelicops annamensis</emphasis>
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<bibRefCitation id="D40220A6FF93FFF6FE58FD7AFD80CAAC" author="Bourret, R." box="[421,627,651,675]" pageId="28" pageNumber="29" pagination="167 - 176" refId="ref45586" refString="Bourret, R. (1934 c) Notes herpetologiques sur l'Indochine francaise. III. Ophidiens d'Annam et du Moyen Laos. Bulletin general de l'Instruction Publique, 13 e Annee (1933 - 1934), 9 Mai 1934, 167 - 176. [Separate reprint: 3 - 12]" type="book chapter" year="1934">Bourret, 1934c: 167</bibRefCitation>
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</taxonomicName>
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(Separate: p. 3) & 170 (Separate: p. 6),
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<figureCitation id="28A841D2FF93FFF6FBEBFD7DFBA1CAAC" box="[1046,1106,651,675]" captionStart="FIGURE 2" captionStartId="8.[151,260,1942,1968]" captionTargetBox="[383,1205,179,1918]" captionTargetId="figure-28@8.[383,1205,179,1918]" captionTargetPageId="8" captionText="FIGURE 2. Hebius venningi Wall, alive—A & C: specimen from Changlang District, Arunachal Pradesh, India. Photographs: Ashok Captain.—B: specimen CAS 233206, Chun Kyone, Hakha Township, Hakha District, Chin State, Myanmar. Photograph: Hla Tun." figureDoi="http://doi.org/10.5281/zenodo.4463744" httpUri="https://zenodo.org/record/4463744/files/figure.png" pageId="28" pageNumber="29">Fig. 2</figureCitation>
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.—
|
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</paragraph>
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</subSubSection>
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<subSubSection id="F8890EDCFF93FFF6FB8FFD7DFDA8CAF0" pageId="28" pageNumber="29" type="materials_examined">
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<paragraph id="B02C5D57FF93FFF6FB8FFD7DFDA8CAF0" blockId="28.[151,1437,651,1105]" pageId="28" pageNumber="29">
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<materialsCitation id="00FB570AFF93FFF6FB8FFD7DFDA8CAF0" collectionCode="MNHN" collectorName="Rene Bourret" country="Vietnam" elevation="0" location="Ba Na Nature Reserve" municipality="Da Nang City" pageId="28" pageNumber="29" specimenCode="MNHN 1938.0117" specimenCount="2" specimenCount-adult="1" specimenCount-male="1" typeStatus="holotype">
|
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<emphasis id="82E78145FF93FFF6FB8FFD7DFB0DCAAC" bold="true" box="[1138,1278,651,675]" pageId="28" pageNumber="29">Type locality</emphasis>
|
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. “Bana, station d’altitude située à 1 500 m d’altitude au bord de la Baie de Tourane”, now
|
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<location id="B54C0B8CFF93FFF6FC47FD5CFB6CCACE" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FC47FD5CFB6CCACE" box="[954,1183,682,705]" country="Vietnam" municipality="Da Nang City" name="Ba Na Nature Reserve" pageId="28" pageNumber="29">Ba Na Nature Reserve</location>
|
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,
|
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<collectingMunicipality id="5048C72DFF93FFF6FB54FD5CFAC4CACD" box="[1193,1335,682,706]" pageId="28" pageNumber="29">Da Nang City</collectingMunicipality>
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,
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<collectingCountry id="C8841DC7FF93FFF6FABDFD5CFA6BCACD" box="[1344,1432,682,706]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
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,
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<quantity id="776BF0B2FF93FFF6FF3AFD3EFEE9CAEF" box="[199,282,712,736]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="m" value="1500.0">1,500 m</quantity>
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a.s.l.—
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<emphasis id="82E78145FF93FFF6FE9AFD3FFE39CAEE" bold="true" box="[359,458,713,737]" pageId="28" pageNumber="29">
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<typeStatus id="6F28E3F5FF93FFF6FE9AFD3FFE39CAEE" box="[359,458,713,737]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
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</emphasis>
|
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.
|
||
<specimenCode id="E035F52CFF93FFF6FE2BFD3FFD65CAEE" box="[470,662,713,737]" pageId="28" pageNumber="29">MNHN 1938.0117</specimenCode>
|
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,
|
||
<specimenCount id="A69596DEFF93FFF6FD62FD3EFD21CAEF" box="[671,722,712,736]" pageId="28" pageNumber="29" type="adult">adult</specimenCount>
|
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<specimenCount id="A69596DEFF93FFF6FD25FD3EFCFACAEF" box="[728,777,712,736]" pageId="28" pageNumber="29" type="male">male</specimenCount>
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||
(Muséum National d’Histoire Naturelle, Paris, France); collected and deposited by
|
||
<collectorName id="1D663881FF93FFF6FE84FD11FDF2CAF0" box="[377,513,743,767]" pageId="28" pageNumber="29">René Bourret</collectorName>
|
||
in 1934.
|
||
</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F8890EDCFF93FFF6FF6AFCF0FCD3CC5E" pageId="28" pageNumber="29" type="reference_group">
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<paragraph id="B02C5D57FF93FFF6FF6AFCF0FE7ACBFB" blockId="28.[151,1437,651,1105]" pageId="28" pageNumber="29">
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<taxonomicName id="779326D4FF93FFF6FF6AFCF0FE6CCB12" authorityName="Bourret" authorityYear="1934" box="[151,415,774,798]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF93FFF6FF6AFCF0FE6CCB12" box="[151,415,774,798]" italics="true" pageId="28" pageNumber="29">Parahelicops annamensis</emphasis>
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</taxonomicName>
|
||
.—
|
||
<bibRefCitation id="D40220A6FF93FFF6FE41FCF0FD8ECB11" author="Bourret, R." box="[444,637,774,798]" pageId="28" pageNumber="29" refId="ref45746" refString="Bourret, R. (1936 a) Les Serpents de l'Indochine. Tome I. Etudes sur la faune. Henri Basuyau et Cie, Toulouse, 141 pp." type="book" year="1936">Bourret 1936a: 97</bibRefCitation>
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, 104, 114 & 125;
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<bibRefCitation id="D40220A6FF93FFF6FCC3FCF0FBF8CB11" author="Bourret, R." box="[830,1035,774,798]" pageId="28" pageNumber="29" refId="ref45777" refString="Bourret, R. (1936 b) Les Serpents de l'Indochine. Tome II. Catalogue systematique descriptif. Henri Basuyau et Cie, Toulouse, 505 pp." type="book" year="1936">Bourret 1936b: 122</bibRefCitation>
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, Fig. 50;
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<bibRefCitation id="D40220A6FF93FFF6FB8CFCF0FAC1CB11" author="Bourret, R." box="[1137,1330,774,798]" pageId="28" pageNumber="29" pagination="49 - 60" refId="ref45906" refString="Bourret, R. (1939 b) Notes herpetologiques sur l'Indochine francaise. XX. Liste des Reptiles et Batraciens actuellement connus en Indochine Francaise. Bulletin general de l'Instruction Publique, 19 e Annee (1939 - 1940), 4 Decembre 1939, Annexe, 49 - 60." type="book chapter" year="1939">Bourret 1939b: 54</bibRefCitation>
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;
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<bibRefCitation id="D40220A6FF93FFF6FAC3FCF0FE8BCB33" author="Taylor, E. H. & Elbel, R. E." pageId="28" pageNumber="29" pagination="1033 - 1189" refId="ref51290" refString="Taylor, E. H. & Elbel, R. E. (1958) Contribution to the herpetology of Thailand. University of Kansas Science Bulletin, 38 (2), 1033 - 1189. https: // doi. org / 10.5962 / bhl. part. 10972" type="journal article" year="1958">Taylor & Elbel 1958: 1158</bibRefCitation>
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;
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<bibRefCitation id="D40220A6FF93FFF6FE7CFCD2FD71CB33" author="Campden-Main, S. M." box="[385,642,804,829]" pageId="28" pageNumber="29" refId="ref46009" refString="Campden-Main, S. M. (1970) A field guide to the Snakes of South Vietnam. Smithsonian Institution, Washington, D. C., 114 pp." type="book" year="1970">Campden-Main 1970: 46</bibRefCitation>
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, Fig.;
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<bibRefCitation id="D40220A6FF93FFF6FD3CFCD2FC90CB32" author="Welch, K. R. G." box="[705,867,804,829]" pageId="28" pageNumber="29" refId="ref52236" refString="Welch, K. R. G. (1988) Snakes of the Orient: a checklist. Robert F. Krieger Publ. Co., Malabar, Florida, vii + 183 pp." type="book" year="1988">Welch 1988: 88</bibRefCitation>
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;
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Orlov
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<emphasis id="82E78145FF93FFF6FB57FCD3FB25CB33" box="[1194,1238,804,828]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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<bibRefCitation id="D40220A6FF93FFF6FEC6FCB5FD91CB54" author="Iskandar, D. T. & Colijn, E." box="[315,610,835,859]" pageId="28" pageNumber="29" refId="ref48267" refString="Iskandar, D. T. & Colijn, E. (2001) A checklist of Southeast Asian and New Guinean Reptiles. Part I: Serpentes. Biodiversity Conservation Project (Indonesian Institute of Sciences - Japan International Cooperation Agency-The Ministry of Forestry), The Gibbon Foundation and Institut of Technology, Bandung, 195 pp." type="book" year="2001">Iskandar & Colijn 2001: 102</bibRefCitation>
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;
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<bibRefCitation id="D40220A6FF93FFF6FD91FCB5FCA3CB54" author="Orlov, N. L. & Ryabov, S. A. & Nguyen, S. V. & Nguyen, T. Q." box="[620,848,835,859]" pageId="28" pageNumber="29" pagination="217 - 240" refId="ref50138" refString="Orlov, N. L., Ryabov, S. A., Nguyen, S. V. & Nguyen, T. Q. (2003) New records and data of the poorly known snakes of Vietnam. Russian Journal of Herpetology, 10 (3), 217 - 240." type="journal article" year="2003">
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Orlov
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;
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<bibRefCitation id="D40220A6FF93FFF6FCA7FCB2FBA1CB54" author="Nguyen, S. V. & Ho, C. T. & Nguyen, T. Q." box="[858,1106,835,860]" pageId="28" pageNumber="29" refId="ref49709" refString="Nguyen, S. V., Ho, C. T. & Nguyen, T. Q. (2005 a) Danh luc Ech nhai va Bo Sat Viet Nam. A checklist of Amphibians and Reptiles of Vietnam. Nha Xuat Ban Nong Nghiep, Hanoi, 180 pp. [in Vietnamese]" type="book" year="2005">
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<emphasis id="82E78145FF93FFF6FC4CFCB2FC2DCB54" box="[945,990,835,859]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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;
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<bibRefCitation id="D40220A6FF93FFF6FABBFCB5FE52CB75" author="Stuart, B. L. & Chuaynkern, Y." pageId="28" pageNumber="29" pagination="3 - 40" refId="ref51045" refString="Stuart, B. L. & Chuaynkern, Y. (2007) A new Opisthotropis (Serpentes: Colubridae: Natricinae) from northeastern Thailand. Current Herpetology, 26, 3 - 40. https: // doi. org / 10.3105 / 1345 - 5834 (2007) 26 [35: ANOSCN] 2.0. CO; 2" type="journal article" year="2007">Stuart & Chuaynkern 2007: 35</bibRefCitation>
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(implicitely, only generic name cited);
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<bibRefCitation id="D40220A6FF93FFF6FCD1FC94FBD0CB75" author="Murphy, J. C. & Mekchai, S. & Cota, M. & Voris, H. K." box="[812,1059,866,890]" pageId="28" pageNumber="29" pagination="169 - 183" refId="ref49260" refString="Murphy, J. C., Chan-ard, T., Mekchai, S., Cota, M. & Voris, H. K. (2008) The Rediscovery of Angel's Stream Snake, Paratapinophis praemaxillaris Angel, 1929 (Reptilia: Serpentes: Natricidae). The Natural History Journal of Chulalongkorn University, 8 (2), 169 - 183." type="journal article" year="2008">
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<bibRefCitation id="D40220A6FF93FFF6FBD1FC94FABACB75" author="Stuart, B. L. & Heatwole, H." box="[1068,1353,866,890]" pageId="28" pageNumber="29" pagination="97 - 106" refId="ref51108" refString="Stuart, B. L. & Heatwole, H. (2008) Country records of snakes from Laos. Hamadryad, 33, 97 - 106." type="journal article" year="2008">Stuart & Heatwole 2008: 97</bibRefCitation>
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<bibRefCitation id="D40220A6FF93FFF6FAAEFC94FE9ACB96" author="Ziegler, T. & David, P. & Vu, T. N." pageId="28" pageNumber="29" pagination="197 - 203" refId="ref53290" refString="Ziegler, T., David, P., & Vu, T. N. (2008) A new natricine snake of the genus Opisthotropis from Tam Dao, Vinh Phuc Province, northern Vietnam (Squamata, Colubridae). Zoosystematics and Evolution, 84 (2), 197 - 203. https: // doi. org / 10.1002 / zoos. 200800004" type="journal article" year="2008">
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<bibRefCitation id="D40220A6FF93FFF6FD90FC69FCACCBB8" author="Teynie, A. & Lottier, A. & David, P. & Nguyen, T. Q. & Vogel, G." box="[621,863,927,951]" pageId="28" pageNumber="29" pagination="165 - 182" refId="ref51342" refString="Teynie, A., Lottier, A., David, P., Nguyen, T. Q. & Vogel, G. (2013) A new species of the genus Opisthotropis G ʾ nther, 1872 from northern Laos (Squamata: Natricidae). Zootaxa, 3774 (2), 165 - 182. https: // doi. org / 10.11646 / zootaxa. 3774.2.4" type="journal article" year="2013">
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Teynié
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<bibRefCitation id="D40220A6FF93FFF6FBAFFC69FAA1CBB8" author="Wallach, V. & Williams, K. L. & Boundy, J." box="[1106,1362,927,951]" pageId="28" pageNumber="29" refId="ref52073" refString="Wallach, V., Williams, K. L. & Boundy, J. (2014) Snakes of the world. A catalogue of living and extinct species. CRC Press, Boca Raton, Florida, xxvii + 1209 pp." type="book" year="2014">
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David
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<emphasis id="82E78145FF93FFF6FF3AFC49FF06CBD9" box="[199,245,958,982]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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. 2015b: 203–205
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<bibRefCitation id="D40220A6FF93FFF6FDE0FC48FCEFCBD9" author="Kizirian, D. & Nguyen, T. Q. & Ngo, T. H. & Le, D. M." box="[541,796,958,982]" pageId="28" pageNumber="29" pagination="1 - 7" refId="ref48356" refString="Kizirian, D., Nguyen, T. Q., Ngo, T. H. & Le, D. M. (2018) Parahelicops, Pararhabdophis, Paraphyly: phylogenetic relationships among certain Southeast Asian natricine snakes (Hebius). American Museum Novitates, 3906, 1 - 7. https: // doi. org / 10.1206 / 3906.1" type="journal article" year="2018">
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Kizirian
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<emphasis id="82E78145FF93FFF6FD85FC49FD56CBD9" box="[632,677,958,982]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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. 2018: 1–3
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;
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<bibRefCitation id="D40220A6FF93FFF6FCD5FC48FC35CBD9" author="Ren, J. & Wang, K. & Nguyen, T. T. & Hoang, C. V. & Zhong, G. & Jiang, K. & Guo, P. & Li, J." box="[808,966,958,982]" pageId="28" pageNumber="29" pagination="31 - 56" refId="ref50439" refString="Ren, J., Wang, K., Nguyen, T. T., Hoang, C. V., Zhong, G., Jiang, K., Guo, P. & Li, J. (2018) Taxonomic re-evaluation of the monotypic genus Pararhabdophis Bourret, 1934 (Squamata: Colubridae: Natricinae) with discovery of its type species, P. chapaensis, from China. Zootaxa, 4486 (1), 31 - 56. https: // doi. org / 10.11646 / zootaxa. 4486.1.2" type="journal article" year="2018">
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Ren
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<emphasis id="82E78145FF93FFF6FCA5FC49FC75CBD9" box="[856,902,958,982]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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. 2018
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</bibRefCitation>
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(implicitely, only generic name cited): 37, 46 (Table 5), 47 & 51.
|
||
</paragraph>
|
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<paragraph id="B02C5D57FF93FFF6FF6AFC0DFBF8CC1C" blockId="28.[151,1437,651,1105]" box="[151,1035,1019,1043]" pageId="28" pageNumber="29">
|
||
<taxonomicName id="779326D4FF93FFF6FF6AFC0DFE53CC1C" authorityName="Smith" authorityYear="1943" box="[151,416,1019,1043]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF93FFF6FF6AFC0DFE53CC1C" box="[151,416,1019,1043]" italics="true" pageId="28" pageNumber="29">Opisthotropis annamensis</emphasis>
|
||
</taxonomicName>
|
||
.—
|
||
<bibRefCitation id="D40220A6FF93FFF6FE40FC0DFD9BCC1C" author="Smith, M. A." box="[445,616,1019,1043]" pageId="28" pageNumber="29" refId="ref50894" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III. Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith 1943: 331</bibRefCitation>
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& 334;
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||
<bibRefCitation id="D40220A6FF93FFF6FD41FC0DFC4DCC1C" author="Bain, R. H. & Hurley, M. M." box="[700,958,1019,1043]" pageId="28" pageNumber="29" pagination="1 - 138" refId="ref45030" refString="Bain, R. H. & Hurley, M. M. (2011) A biogeographic synthesis of the Amphibians and Reptiles of Indochina. Bulletin of the American Museum of Natural History, 360, 1 - 138. https: // doi. org / 10.1206 / 360.1" type="journal article" year="2011">Bain & Hurley 2011: 104</bibRefCitation>
|
||
& 128.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF93FFF6FF6AFBECFC47CC3D" blockId="28.[151,1437,651,1105]" box="[151,948,1050,1074]" pageId="28" pageNumber="29">
|
||
<taxonomicName id="779326D4FF93FFF6FF6AFBECFE75CC3D" box="[151,390,1050,1074]" class="Reptilia" family="Colubridae" genus="Amphiesma" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamense">
|
||
<emphasis id="82E78145FF93FFF6FF6AFBECFE75CC3D" box="[151,390,1050,1074]" italics="true" pageId="28" pageNumber="29">Amphiesma annamense</emphasis>
|
||
</taxonomicName>
|
||
.—
|
||
<bibRefCitation id="D40220A6FF93FFF6FE5EFBECFD7CCC3D" author="Teynie, A. & Lottier, A. & David, P. & Nguyen, T. Q. & Vogel, G." box="[419,655,1050,1074]" pageId="28" pageNumber="29" pagination="165 - 182" refId="ref51342" refString="Teynie, A., Lottier, A., David, P., Nguyen, T. Q. & Vogel, G. (2013) A new species of the genus Opisthotropis G ʾ nther, 1872 from northern Laos (Squamata: Natricidae). Zootaxa, 3774 (2), 165 - 182. https: // doi. org / 10.11646 / zootaxa. 3774.2.4" type="journal article" year="2013">
|
||
Teynié
|
||
<emphasis id="82E78145FF93FFF6FE0DFBEDFDEECC3D" box="[496,541,1050,1074]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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||
. 2013: 178
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||
</bibRefCitation>
|
||
(generic allocation in error).
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF93FFF6FF6AFBCEFCD3CC5E" blockId="28.[151,1437,651,1105]" box="[151,800,1080,1105]" pageId="28" pageNumber="29">
|
||
<taxonomicName id="779326D4FF93FFF6FF6AFBCEFEAECC5F" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[151,349,1080,1104]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF93FFF6FF6AFBCEFEAECC5F" box="[151,349,1080,1104]" italics="true" pageId="28" pageNumber="29">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
.—
|
||
<bibRefCitation id="D40220A6FF93FFF6FE87FBCEFDADCC5F" author="Kizirian, D. & Nguyen, T. Q. & Ngo, T. H. & Le, D. M." box="[378,606,1080,1105]" pageId="28" pageNumber="29" pagination="1 - 7" refId="ref48356" refString="Kizirian, D., Nguyen, T. Q., Ngo, T. H. & Le, D. M. (2018) Parahelicops, Pararhabdophis, Paraphyly: phylogenetic relationships among certain Southeast Asian natricine snakes (Hebius). American Museum Novitates, 3906, 1 - 7. https: // doi. org / 10.1206 / 3906.1" type="journal article" year="2018">
|
||
Kizirian
|
||
<emphasis id="82E78145FF93FFF6FE2BFBCFFDF0CC5F" box="[470,515,1080,1104]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
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||
. 2018: 5
|
||
</bibRefCitation>
|
||
;
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||
<bibRefCitation id="D40220A6FF93FFF6FD94FBCEFCEFCC5E" author="Boundy, J." box="[617,796,1080,1105]" pageId="28" pageNumber="29" refId="ref45410" refString="Boundy, J. (2020) Snakes of the World. A Supplement. CRC Press, Boca Raton - London - New York, vii + 273 pp." type="book" year="2020">Boundy 2020: 91</bibRefCitation>
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.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8890EDCFF93FFF6FF6AFB88FCCCCE62" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="B02C5D57FF93FFF6FF6AFB88FCCCCE62" blockId="28.[151,1437,1150,2041]" pageId="28" pageNumber="29">
|
||
<emphasis id="82E78145FF93FFF6FF6AFB88FE37CC97" bold="true" box="[151,452,1150,1176]" pageId="28" pageNumber="29">Specimens examined (16)</emphasis>
|
||
.—
|
||
<materialsCitation id="00FB570AFF93FFF6FE15FB88FD17CCB2" country="Vietnam" elevation="1500" location="Ba Na Nature Reserve" municipality="City." pageId="28" pageNumber="29" specimenCount="1" stateProvince="Da Nang" typeStatus="holotype">
|
||
<emphasis id="82E78145FF93FFF6FE15FB88FDBDCC97" bold="true" box="[488,590,1150,1176]" pageId="28" pageNumber="29">
|
||
<collectingCountry id="C8841DC7FF93FFF6FE15FB88FDBDCC97" box="[488,590,1150,1176]" name="Vietnam" pageId="28" pageNumber="29">Vietnam</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<collectingRegion id="725793B5FF93FFF6FDA6FB89FD4CCC96" box="[603,703,1151,1177]" country="Vietnam" name="Da Nang" pageId="28" pageNumber="29">Da Nang</collectingRegion>
|
||
<collectingMunicipality id="5048C72DFF93FFF6FD3BFB89FD0FCC96" box="[710,764,1151,1177]" pageId="28" pageNumber="29">City.</collectingMunicipality>
|
||
MNHN 1938.0117 (
|
||
<typeStatus id="6F28E3F5FF93FFF6FC1EFB89FBB4CC96" box="[995,1095,1151,1177]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="779326D4FF93FFF6FB91FB89FA7FCC97" authorityName="Bourret" authorityYear="1934" box="[1132,1420,1151,1177]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF93FFF6FB91FB89FA7FCC97" box="[1132,1420,1151,1177]" italics="true" pageId="28" pageNumber="29">Parahelicops annamensis</emphasis>
|
||
</taxonomicName>
|
||
), “Bana”, now
|
||
<location id="B54C0B8CFF93FFF6FECEFB55FDCBCCB2" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FECEFB55FDCBCCB2" box="[307,568,1187,1213]" country="Vietnam" municipality="City." name="Ba Na Nature Reserve" pageId="28" pageNumber="29" stateProvince="Da Nang">Ba Na Nature Reserve</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FD7FFB54FD17CCB2" box="[642,740,1186,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="m" value="1500.0">
|
||
<elevation id="3BBEBA64FF93FFF6FD7FFB54FD17CCB2" box="[642,740,1186,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="m" value="1500.0">1,500 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="00FB570AFF93FFF6FD0CFB55FEB3CCEE" collectorName="Rao An" country="Vietnam" county="Po Mu forest" elevation="1400" latitude="18.340557" location="Ba Na Nature Reserve" longLatPrecision="21" longitude="105.23694" municipality="Rao An" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Da Nang" typeStatus="holotype">
|
||
ZISP NLO-2831,
|
||
<location id="B54C0B8CFF93FFF6FC43FB55FB30CCB2" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FC43FB55FB30CCB2" box="[958,1219,1187,1213]" country="Vietnam" name="Ba Na Nature Reserve" pageId="28" pageNumber="29" stateProvince="Da Nang">Ba Na Nature Reserve</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FAF0FB55FA9CCCB2" box="[1293,1391,1187,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="29" unit="m" value="1400.0">
|
||
<elevation id="3BBEBA64FF93FFF6FAF0FB55FA9CCCB2" box="[1293,1391,1187,1213]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="28" pageNumber="29" unit="m" value="1400.0">1,400 m</elevation>
|
||
</quantity>
|
||
.
|
||
<collectingRegion id="725793B5FF93FFF6FA81FB55FECFCCEE" country="Vietnam" name="Ha Tinh" pageId="28" pageNumber="29">Ha Tinh Province</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FEBAFB31FA6ECCEE" box="[327,1437,1222,1249]" collectorName="Rao An" country="Vietnam" county="Po Mu forest" elevation="870" latitude="18.340557" location="Huong Son District" longLatPrecision="21" longitude="105.23694" municipality="Rao An" pageId="28" pageNumber="29" specimenCount="300" stateProvince="Ha Tinh" typeStatus="holotype">
|
||
AMNH 147129,
|
||
<collectingCounty id="594D25DBFF93FFF6FDF7FB31FD56CCEE" box="[522,677,1223,1249]" pageId="28" pageNumber="29">Po Mu forest</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FD4FFB31FCFFCCEE" box="[690,780,1223,1249]" pageId="28" pageNumber="29">Rao An</collectingMunicipality>
|
||
,
|
||
<location id="B54C0B8CFF93FFF6FCE4FB31FC0BCCEE" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FCE4FB31FC0BCCEE" box="[793,1016,1223,1249]" country="Vietnam" county="Po Mu forest" latitude="18.340557" longLatPrecision="21" longitude="105.23694" municipality="Rao An" name="Huong Son District" pageId="28" pageNumber="29" stateProvince="Ha Tinh">Huong Son District</location>
|
||
(
|
||
<geoCoordinate id="D5A73B90FF93FFF6FBF7FB31FB67CCEE" box="[1034,1172,1223,1249]" degrees="18" direction="north" minutes="20" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="26" value="18.340557">18°20’26”N</geoCoordinate>
|
||
<geoCoordinate id="D5A73B90FF93FFF6FB60FB30FAC0CCEE" box="[1181,1331,1222,1249]" degrees="105" direction="east" minutes="14" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="13" value="105.23694">105°14’13”E</geoCoordinate>
|
||
,
|
||
<quantity id="776BF0B2FF93FFF6FABFFB31FA7CCCEE" box="[1346,1423,1223,1249]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="28" pageNumber="29" unit="m" value="870.0">
|
||
<elevation id="3BBEBA64FF93FFF6FABFFB31FA7CCCEE" box="[1346,1423,1223,1249]" metricMagnitude="2" metricUnit="m" metricValue="8.7" pageId="28" pageNumber="29" unit="m" value="870.0">870 m</elevation>
|
||
</quantity>
|
||
);
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FF6AFB1DFBE3CD0A" box="[151,1040,1259,1285]" collectorName="Rao An" country="Vietnam" county="Po Mu forest" elevation="870" latitude="18.340557" location="Huong Son District" longLatPrecision="21" longitude="105.23694" municipality="Rao An" pageId="28" pageNumber="29" specimenCount="300" stateProvince="Ha Tinh" typeStatus="holotype">
|
||
ZISP NLO-2478,
|
||
<collectorName id="1D663881FF93FFF6FEA2FB1DFE45CD0A" box="[351,438,1259,1285]" pageId="28" pageNumber="29">Rao An</collectorName>
|
||
,
|
||
<location id="B54C0B8CFF93FFF6FE3DFB1DFD6FCD0A" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FE3DFB1DFD6FCD0A" box="[448,668,1259,1285]" country="Vietnam" county="Po Mu forest" latitude="18.340557" longLatPrecision="21" longitude="105.23694" municipality="Rao An" name="Huong Son District" pageId="28" pageNumber="29" stateProvince="Ha Tinh">Huong Son District</location>
|
||
, elev.
|
||
<specimenCount id="A69596DEFF93FFF6FD1CFB1DFCDDCD0A" box="[737,814,1259,1285]" pageId="28" pageNumber="29" type="generic">
|
||
<quantity id="776BF0B2FF93FFF6FD1CFB1DFCD9CD0A" box="[737,810,1259,1285]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="300.0">
|
||
<elevation id="3BBEBA64FF93FFF6FD1CFB1DFCD9CD0A" box="[737,810,1259,1285]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="28" pageNumber="29" unit="m" value="300.0">300 m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
<collectingRegion id="725793B5FF93FFF6FCC9FB1DFBFFCD0A" box="[820,1036,1259,1285]" country="Vietnam" name="Kon Tum" pageId="28" pageNumber="29">Kon Tum Province</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FBEBFB1DFAA6CD26" collectorName="Rao An" country="Vietnam" county="Po Mu forest" elevation="870" latitude="18.340557" location="Huong Son District" longLatPrecision="21" longitude="105.23694" municipality="Rao An" pageId="28" pageNumber="29" specimenCount="300" stateProvince="Ha Tinh" typeStatus="holotype">
|
||
IEBR 371, ZISP NLO-3435,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FAA6FB1DFED5CD26" pageId="28" pageNumber="29">Mang Canh village</collectingMunicipality>
|
||
(14°41’950N, 108°14’642E),
|
||
<location id="B54C0B8CFF93FFF6FD7DFAF9FCABCD26" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FD7DFAF9FCABCD26" box="[640,856,1295,1321]" country="Vietnam" latitude="1.2" longLatPrecision="78" longitude="-1.25" municipality="Mang Canh village" name="Kon Plong District" pageId="28" pageNumber="29" stateProvince="Kon Tum">Kon Plong District</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FC5DFAF8FBBECD26" box="[928,1101,1294,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="28" pageNumber="29" unit="m" value="1225.0" valueMax="1250.0" valueMin="1200.0">
|
||
<elevation id="3BBEBA64FF93FFF6FC5DFAF8FBBECD26" box="[928,1101,1294,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.225" metricValueMax="1.25" metricValueMin="1.2" pageId="28" pageNumber="29" unit="m" value="1225.0" valueMax="1250.0" valueMin="1200.0">
|
||
<geoCoordinate id="D5A73B90FF93FFF6FC5DFAF8FC2CCD26" box="[928,991,1294,1321]" degrees="1,200" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="55" value="1.2">1,200</geoCoordinate>
|
||
<geoCoordinate id="D5A73B90FF93FFF6FC22FAF8FBDFCD26" box="[991,1068,1294,1321]" degrees="1,250" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="55" value="-1.25">–1,250</geoCoordinate>
|
||
m
|
||
</elevation>
|
||
</quantity>
|
||
.
|
||
<collectingRegion id="725793B5FF93FFF6FBA4FAF9FAA2CD26" box="[1113,1361,1295,1321]" country="Vietnam" name="Quang Binh" pageId="28" pageNumber="29">Quang Binh Province</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FAA1FAF9FF00CD7E" country="Vietnam" elevation="696" latitude="16.9684" location="Dong Chau Forest" longLatPrecision="1" longitude="106.59299" municipality="Field" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Quang Binh" typeStatus="holotype">
|
||
IEBR 4580 (
|
||
<collectingMunicipality id="5048C72DFF93FFF6FF1CFAC5FEEFCD42" box="[225,284,1331,1357]" pageId="28" pageNumber="29">Field</collectingMunicipality>
|
||
number QB.2015.377),
|
||
<location id="B54C0B8CFF93FFF6FDC5FAC5FCFCCD42" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FDC5FAC5FCFCCD42" box="[568,783,1331,1357]" country="Vietnam" latitude="16.9684" longLatPrecision="1" longitude="106.59299" municipality="Field" name="Dong Chau Forest" pageId="28" pageNumber="29" stateProvince="Quang Binh">Dong Chau Forest</location>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FCE0FAC4FC42CD42" box="[797,945,1330,1357]" degrees="16" direction="north" minutes="58.104" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="16.9684">16°58.104’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FC3DFAC4FBACCD42" box="[960,1119,1330,1357]" degrees="106" direction="east" minutes="35.579" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="106.59299">106°35.579’E</geoCoordinate>
|
||
,
|
||
<elevation id="3BBEBA64FF93FFF6FB93FAC5FAD8CD42" box="[1134,1323,1331,1357]" metricMagnitude="2" metricUnit="m" metricValue="6.959999999999999" pageId="28" pageNumber="29" unit="m" value="696.0">
|
||
elevation
|
||
<quantity id="776BF0B2FF93FFF6FB22FAC5FAD8CD42" box="[1247,1323,1331,1357]" metricMagnitude="2" metricUnit="m" metricValue="6.959999999999999" pageId="28" pageNumber="29" unit="m" value="696.0">696 m</quantity>
|
||
</elevation>
|
||
,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FAC7FAC5FF1CCD7E" pageId="28" pageNumber="29">Lê Thuy District</collectingMunicipality>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FF06FAA1FC6DCD9A" country="Vietnam" elevation="527" latitude="17.7109" location="Phong Nha - Ke Bang National Park" longLatPrecision="1" longitude="105.87846" municipality="Le Thuy District" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Quang Binh Province" typeStatus="holotype">
|
||
ZFMK 86457, ZISP NLO-2714,
|
||
<location id="B54C0B8CFF93FFF6FD92FAA1FBE3CD7E" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FD92FAA1FBE3CD7E" box="[623,1040,1367,1393]" country="Vietnam" latitude="17.7109" longLatPrecision="1" longitude="105.87846" municipality="Le Thuy District" name="Phong Nha - Ke Bang National Park" pageId="28" pageNumber="29" stateProvince="Quang Binh Province">Phong Nha - Ke Bang National Park</location>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FBE1FAA0FB42CD7E" box="[1052,1201,1366,1393]" degrees="17" direction="north" minutes="42.654" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="17.7109">17°42.654’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FB40FAA0FAAFCD7E" box="[1213,1372,1366,1393]" degrees="105" direction="east" minutes="52.708" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="105.87846">105°52.708’E</geoCoordinate>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FF6AFA8CFF12CD9A" box="[151,225,1402,1429]" metricMagnitude="2" metricUnit="m" metricValue="5.27" pageId="28" pageNumber="29" unit="m" value="527.0">
|
||
<elevation id="3BBEBA64FF93FFF6FF6AFA8CFF12CD9A" box="[151,225,1402,1429]" metricMagnitude="2" metricUnit="m" metricValue="5.27" pageId="28" pageNumber="29" unit="m" value="527.0">527 m</elevation>
|
||
</quantity>
|
||
, Hoa Son Commune,
|
||
<location id="B54C0B8CFF93FFF6FE26FA8DFD5ECD9A" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FE26FA8DFD5ECD9A" box="[475,685,1403,1429]" country="Vietnam" name="Minh Hoa District" pageId="28" pageNumber="29" stateProvince="Quang Binh Province">Minh Hoa District</location>
|
||
.
|
||
<collectingRegion id="725793B5FF93FFF6FD45FA8DFC69CD9A" box="[696,922,1403,1429]" country="Vietnam" name="Quang Tri" pageId="28" pageNumber="29">Quang Tri Province</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FC58FA8DFBEACDB6" country="Vietnam" elevation="527" latitude="17.7109" location="Phong Nha - Ke Bang National Park" longLatPrecision="1" longitude="105.87846" municipality="Le Thuy District" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Quang Binh Province" typeStatus="holotype">
|
||
ZISP NLO-3257,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FB92FA8DFB26CD9A" box="[1135,1237,1403,1429]" pageId="28" pageNumber="29">Ban Cup</collectingMunicipality>
|
||
,
|
||
<location id="B54C0B8CFF93FFF6FB1FFA8DFF34CDB6" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FB1FFA8DFF34CDB6" country="Vietnam" latitude="16.957233" longLatPrecision="1" longitude="106.58881" municipality="Ban Cup" name="Huong Hoa District" pageId="28" pageNumber="29" stateProvince="Quang Tri">Huong Hoa District</location>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FF29FA68FE9BCDB6" box="[212,360,1438,1465]" degrees="16" direction="north" minutes="57.434" orientation="latitude" pageId="28" pageNumber="29" precision="1" value="16.957233">16°57.434’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FE88FA68FDE7CDB6" box="[373,532,1438,1465]" degrees="106" direction="east" minutes="35.329" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="106.58881">106°35.329’E</geoCoordinate>
|
||
, elev.
|
||
<elevation id="3BBEBA64FF93FFF6FDA0FA69FD10CDB6" box="[605,739,1439,1465]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="28" pageNumber="29" unit="m" value="490.0">
|
||
<quantity id="776BF0B2FF93FFF6FDA0FA69FD55CDB6" box="[605,678,1439,1465]" metricMagnitude="2" metricUnit="m" metricValue="4.9" pageId="28" pageNumber="29" unit="m" value="490.0">490 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
<collectingRegion id="725793B5FF93FFF6FD17FA69FBE7CDB6" box="[746,1044,1439,1465]" country="Vietnam" name="Thua Thien-Hue" pageId="28" pageNumber="29">Thua Thien-Hue Province</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FBDDFA69FF21CDD2" country="Vietnam" elevation="527" latitude="17.7109" location="Phong Nha - Ke Bang National Park" longLatPrecision="1" longitude="105.87846" municipality="Le Thuy District" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Quang Binh Province" typeStatus="holotype">
|
||
ZFMK 83513,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FB31FA69FF3DCDD2" pageId="28" pageNumber="29">Bach Ma National Park</collectingMunicipality>
|
||
;
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FF25FA35FCB4CDD2" box="[216,839,1474,1501]" country="Vietnam" elevation="527" latitude="17.7109" location="Phong Nha - Ke Bang National Park" longLatPrecision="1" longitude="105.87846" municipality="Le Thuy District" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Quang Binh Province" typeStatus="holotype">
|
||
ZISP NLO-2790,
|
||
<location id="B54C0B8CFF93FFF6FE62FA35FD54CDD2" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FE62FA35FD54CDD2" box="[415,679,1475,1501]" country="Vietnam" municipality="Bach Ma National Park" name="Bach Ma National Park" pageId="28" pageNumber="29" stateProvince="Thua Thien-Hue">Bach Ma National Park</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FD16FA34FCB4CDD2" box="[747,839,1474,1501]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="28" pageNumber="29" unit="m" value="1350.0">
|
||
<elevation id="3BBEBA64FF93FFF6FD16FA34FCB4CDD2" box="[747,839,1474,1501]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="28" pageNumber="29" unit="m" value="1350.0">1,350 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.—
|
||
<materialsCitation id="00FB570AFF93FFF6FC94FA35FB43CE0E" country="Laos" county="Houay Tad Seua" elevation="1245" latitude="1.28" location="Pakxong District" longLatPrecision="78" longitude="-1.5" municipality="Dong Hua Sao" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Champasak" typeStatus="holotype">
|
||
<emphasis id="82E78145FF93FFF6FC94FA35FC50CDD2" bold="true" box="[873,931,1475,1501]" pageId="28" pageNumber="29">
|
||
<collectingCountry id="C8841DC7FF93FFF6FC94FA35FC50CDD2" box="[873,931,1475,1501]" name="Laos" pageId="28" pageNumber="29">Laos</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<collectingRegion id="725793B5FF93FFF6FC4DFA35FB6CCDD2" box="[944,1183,1475,1501]" country="Laos" name="Champasak" pageId="28" pageNumber="29">Champasak Province</collectingRegion>
|
||
. NCSM 77919.
|
||
<collectingCounty id="594D25DBFF93FFF6FAAFFA35FEF2CE0E" pageId="28" pageNumber="29">Houay Tad Seua</collectingCounty>
|
||
stream,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FE94FA11FDE6CE0E" box="[361,533,1511,1537]" pageId="28" pageNumber="29">Dong Hua Sao</collectingMunicipality>
|
||
NPA,
|
||
<location id="B54C0B8CFF93FFF6FD9AFA11FCD9CE0E" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FD9AFA11FCD9CE0E" box="[615,810,1511,1537]" country="Laos" county="Houay Tad Seua" municipality="Dong Hua Sao" name="Pakxong District" pageId="28" pageNumber="29" stateProvince="Champasak">Pakxong District</location>
|
||
,
|
||
<elevation id="3BBEBA64FF93FFF6FCC4FA10FC2BCE0E" box="[825,984,1510,1537]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="28" pageNumber="29" unit="m" value="1245.0">
|
||
<quantity id="776BF0B2FF93FFF6FCC4FA10FC6ACE0E" box="[825,921,1510,1537]" metricMagnitude="3" metricUnit="m" metricValue="1.2449999999999999" pageId="28" pageNumber="29" unit="m" value="1245.0">1,245 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
<collectingRegion id="725793B5FF93FFF6FC1FFA11FB5FCE0E" box="[994,1196,1511,1537]" country="Laos" name="Xekong" pageId="28" pageNumber="29">Xékong Province</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="00FB570AFF93FFF6FB47FA11FA6BCE2A" country="Laos" elevation="1390" latitude="1.28" location="Kaleum District" longLatPrecision="78" longitude="-1.5" municipality="Xe Sap National Biodiversity Conservation Area" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Xekong" typeStatus="holotype">
|
||
FMNH 258637,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FA86FA11FD68CE2A" pageId="28" pageNumber="29">Xe Sap National Biodiversity Conservation Area</collectingMunicipality>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FD55F9FDFCCBCE2A" box="[680,824,1547,1573]" degrees="16" direction="north" minutes="04" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="10" value="16.069445">16°04’10’’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D5A73B90FF93FFF6FCB8F9FCFC28CE2A" box="[837,987,1546,1573]" degrees="106" direction="east" minutes="58" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="45" value="106.979164">106°58’45”E</geoCoordinate>
|
||
,
|
||
<location id="B54C0B8CFF93FFF6FC14F9FDFB52CE2A" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FC14F9FDFB52CE2A" box="[1001,1185,1547,1573]" country="Laos" latitude="1.28" longLatPrecision="78" longitude="-1.5" municipality="Xe Sap National Biodiversity Conservation Area" name="Kaleum District" pageId="28" pageNumber="29" stateProvince="Xekong">Kaleum District</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FB17F9FCFA6BCE2A" box="[1258,1432,1546,1573]" metricMagnitude="3" metricUnit="m" metricValue="1.39" metricValueMax="1.5" metricValueMin="1.28" pageId="28" pageNumber="29" unit="m" value="1390.0" valueMax="1500.0" valueMin="1280.0">
|
||
<elevation id="3BBEBA64FF93FFF6FB17F9FCFA6BCE2A" box="[1258,1432,1546,1573]" metricMagnitude="3" metricUnit="m" metricValue="1.39" metricValueMax="1.5" metricValueMin="1.28" pageId="28" pageNumber="29" unit="m" value="1390.0" valueMax="1500.0" valueMin="1280.0">1,280–1,500 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="00FB570AFF93FFF6FF6AF9D9FB98CE46" box="[151,1131,1582,1609]" country="Laos" elevation="1475" latitude="15.68239" location="Dakchung District" municipality="Phou Ajol" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Xekong Province" typeStatus="holotype">
|
||
NCSM 78630,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FEBEF9D9FE4BCE46" box="[323,440,1583,1609]" pageId="28" pageNumber="29">Phou Ajol</collectingMunicipality>
|
||
, 15.68239°N, 107°1942°E,
|
||
<location id="B54C0B8CFF93FFF6FD0EF9D9FC37CE46" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FD0EF9D9FC37CE46" box="[755,964,1583,1609]" country="Laos" latitude="15.68239" municipality="Phou Ajol" name="Dakchung District" pageId="28" pageNumber="29" stateProvince="Xekong Province">Dakchung District</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FBF1F9D8FB98CE46" box="[1036,1131,1582,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.475" pageId="28" pageNumber="29" unit="m" value="1475.0">
|
||
<elevation id="3BBEBA64FF93FFF6FBF1F9D8FB98CE46" box="[1036,1131,1582,1609]" metricMagnitude="3" metricUnit="m" metricValue="1.475" pageId="28" pageNumber="29" unit="m" value="1475.0">1,475 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="00FB570AFF93FFF6FB85F9D9FCC8CE62" country="Laos" elevation="1450" latitude="15.68516" location="Dakchung District" municipality="Phou Ajol" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Xekong Province" typeStatus="holotype">
|
||
NCSM 78632,
|
||
<collectingMunicipality id="5048C72DFF93FFF6FAD9F9D9FA6BCE46" box="[1316,1432,1583,1609]" pageId="28" pageNumber="29">Phou Ajol</collectingMunicipality>
|
||
, 15.68516°N, 107°1981°E,
|
||
<location id="B54C0B8CFF93FFF6FE38F9A5FD65CE62" LSID="urn:lsid:plazi:treatment:383AEC41FF93FFC8FF6AFDD5FDB4CE5E:B54C0B8CFF93FFF6FE38F9A5FD65CE62" box="[453,662,1619,1645]" country="Laos" latitude="15.68516" municipality="Phou Ajol" name="Dakchung District" pageId="28" pageNumber="29" stateProvince="Xekong Province">Dakchung District</location>
|
||
, elev.
|
||
<quantity id="776BF0B2FF93FFF6FD21F9A4FCC8CE62" box="[732,827,1618,1645]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="28" pageNumber="29" unit="m" value="1450.0">
|
||
<elevation id="3BBEBA64FF93FFF6FD21F9A4FCC8CE62" box="[732,827,1618,1645]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="28" pageNumber="29" unit="m" value="1450.0">1,450 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8890EDCFF93FFF7FF3AF980FCE4CBAA" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="B02C5D57FF93FFF6FF3AF980FDFBCFBE" blockId="28.[151,1437,1150,2041]" pageId="28" pageNumber="29">
|
||
<emphasis id="82E78145FF93FFF6FF3AF980FE20CE9E" bold="true" box="[199,467,1654,1681]" pageId="28" pageNumber="29">Taxonomic comments.</emphasis>
|
||
—
|
||
<bibRefCitation id="D40220A6FF93FFF6FE12F981FD50CE9E" author="Bourret, R." box="[495,675,1655,1681]" pageId="28" pageNumber="29" pagination="167 - 176" refId="ref45586" refString="Bourret, R. (1934 c) Notes herpetologiques sur l'Indochine francaise. III. Ophidiens d'Annam et du Moyen Laos. Bulletin general de l'Instruction Publique, 13 e Annee (1933 - 1934), 9 Mai 1934, 167 - 176. [Separate reprint: 3 - 12]" type="book chapter" year="1934">Bourret (1934c)</bibRefCitation>
|
||
erected the genus
|
||
<taxonomicName id="779326D4FF93FFF6FC8BF981FBF8CE9E" authorityName="Bourret" authorityYear="1934" box="[886,1035,1655,1681]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF93FFF6FC8BF981FBF8CE9E" box="[886,1035,1655,1681]" italics="true" pageId="28" pageNumber="29">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
for the sole included species
|
||
<taxonomicName id="779326D4FF93FFF6FAA6F981FDD8CEBA" authority="Bourret, 1934" authorityName="Bourret" authorityYear="1934" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF93FFF6FAA6F981FE72CEBB" italics="true" pageId="28" pageNumber="29">Parahelicops annamensis</emphasis>
|
||
Bourret, 1934
|
||
</taxonomicName>
|
||
, described in the same paper from a single specimen, on the basis of the following characters: 25 subequal maxillary teeth, of which the two last ones are enlarged, a small eye with a round pupil, nostrils directed upwards, and a single prefrontal; other characters included a head distinct from the neck, 2 internasals, body elongate, slightly laterally compressed, dorsal scales keeled, without apical pits, in 15 rows, tail long, subcaudals paired and hypapophyses developped throughout the vertebral column. The position of the genus
|
||
<taxonomicName id="779326D4FF93FFF6FF6AF8B9FEDFCF66" authorityName="Bourret" authorityYear="1934" box="[151,300,1871,1897]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF93FFF6FF6AF8B9FEDFCF66" box="[151,300,1871,1897]" italics="true" pageId="28" pageNumber="29">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
has long been controversial (
|
||
<bibRefCitation id="D40220A6FF93FFF6FD86F8B9FCA6CF66" author="David, P. & Pauwels, O. S. G. & Nguyen, T. Q. & Vogel, G." box="[635,853,1870,1897]" pageId="28" pageNumber="29" pagination="203 - 217" refId="ref47072" refString="David, P., Pauwels, O. S. G., Nguyen, T. Q. & Vogel, G. (2015 b) On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae). Zootaxa, 3948 (2), 203 - 217. https: // doi. org / 10.11646 / zootaxa. 3948.2.3" type="journal article" year="2015">
|
||
David
|
||
<emphasis id="82E78145FF93FFF6FD34F8B9FD08CF66" box="[713,763,1871,1897]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
. 2015b
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="D40220A6FF93FFF6FC94F8B9FBEECF66" author="Bourret, R." box="[873,1053,1871,1897]" pageId="28" pageNumber="29" pagination="167 - 176" refId="ref45586" refString="Bourret, R. (1934 c) Notes herpetologiques sur l'Indochine francaise. III. Ophidiens d'Annam et du Moyen Laos. Bulletin general de l'Instruction Publique, 13 e Annee (1933 - 1934), 9 Mai 1934, 167 - 176. [Separate reprint: 3 - 12]" type="book chapter" year="1934">Bourret (1934c)</bibRefCitation>
|
||
stated that this genus was similar to members of the genus
|
||
<taxonomicName id="779326D4FF93FFF6FE4DF885FDBACF82" box="[432,585,1907,1933]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF93FFF6FE4DF885FDBACF82" box="[432,585,1907,1933]" italics="true" pageId="28" pageNumber="29">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
G̹nther, 1872 but differed by its dentition, its head clearly distinct from the neck and its more elongate body.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF93FFF7FF3AF84DFA82C912" blockId="28.[151,1437,1150,2041]" lastBlockId="29.[151,1437,150,2013]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
|
||
<bibRefCitation id="D40220A6FF93FFF6FF3AF84DFE64CFDA" author="Smith, M. A." box="[199,407,1979,2005]" pageId="28" pageNumber="29" refId="ref50894" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III. Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith (1943: 334)</bibRefCitation>
|
||
, although recognizing the validity of the characters on which
|
||
<bibRefCitation id="D40220A6FF93FFF6FBB3F84DFAF1CFDA" author="Bourret, R." box="[1102,1282,1979,2005]" pageId="28" pageNumber="29" pagination="167 - 176" refId="ref45586" refString="Bourret, R. (1934 c) Notes herpetologiques sur l'Indochine francaise. III. Ophidiens d'Annam et du Moyen Laos. Bulletin general de l'Instruction Publique, 13 e Annee (1933 - 1934), 9 Mai 1934, 167 - 176. [Separate reprint: 3 - 12]" type="book chapter" year="1934">Bourret (1934c)</bibRefCitation>
|
||
based his description, synonymized
|
||
<taxonomicName id="779326D4FF93FFF6FE59F829FDCACFF6" authorityName="Bourret" authorityYear="1934" box="[420,569,2015,2041]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF93FFF6FE59F829FDCACFF6" box="[420,569,2015,2041]" italics="true" pageId="28" pageNumber="29">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
with the genus
|
||
<taxonomicName id="779326D4FF93FFF6FD10F829FC75CFF6" box="[749,902,2015,2041]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF93FFF6FD10F829FC75CFF6" box="[749,902,2015,2041]" italics="true" pageId="28" pageNumber="29">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
in extending the definition of this latter one. In contrast,
|
||
<bibRefCitation id="D40220A6FF92FFF7FF00FF61FDC9C8BE" author="Taylor, E. H. & Elbel, R. E." box="[253,570,150,177]" pageId="29" pageNumber="30" pagination="1033 - 1189" refId="ref51290" refString="Taylor, E. H. & Elbel, R. E. (1958) Contribution to the herpetology of Thailand. University of Kansas Science Bulletin, 38 (2), 1033 - 1189. https: // doi. org / 10.5962 / bhl. part. 10972" type="journal article" year="1958">Taylor & Elbel (1958: 1156)</bibRefCitation>
|
||
accepted the validity of
|
||
<taxonomicName id="779326D4FF92FFF7FCB7FF61FC2CC8BE" authorityName="Bourret" authorityYear="1934" box="[842,991,151,177]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FCB7FF61FC2CC8BE" box="[842,991,151,177]" italics="true" pageId="29" pageNumber="30">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
on the basis of Bourret’s characters and placed in this latter genus a new species from north-eastern
|
||
<collectingCountry id="C8841DC7FF92FFF7FCD0FF4DFC60C8DA" box="[813,915,187,213]" name="Thailand" pageId="29" pageNumber="30">Thailand</collectingCountry>
|
||
,
|
||
<taxonomicName id="779326D4FF92FFF7FC63FF4DFB5CC8DA" authorityName="Taylor & Elbel" authorityYear="1958" box="[926,1199,187,213]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="boonsongi">
|
||
<emphasis id="82E78145FF92FFF7FC63FF4DFB5CC8DA" box="[926,1199,187,213]" italics="true" pageId="29" pageNumber="30">Parahelicops boonsongi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="D40220A6FF92FFF7FB46FF4DFA6FC8DA" author="David, P. & Pauwels, O. S. G. & Nguyen, T. Q. & Vogel, G." box="[1211,1436,186,213]" pageId="29" pageNumber="30" pagination="203 - 217" refId="ref47072" refString="David, P., Pauwels, O. S. G., Nguyen, T. Q. & Vogel, G. (2015 b) On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae). Zootaxa, 3948 (2), 203 - 217. https: // doi. org / 10.11646 / zootaxa. 3948.2.3" type="journal article" year="2015">
|
||
David
|
||
<emphasis id="82E78145FF92FFF7FAFBFF4DFAC4C8DA" box="[1286,1335,187,213]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
. (2015b)
|
||
</bibRefCitation>
|
||
showed that this species is obviously not congeneric either with
|
||
<taxonomicName id="779326D4FF92FFF7FCA0FF29FC08C8F7" authorityName="Bourret" authorityYear="1934" box="[861,1019,223,248]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FCA0FF29FC08C8F7" box="[861,1019,223,248]" italics="true" pageId="29" pageNumber="30">P. annamensis</emphasis>
|
||
</taxonomicName>
|
||
, or with any other natricid genera. As a consequence,
|
||
<taxonomicName id="779326D4FF92FFF7FEB5FEF5FE2BC912" authorityName="Taylor & Elbel" authorityYear="1958" box="[328,472,259,285]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="boonsongi">
|
||
<emphasis id="82E78145FF92FFF7FEB5FEF5FE2BC912" box="[328,472,259,285]" italics="true" pageId="29" pageNumber="30">P. boonsongi</emphasis>
|
||
</taxonomicName>
|
||
was referred to the new genus
|
||
<taxonomicName id="779326D4FF92FFF7FCC8FEF5FA9EC912" authority="David, Pauwels, Nguyen & Vogel, 2015" authorityName="David, Pauwels, Nguyen & Vogel" authorityYear="2015" box="[821,1389,258,285]" class="Reptilia" family="Colubridae" genus="Isanophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FCC8FEF5FC51C912" box="[821,930,259,285]" italics="true" pageId="29" pageNumber="30">Isanophis</emphasis>
|
||
<bibRefCitation id="D40220A6FF92FFF7FC54FEF5FA9EC912" author="David, P. & Pauwels, O. S. G. & Nguyen, T. Q. & Vogel, G." box="[937,1389,258,285]" pageId="29" pageNumber="30" pagination="203 - 217" refId="ref47072" refString="David, P., Pauwels, O. S. G., Nguyen, T. Q. & Vogel, G. (2015 b) On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae). Zootaxa, 3948 (2), 203 - 217. https: // doi. org / 10.11646 / zootaxa. 3948.2.3" type="journal article" year="2015">David, Pauwels, Nguyen & Vogel, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF92FFF7FF3AFED1FAB3CB1A" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
<bibRefCitation id="D40220A6FF92FFF7FF3AFED1FEACC94D" author="Stuart, B. L." box="[199,351,295,322]" pageId="29" pageNumber="30" pagination="167 - 171" refId="ref51006" refString="Stuart, B. L. (2006) A second specimen of Parahelicops annamensis Bourret, 1934 (Serpentes: Colubridae / Natricinae). Hamadryad, 30 (1 - 2), 167 - 171." type="journal article" year="2006">Stuart (2006)</bibRefCitation>
|
||
, although recognizing the validity of the genus
|
||
<taxonomicName id="779326D4FF92FFF7FC8BFED1FBF8C94E" authorityName="Bourret" authorityYear="1934" box="[886,1035,295,321]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FC8BFED1FBF8C94E" box="[886,1035,295,321]" italics="true" pageId="29" pageNumber="30">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
, also suggested a close relationship with
|
||
<taxonomicName id="779326D4FF92FFF7FF32FEBDFE9BC96A" box="[207,360,331,357]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FF32FEBDFE9BC96A" box="[207,360,331,357]" italics="true" pageId="29" pageNumber="30">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
. In contrast,
|
||
<bibRefCitation id="D40220A6FF92FFF7FE05FEBDFD28C96A" author="David, P. & Pauwels, O. S. G. & Nguyen, T. Q. & Vogel, G." box="[504,731,330,357]" pageId="29" pageNumber="30" pagination="203 - 217" refId="ref47072" refString="David, P., Pauwels, O. S. G., Nguyen, T. Q. & Vogel, G. (2015 b) On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae). Zootaxa, 3948 (2), 203 - 217. https: // doi. org / 10.11646 / zootaxa. 3948.2.3" type="journal article" year="2015">
|
||
David
|
||
<emphasis id="82E78145FF92FFF7FDBEFEBDFD80C96A" box="[579,627,331,357]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
. (2015b)
|
||
</bibRefCitation>
|
||
, on the basis of morphological characters, considered the genus
|
||
<taxonomicName id="779326D4FF92FFF7FF6AFE99FEDFC986" authorityName="Bourret" authorityYear="1934" box="[151,300,367,393]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FF6AFE99FEDFC986" box="[151,300,367,393]" italics="true" pageId="29" pageNumber="30">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
to be valid and distinct from
|
||
<taxonomicName id="779326D4FF92FFF7FD8CFE99FCF9C986" box="[625,778,367,393]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FD8CFE99FCF9C986" box="[625,778,367,393]" italics="true" pageId="29" pageNumber="30">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
. Although “
|
||
<taxonomicName id="779326D4FF92FFF7FC6FFE99FBD4C986" authorityName="Bourret" authorityYear="1934" box="[914,1063,367,393]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FC6FFE99FBD4C986" box="[914,1063,367,393]" italics="true" pageId="29" pageNumber="30">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
”
|
||
<taxonomicName id="779326D4FF92FFF7FBC7FE99FB4DC987" authorityName="Bourret" authorityYear="1934" box="[1082,1214,367,392]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FBC7FE99FB4DC987" box="[1082,1214,367,392]" italics="true" pageId="29" pageNumber="30">annamensis</emphasis>
|
||
</taxonomicName>
|
||
has usually a single prefrontal, as in most species of
|
||
<taxonomicName id="779326D4FF92FFF7FDF3FE65FD54C9A2" box="[526,679,403,429]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FDF3FE65FD54C9A2" box="[526,679,403,429]" italics="true" pageId="29" pageNumber="30">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
, it differs from the latter genus, as defined by
|
||
<bibRefCitation id="D40220A6FF92FFF7FB3CFE65FA64C9A2" author="Boulenger, G. A." box="[1217,1431,403,429]" pageId="29" pageNumber="30" refId="ref45260" refString="Boulenger, G. A. (1893 a) Catalogue of the snakes in the British Museum (Natural History), Vol. I., containing the families Typhlopidae, Glauconiidae, Boidae, Ilysiidae, Uropeltidae, Xenopeltidae, and Colubridae Aglyphae, part. British Museum of Natural History, London, xiii + 448 pp., 28 pls." type="book" year="1893">Boulenger (1893a)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D40220A6FF92FFF7FF6AFE41FED6C9DE" author="Pope, C. H." box="[151,293,438,465]" pageId="29" pageNumber="30" pagination="27" refId="ref50250" refString="Pope, C. H. (1935) The Reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia. Vol. X. American Museum of Natural History, New York, xlii + 604, 27 pls." type="journal article" year="1935">Pope (1935)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D40220A6FF92FFF7FEC9FE41FE3EC9DE" author="Smith, M. A." box="[308,461,439,465]" pageId="29" pageNumber="30" refId="ref50894" refString="Smith, M. A. (1943) The Fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese Sub-region. Reptilia and Amphibia. Vol. III. Serpentes. Taylor & Francis, London, xii + 583 pp." type="book" year="1943">Smith (1943)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="D40220A6FF92FFF7FE26FE41FD60C9DD" author="Bourret, R." box="[475,659,439,466]" pageId="29" pageNumber="30" refId="ref45777" refString="Bourret, R. (1936 b) Les Serpents de l'Indochine. Tome II. Catalogue systematique descriptif. Henri Basuyau et Cie, Toulouse, 505 pp." type="book" year="1936">Bourret (1936b)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="D40220A6FF92FFF7FD32FE41FC01C9DD" author="Brown, W. C. & Leviton, A. E." box="[719,1010,439,466]" pageId="29" pageNumber="30" pagination="1 - 5" refId="ref45958" refString="Brown, W. C. & Leviton, A. E. (1961) Discovery of the snake genus Opisthotropis in the Philippine Islands, with description of a new species. Occasional Papers of the Natural History Museum of Stanford University, 8, 1 - 5." type="journal article" year="1961">Brown & Leviton (1961)</bibRefCitation>
|
||
, by several morphological characters such as (1) 2 distinctly enlarged maxillary teeth vs. subequal teeth in
|
||
<taxonomicName id="779326D4FF92FFF7FC43FE2DFBA4C9FA" box="[958,1111,475,501]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FC43FE2DFBA4C9FA" box="[958,1111,475,501]" italics="true" pageId="29" pageNumber="30">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
, (2) dorsolateral nostrils vs. directed distinctly upwards in
|
||
<taxonomicName id="779326D4FF92FFF7FE17FE09FD70CA16" box="[490,643,511,537]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FE17FE09FD70CA16" box="[490,643,511,537]" italics="true" pageId="29" pageNumber="30">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
, and (3) supralabials longer than high vs. distinctly higher than long in
|
||
<taxonomicName id="779326D4FF92FFF7FF6AFDD5FEC3CA32" box="[151,304,547,573]" class="Reptilia" family="Colubridae" genus="Opisthotropis" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FF6AFDD5FEC3CA32" box="[151,304,547,573]" italics="true" pageId="29" pageNumber="30">Opisthotropis</emphasis>
|
||
</taxonomicName>
|
||
. In contrast, some characters of the genus
|
||
<taxonomicName id="779326D4FF92FFF7FCF6FDD5FC53CA32" authorityName="Bourret" authorityYear="1934" box="[779,928,547,573]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FCF6FDD5FC53CA32" box="[779,928,547,573]" italics="true" pageId="29" pageNumber="30">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
, i.e., the dentition of the upper maxilla, made of subequal small teeth followed by two enlarged posterior teeth, the average eye with a round pupil, the dorsolateral nostrils, the keeled dorsal scales, the long, keeled tail, the supralabials longer than high and the general pattern, were obviously typical of the genus
|
||
<taxonomicName id="779326D4FF92FFF7FDCEFD79FD71CAA6" authorityName="Thompson" authorityYear="1913" box="[563,642,655,681]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FDCEFD79FD71CAA6" box="[563,642,655,681]" italics="true" pageId="29" pageNumber="30">Hebius</emphasis>
|
||
</taxonomicName>
|
||
as defined by
|
||
<bibRefCitation id="D40220A6FF92FFF7FCD4FD79FC1CCAA6" author="Guo, P. & Zhu, F. & Liu, Q. & Zhang, L. & Li, J. X. & Huang, Y. Y. & Pyron, R. A." box="[809,1007,655,681]" pageId="29" pageNumber="30" pagination="425 - 440" refId="ref47938" refString="Guo, P., Zhu, F., Liu, Q., Zhang, L., Li, J. X., Huang, Y. Y. & Pyron, R. A. (2014) A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa, 3873 (4), 425 - 440. https: // doi. org / 10.11646 / zootaxa. 3873.4.5" type="journal article" year="2014">
|
||
Guo
|
||
<emphasis id="82E78145FF92FFF7FC9CFD79FC60CAA6" box="[865,915,655,681]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
. (2014)
|
||
</bibRefCitation>
|
||
, and especially the “
|
||
<taxonomicName id="779326D4FF92FFF7FB27FD79FA96CAA6" box="[1242,1381,655,681]" class="Squamata" family="Colubridae" genus="Tropidonotus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="82E78145FF92FFF7FB27FD79FA96CAA6" box="[1242,1381,655,681]" italics="true" pageId="29" pageNumber="30">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
species group”, the group of Indo-Himalayan species which is treated in the present paper. Eventually,
|
||
<bibRefCitation id="D40220A6FF92FFF7FAFDFD45FF17CAFE" author="Kizirian, D. & Nguyen, T. Q. & Ngo, T. H. & Le, D. M." pageId="29" pageNumber="30" pagination="1 - 7" refId="ref48356" refString="Kizirian, D., Nguyen, T. Q., Ngo, T. H. & Le, D. M. (2018) Parahelicops, Pararhabdophis, Paraphyly: phylogenetic relationships among certain Southeast Asian natricine snakes (Hebius). American Museum Novitates, 3906, 1 - 7. https: // doi. org / 10.1206 / 3906.1" type="journal article" year="2018">
|
||
Kizirian
|
||
<emphasis id="82E78145FF92FFF7FA9EFD45FA66CAC2" box="[1379,1429,691,717]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
. (2018)
|
||
</bibRefCitation>
|
||
, on the basis of molecular data, showed that
|
||
<taxonomicName id="779326D4FF92FFF7FD2AFD21FC04CAFF" authorityName="Bourret" authorityYear="1934" box="[727,1015,727,753]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FD2AFD21FC04CAFF" box="[727,1015,727,753]" italics="true" pageId="29" pageNumber="30">Parahelicops annamensis</emphasis>
|
||
</taxonomicName>
|
||
was nested in the genus
|
||
<taxonomicName id="779326D4FF92FFF7FAF6FD21FAA9CAFE" authorityName="Thompson" authorityYear="1913" box="[1291,1370,727,753]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FAF6FD21FAA9CAFE" box="[1291,1370,727,753]" italics="true" pageId="29" pageNumber="30">Hebius</emphasis>
|
||
</taxonomicName>
|
||
and a synonym thereof. We here follow these authors in referring
|
||
<taxonomicName id="779326D4FF92FFF7FCC9FD0DFBA6CB1B" authorityName="Bourret" authorityYear="1934" box="[820,1109,763,789]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FCC9FD0DFBA6CB1B" box="[820,1109,763,789]" italics="true" pageId="29" pageNumber="30">Parahelicops annamensis</emphasis>
|
||
</taxonomicName>
|
||
to the genus
|
||
<taxonomicName id="779326D4FF92FFF7FB17FD0DFACACB1A" authorityName="Thompson" authorityYear="1913" box="[1258,1337,763,789]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FB17FD0DFACACB1A" box="[1258,1337,763,789]" italics="true" pageId="29" pageNumber="30">Hebius</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF92FFF7FF3AFCE9FCE4CBAA" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
For long,
|
||
<taxonomicName id="779326D4FF92FFF7FEC7FCE9FDE4CB37" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[314,535,799,825]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FEC7FCE9FDE4CB37" box="[314,535,799,825]" italics="true" pageId="29" pageNumber="30">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
has been known from its sole
|
||
<typeStatus id="6F28E3F5FF92FFF7FC80FCE9FC2CCB36" box="[893,991,799,825]" pageId="29" pageNumber="30" type="holotype">holotype</typeStatus>
|
||
collected in central
|
||
<collectingCountry id="C8841DC7FF92FFF7FB36FCE9FADDCB36" box="[1227,1326,799,825]" name="Vietnam" pageId="29" pageNumber="30">Vietnam</collectingCountry>
|
||
. The second known specimen, and first record from
|
||
<collectingCountry id="C8841DC7FF92FFF7FD76FCB5FD37CB52" box="[651,708,835,861]" name="Laos" pageId="29" pageNumber="30">Laos</collectingCountry>
|
||
, was described by
|
||
<bibRefCitation id="D40220A6FF92FFF7FC61FCB5FBC5CB51" author="Stuart, B. L." box="[924,1078,835,862]" pageId="29" pageNumber="30" pagination="167 - 171" refId="ref51006" refString="Stuart, B. L. (2006) A second specimen of Parahelicops annamensis Bourret, 1934 (Serpentes: Colubridae / Natricinae). Hamadryad, 30 (1 - 2), 167 - 171." type="journal article" year="2006">Stuart (2006)</bibRefCitation>
|
||
. Subsequently, more specimens became available, especially from central
|
||
<collectingCountry id="C8841DC7FF92FFF7FD8CFC91FD27CB8E" box="[625,724,871,897]" name="Vietnam" pageId="29" pageNumber="30">Vietnam</collectingCountry>
|
||
.
|
||
<bibRefCitation id="D40220A6FF92FFF7FD22FC91FC35CB8E" author="David, P. & Pauwels, O. S. G. & Nguyen, T. Q. & Vogel, G." box="[735,966,870,897]" pageId="29" pageNumber="30" pagination="203 - 217" refId="ref47072" refString="David, P., Pauwels, O. S. G., Nguyen, T. Q. & Vogel, G. (2015 b) On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae). Zootaxa, 3948 (2), 203 - 217. https: // doi. org / 10.11646 / zootaxa. 3948.2.3" type="journal article" year="2015">
|
||
David
|
||
<emphasis id="82E78145FF92FFF7FCD6FC91FCAECB8E" box="[811,861,871,897]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
|
||
. (2015b)
|
||
</bibRefCitation>
|
||
, in the genus
|
||
<taxonomicName id="779326D4FF92FFF7FB9EFC91FB0BCB8E" authorityName="Bourret" authorityYear="1934" box="[1123,1272,871,897]" class="Reptilia" family="Colubridae" genus="Parahelicops" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FB9EFC91FB0BCB8E" box="[1123,1272,871,897]" italics="true" pageId="29" pageNumber="30">Parahelicops</emphasis>
|
||
</taxonomicName>
|
||
, expanded the description of
|
||
<taxonomicName id="779326D4FF92FFF7FEC4FC7DFE2CCBAB" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[313,479,907,932]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FEC4FC7DFE2CCBAB" box="[313,479,907,932]" italics="true" pageId="29" pageNumber="30">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
. This species is monotypic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8890EDCFF92FFCAFF3AFC58FB66CE51" lastPageId="32" lastPageNumber="33" pageId="29" pageNumber="30" type="diagnosis">
|
||
<paragraph id="B02C5D57FF92FFF7FF3AFC58FDE0CDCE" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="82E78145FF92FFF7FF3AFC58FEC9CBC7" bold="true" box="[199,314,942,968]" pageId="29" pageNumber="30">Diagnosis</emphasis>
|
||
.—A moderately sized species of the genus
|
||
<taxonomicName id="779326D4FF92FFF7FCC5FC59FC74CBC6" authorityName="Thompson" authorityYear="1913" box="[824,903,943,969]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FCC5FC59FC74CBC6" box="[824,903,943,969]" italics="true" pageId="29" pageNumber="30">Hebius</emphasis>
|
||
</taxonomicName>
|
||
characterized by the combination of
|
||
<emphasis id="82E78145FF92FFF7FAC7FC58FAA9CBC7" bold="true" box="[1338,1370,942,968]" pageId="29" pageNumber="30">(1)</emphasis>
|
||
body elongate in males and females;
|
||
<emphasis id="82E78145FF92FFF7FE06FC24FDEFCBE3" bold="true" box="[507,540,978,1004]" pageId="29" pageNumber="30">(2)</emphasis>
|
||
head distinct from the neck;
|
||
<emphasis id="82E78145FF92FFF7FC9AFC24FC7BCBE3" bold="true" box="[871,904,978,1004]" pageId="29" pageNumber="30">(3)</emphasis>
|
||
15 or 17 dorsal scale rows at midbody, moderately keeled at midbody, strongly keeled posteriorly, usually smooth on the 1
|
||
<superScript id="47E6F01FFF92FFF7FBFDFC01FBF8CC0A" attach="left" box="[1024,1035,1015,1029]" fontSize="6" pageId="29" pageNumber="30">st</superScript>
|
||
dorsal scale row;
|
||
<emphasis id="82E78145FF92FFF7FB26FC00FB0FCC1F" bold="true" box="[1243,1276,1014,1040]" pageId="29" pageNumber="30">(4)</emphasis>
|
||
scales around the base of the tail very strongly keeled;
|
||
<emphasis id="82E78145FF92FFF7FD95FBECFD7BCC3B" bold="true" box="[616,648,1050,1076]" pageId="29" pageNumber="30">(5)</emphasis>
|
||
eye moderate with a round pupil, 1.2–1.4 times the distance between the lower margins of eye and of lip;
|
||
<emphasis id="82E78145FF92FFF7FDB9FBC8FD97CC57" bold="true" box="[580,612,1086,1112]" pageId="29" pageNumber="30">(6)</emphasis>
|
||
maxillary teeth 28–34, progressively increasing in size in a continuous series, the last two moderately enlarged and not separated by a diastema from anterior teeth;
|
||
<emphasis id="82E78145FF92FFF7FB41FB94FB2ECC73" bold="true" box="[1212,1245,1122,1148]" pageId="29" pageNumber="30">(7)</emphasis>
|
||
nostrils piercing dorsolaterally;
|
||
<emphasis id="82E78145FF92FFF7FEBDFB70FE92CCAF" bold="true" box="[320,353,1158,1184]" pageId="29" pageNumber="30">(8)</emphasis>
|
||
tail long with a ratio Tal/TL equal to 0.30 and above;
|
||
<emphasis id="82E78145FF92FFF7FC38FB70FC15CCAF" bold="true" box="[965,998,1158,1184]" pageId="29" pageNumber="30">(9)</emphasis>
|
||
VEN 158–172;
|
||
<emphasis id="82E78145FF92FFF7FB62FB70FB3ECCAF" bold="true" box="[1183,1229,1158,1184]" pageId="29" pageNumber="30">(10)</emphasis>
|
||
SC 116–146;
|
||
<emphasis id="82E78145FF92FFF7FA92FB70FA6FCCAF" bold="true" box="[1391,1436,1158,1184]" pageId="29" pageNumber="30">(11)</emphasis>
|
||
internasals distinctly narrowing anteriorly;
|
||
<emphasis id="82E78145FF92FFF7FD82FB5CFD5ECCCB" bold="true" box="[639,685,1194,1220]" pageId="29" pageNumber="30">(12)</emphasis>
|
||
prefrontal single in most known specimens,
|
||
<emphasis id="82E78145FF92FFF7FB57FB5CFB29CCCB" bold="true" box="[1194,1242,1194,1220]" pageId="29" pageNumber="30">(13)</emphasis>
|
||
8 or 9 supralabials;
|
||
<emphasis id="82E78145FF92FFF7FF3BFB38FF07CCE7" bold="true" box="[198,244,1230,1256]" pageId="29" pageNumber="30">(14)</emphasis>
|
||
1 anterior temporal,
|
||
<emphasis id="82E78145FF92FFF7FE1FFB38FDE2CCE7" bold="true" box="[482,529,1230,1256]" pageId="29" pageNumber="30">(15)</emphasis>
|
||
venter pale on its inner three quarters, with tips of ventral scales very dark;
|
||
<emphasis id="82E78145FF92FFF7FA93FB38FA6FCCE7" bold="true" box="[1390,1436,1230,1256]" pageId="29" pageNumber="30">(16)</emphasis>
|
||
background colour chestnut brown, dark brown or dark greyish-brown;
|
||
<emphasis id="82E78145FF92FFF7FC3EFB04FC01CD03" bold="true" box="[963,1010,1266,1292]" pageId="29" pageNumber="30">(17)</emphasis>
|
||
dorsal pattern made of isolate, cream or yellowish-ochre blotches (bright orange or rusty-red blotches in life), usually transversally elongate as crossbars anteriorly, then becoming irregular, broken spots posteriorly;
|
||
<emphasis id="82E78145FF92FFF7FCA1FACCFC78CD5B" bold="true" box="[860,907,1338,1364]" pageId="29" pageNumber="30">(18)</emphasis>
|
||
a short and narrow, oblique postocular streak reaching the neck behind the corner of the mouth, sometimes extending on the nape as a broad, oblique and elongate cream or yellowish-ochre streak (orange or rusty-red in life); and
|
||
<emphasis id="82E78145FF92FFF7FC81FA74FC59CD93" bold="true" box="[892,938,1410,1436]" pageId="29" pageNumber="30">(19)</emphasis>
|
||
supralabials dark as the upper head surface, without defined spots or blotches.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF92FFF7FF3AFA3CFBB7CE22" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
<emphasis id="82E78145FF92FFF7FF3AFA3CFEA8CDEB" bold="true" box="[199,347,1482,1508]" pageId="29" pageNumber="30">Comparison</emphasis>
|
||
. In preservative, specimens of
|
||
<taxonomicName id="779326D4FF92FFF7FD39FA3DFC53CDEB" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[708,928,1483,1509]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FD39FA3DFC53CDEB" box="[708,928,1483,1509]" italics="true" pageId="29" pageNumber="30">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
may be easily misidentified as
|
||
<taxonomicName id="779326D4FF92FFF7FAF2FA3DFA6FCDEA" box="[1295,1436,1483,1509]" class="Squamata" family="Colubridae" genus="Tropidonotus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="82E78145FF92FFF7FAF2FA3DFA6FCDEA" box="[1295,1436,1483,1509]" italics="true" pageId="29" pageNumber="30">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
(G̹nther, 1875), an Indo-Himalayan species unknown east of
|
||
<collectingCountry id="C8841DC7FF92FFF7FCA8FA19FC37CE06" box="[853,964,1519,1545]" name="Myanmar" pageId="29" pageNumber="30">Myanmar</collectingCountry>
|
||
. However,
|
||
<taxonomicName id="779326D4FF92FFF7FBBEFA19FB3DCE06" box="[1091,1230,1519,1545]" class="Squamata" family="Colubridae" genus="Tropidonotus" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="82E78145FF92FFF7FBBEFA19FB3DCE06" box="[1091,1230,1519,1545]" italics="true" pageId="29" pageNumber="30">H. modestus</emphasis>
|
||
</taxonomicName>
|
||
has 19 scale rows at midbody vs. 17, 143–163 ventrals vs. 158–172, and a ochre or pale brown venter.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF92FFF7FF3AF9C1FB99CE7A" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
With its typical dorsal and ventral pattern and its long tail,
|
||
<taxonomicName id="779326D4FF92FFF7FC92F9C1FBB8CE5F" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[879,1099,1591,1617]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FC92F9C1FBB8CE5F" box="[879,1099,1591,1617]" italics="true" pageId="29" pageNumber="30">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
cannot be confused with any species of the genus
|
||
<taxonomicName id="779326D4FF92FFF7FE82F9ADFE3DCE7A" authorityName="Thompson" authorityYear="1913" box="[383,462,1627,1653]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="genus">
|
||
<emphasis id="82E78145FF92FFF7FE82F9ADFE3DCE7A" box="[383,462,1627,1653]" italics="true" pageId="29" pageNumber="30">Hebius</emphasis>
|
||
</taxonomicName>
|
||
with 17 dorsal scale rows, from which it differs as follows:
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF92FFF7FF3AF989FDBACF9A" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
<taxonomicName id="779326D4FF92FFF7FF3AF989FE53CE97" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[199,416,1663,1689]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FF3AF989FE53CE97" box="[199,416,1663,1689]" italics="true" pageId="29" pageNumber="30">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
readily differs from
|
||
<taxonomicName id="779326D4FF92FFF7FD78F989FCD1CE96" box="[645,802,1663,1689]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="82E78145FF92FFF7FD78F989FCD1CE96" box="[645,802,1663,1689]" italics="true" pageId="29" pageNumber="30">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
by the characters given above in the account of this latter species. The main characters are (1) internasals distinctly narrowing anteriorly (vs. abruptly truncated); (2) number of subcaudals, 116–146 vs. 90–114 subcaudals in
|
||
<taxonomicName id="779326D4FF92FFF7FD32F931FC9CCEEE" box="[719,879,1735,1761]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="82E78145FF92FFF7FD32F931FC9CCEEE" box="[719,879,1735,1761]" italics="true" pageId="29" pageNumber="30">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
; (3) venter largely pale, with only tips of ventral scales dark vs. nearly entirely dark in
|
||
<taxonomicName id="779326D4FF92FFF7FDBDF91DFD2DCF0A" box="[576,734,1771,1797]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="82E78145FF92FFF7FDBDF91DFD2DCF0A" box="[576,734,1771,1797]" italics="true" pageId="29" pageNumber="30">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
; (4) a distinct dorsal pattern, consisting of pales blotches, usually arranged as crossbars anteriorly in
|
||
<taxonomicName id="779326D4FF92FFF7FDABF8F9FD0DCF27" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[598,766,1807,1832]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FDABF8F9FD0DCF27" box="[598,766,1807,1832]" italics="true" pageId="29" pageNumber="30">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
vs. bright, pale blotches on a faint dorsolateral stripe in
|
||
<taxonomicName id="779326D4FF92FFF7FA7CF8F9FEE0CF42" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="82E78145FF92FFF7FA7CF8F9FEE0CF42" italics="true" pageId="29" pageNumber="30">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
; and (5) head with short and narrow postocular streak in
|
||
<taxonomicName id="779326D4FF92FFF7FC6EF8C5FBC9CF43" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[915,1082,1843,1868]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FC6EF8C5FBC9CF43" box="[915,1082,1843,1868]" italics="true" pageId="29" pageNumber="30">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
, as well as with dark supralabials, without defined blotches vs. strong postocular streak in
|
||
<taxonomicName id="779326D4FF92FFF7FCC9F8A1FC21CF7E" box="[820,978,1879,1905]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="chapaensis">
|
||
<emphasis id="82E78145FF92FFF7FCC9F8A1FC21CF7E" box="[820,978,1879,1905]" italics="true" pageId="29" pageNumber="30">H. chapaensis</emphasis>
|
||
</taxonomicName>
|
||
, with pale supralabials, edged with dark, or divided into a pale and a dark areas.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FF92FFF7FF3AF869FC58CFD2" blockId="29.[151,1437,150,2013]" pageId="29" pageNumber="30">
|
||
<taxonomicName id="779326D4FF92FFF7FF3AF869FE57CFB7" baseAuthorityName="Bourret" baseAuthorityYear="1934" box="[199,420,1951,1977]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF92FFF7FF3AF869FE57CFB7" box="[199,420,1951,1977]" italics="true" pageId="29" pageNumber="30">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
differs from both
|
||
<taxonomicName id="779326D4FF92FFF7FD87F869FCE1CFB7" box="[634,786,1951,1976]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="taronensis">
|
||
<emphasis id="82E78145FF92FFF7FD87F869FCE1CFB7" box="[634,786,1951,1976]" italics="true" pageId="29" pageNumber="30">H. taronensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="779326D4FF92FFF7FCB0F869FC1ECFB7" box="[845,1005,1951,1976]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="nigriventer">
|
||
<emphasis id="82E78145FF92FFF7FCB0F869FC1ECFB7" box="[845,1005,1951,1976]" italics="true" pageId="29" pageNumber="30">H. nigriventer</emphasis>
|
||
</taxonomicName>
|
||
by its venter largely pale vs. entirely black or so in these species, plus by characters given in the key above.
|
||
</paragraph>
|
||
<caption id="E4EC0DDFFF91FFF4FF6AF860FD7DCFDB" ID-DOI="http://doi.org/10.5281/zenodo.4463766" ID-Zenodo-Dep="4463766" httpUri="https://zenodo.org/record/4463766/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,260,1942,1968]" targetBox="[151,1436,179,1917]" targetPageId="30">
|
||
<paragraph id="B02C5D57FF91FFF4FF6AF860FD7DCFDB" blockId="30.[151,1436,1942,2004]" pageId="30" pageNumber="31">
|
||
<emphasis id="82E78145FF91FFF4FF6AF860FEDCCFBF" bold="true" box="[151,303,1942,1968]" pageId="30" pageNumber="31">FIGURE 10.</emphasis>
|
||
<taxonomicName id="779326D4FF91FFF4FEC5F860FD86CFBF" authority="Bourret" authorityName="Bourret" box="[312,629,1942,1968]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF91FFF4FEC5F860FDE6CFBF" box="[312,533,1942,1968]" italics="true" pageId="30" pageNumber="31">Hebius annamensis</emphasis>
|
||
Bourret
|
||
</taxonomicName>
|
||
,
|
||
<typeStatus id="6F28E3F5FF91FFF4FD7EF860FD16CFBF" box="[643,741,1942,1968]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
|
||
(MNHN 1938.0117)—A: dorsal view; B: lateral view of the head, right side. Photographs: Patrick David.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E4EC0DDFFF90FFF5FF6AF863FD51CFF7" ID-DOI="http://doi.org/10.5281/zenodo.4463768" ID-Zenodo-Dep="4463768" httpUri="https://zenodo.org/record/4463768/files/figure.png" pageId="31" pageNumber="32" startId="31.[151,260,1941,1967]" targetBox="[330,1257,180,1917]" targetPageId="31">
|
||
<paragraph id="B02C5D57FF90FFF5FF6AF863FD51CFF7" blockId="31.[151,1437,1941,2040]" pageId="31" pageNumber="32">
|
||
<emphasis id="82E78145FF90FFF5FF6AF863FEDCCFBF" bold="true" box="[151,303,1941,1968]" pageId="31" pageNumber="32">FIGURE 11.</emphasis>
|
||
<taxonomicName id="779326D4FF90FFF5FEC4F860FD8BCFBF" authority="Bourret" authorityName="Bourret" box="[313,632,1942,1968]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FF90FFF5FEC4F860FDE5CFBF" box="[313,534,1942,1968]" italics="true" pageId="31" pageNumber="32">Hebius annamensis</emphasis>
|
||
Bourret
|
||
</taxonomicName>
|
||
, alive—A: specimen from
|
||
<collectingRegion id="725793B5FF90FFF5FC4FF860FB65CFBF" box="[946,1174,1942,1968]" country="Vietnam" name="Quang Tri" pageId="31" pageNumber="32">Quang Tri Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="C8841DC7FF90FFF5FB59F860FAF4CFBF" box="[1188,1287,1942,1968]" name="Vietnam" pageId="31" pageNumber="32">Vietnam</collectingCountry>
|
||
. Photograph: Nikolai Orlov. B: specimen from
|
||
<collectingRegion id="725793B5FF90FFF5FDE5F84CFCE2CFDB" box="[536,785,1978,2004]" country="Vietnam" name="Quang Binh" pageId="31" pageNumber="32">Quang Binh Province</collectingRegion>
|
||
. Photograph: Nikolai Orlov. C: specimen from
|
||
<collectingRegion id="725793B5FF90FFF5FACDF84CFF0DCFF7" country="Vietnam" name="Kon Tum" pageId="31" pageNumber="32">Kon Tum Province</collectingRegion>
|
||
. Photograph: Truong Quang Nguyen.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E4EC0DDFFFAFFFCAFF6AFB7DFE41CCE1" ID-DOI="http://doi.org/10.5281/zenodo.4463772" ID-Zenodo-Dep="4463772" httpUri="https://zenodo.org/record/4463772/files/figure.png" pageId="32" pageNumber="33" startId="32.[151,260,1163,1189]" targetBox="[226,1360,191,1129]" targetPageId="32">
|
||
<paragraph id="B02C5D57FFAFFFCAFF6AFB7DFE41CCE1" blockId="32.[151,1437,1163,1262]" pageId="32" pageNumber="33">
|
||
<emphasis id="82E78145FFAFFFCAFF6AFB7DFEDECCAA" bold="true" box="[151,301,1163,1189]" pageId="32" pageNumber="33">FIGURE 12.</emphasis>
|
||
Distribution map of species with 19 dorsal scale rows at midbody.
|
||
<emphasis id="82E78145FFAFFFCAFBEBFB7DFB9ACCAA" bold="true" box="[1046,1129,1163,1189]" pageId="32" pageNumber="33">Circles</emphasis>
|
||
:
|
||
<taxonomicName id="779326D4FFAFFFCAFB8AFB7DFAC6CCAA" baseAuthorityName="Gunther" baseAuthorityYear="1875" box="[1143,1333,1163,1189]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="modestus">
|
||
<emphasis id="82E78145FFAFFFCAFB8AFB7DFAC6CCAA" box="[1143,1333,1163,1189]" italics="true" pageId="32" pageNumber="33">Hebius modestus</emphasis>
|
||
</taxonomicName>
|
||
G̹nther.
|
||
<emphasis id="82E78145FFAFFFCAFF6AFB59FF04CCC6" bold="true" box="[151,247,1199,1225]" pageId="32" pageNumber="33">Squares</emphasis>
|
||
:
|
||
<taxonomicName id="779326D4FFAFFFCAFEF8FB59FD69CCC6" authority="Taylor. Black" authorityName="Taylor. Black" box="[261,666,1199,1225]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="deschauenseei">
|
||
<emphasis id="82E78145FFAFFFCAFEF8FB59FE0FCCC6" box="[261,508,1199,1225]" italics="true" pageId="32" pageNumber="33">Hebius deschauenseei</emphasis>
|
||
Taylor.
|
||
<emphasis id="82E78145FFAFFFCAFDABFB59FD69CCC6" bold="true" box="[598,666,1199,1225]" pageId="32" pageNumber="33">Black</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="82E78145FFAFFFCAFD5DFB59FD0ECCC6" bold="true" box="[672,765,1199,1225]" pageId="32" pageNumber="33">triangle</emphasis>
|
||
:
|
||
<typeStatus id="6F28E3F5FFAFFFCAFCF6FB46FCB0CCC5" box="[779,835,1200,1226]" pageId="32" pageNumber="33">Type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="779326D4FFAFFFCAFC3DFB59FBADCCC6" authority="David & Vogel & Nguyen & Orlov & Pauwels & Teynié & Ziegler, 2021" authorityName="David & Vogel & Nguyen & Orlov & Pauwels & Teynié & Ziegler" authorityYear="2021" box="[960,1118,1199,1225]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="igneus" status="sp. nov.">
|
||
<emphasis id="82E78145FFAFFFCAFC3DFB59FBADCCC6" box="[960,1118,1199,1225]" italics="true" pageId="32" pageNumber="33">Hebius igneus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="99D43C3EFFAFFFCAFB99FB59FB24CCC5" box="[1124,1239,1199,1226]" pageId="32" pageNumber="33" rank="species">
|
||
<emphasis id="82E78145FFAFFFCAFB99FB59FB22CCC6" bold="true" box="[1124,1233,1199,1225]" pageId="32" pageNumber="33">spec. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="82E78145FFAFFFCAFB20FB59FA67CCC6" bold="true" box="[1245,1428,1199,1225]" pageId="32" pageNumber="33">White triangles</emphasis>
|
||
:
|
||
<taxonomicName id="779326D4FFAFFFCAFF6AFB25FEC4CCE2" authority="David & Vogel & Nguyen & Orlov & Pauwels & Teynié & Ziegler, 2021" authorityName="David & Vogel & Nguyen & Orlov & Pauwels & Teynié & Ziegler" authorityYear="2021" box="[151,311,1235,1261]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="igneus" status="sp. nov.">
|
||
<emphasis id="82E78145FFAFFFCAFF6AFB25FEC4CCE2" box="[151,311,1235,1261]" italics="true" pageId="32" pageNumber="33">Hebius igneus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="99D43C3EFFAFFFCAFEC3FB25FE41CCE1" box="[318,434,1235,1262]" pageId="32" pageNumber="33" rank="species">
|
||
<emphasis id="82E78145FFAFFFCAFEC3FB25FE58CCE2" bold="true" box="[318,427,1235,1261]" pageId="32" pageNumber="33">spec. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B02C5D57FFAFFFCAFF3AFAEBFB66CE51" blockId="32.[151,1437,1309,2034]" pageId="32" pageNumber="33">
|
||
<taxonomicName id="779326D4FFAFFFCAFF3AFAEBFE52CD38" box="[199,417,1309,1335]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FFAFFFCAFF3AFAEBFE52CD38" box="[199,417,1309,1335]" italics="true" pageId="32" pageNumber="33">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="779326D4FFAFFFCAFDC9FAE8FD45CD38" authorityName="Wall" authorityYear="1910" box="[564,694,1310,1335]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="venningi">
|
||
<emphasis id="82E78145FFAFFFCAFDC9FAE8FD45CD38" box="[564,694,1310,1335]" italics="true" pageId="32" pageNumber="33">H. venningi</emphasis>
|
||
</taxonomicName>
|
||
especially by (1) a prefrontal usually single vs. always double; (2) the number of postoculars, 2 (rarely 3) in
|
||
<taxonomicName id="779326D4FFAFFFCAFD8CFAB4FCEBCD54" box="[625,792,1346,1371]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FFAFFFCAFD8CFAB4FCEBCD54" box="[625,792,1346,1371]" italics="true" pageId="32" pageNumber="33">H. annamensis</emphasis>
|
||
</taxonomicName>
|
||
vs. usually 3 (rarely 2) in
|
||
<taxonomicName id="779326D4FFAFFFCAFBBBFAB4FB3ACD54" authorityName="Wall" authorityYear="1910" box="[1094,1225,1346,1371]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="venningi">
|
||
<emphasis id="82E78145FFAFFFCAFBBBFAB4FB3ACD54" box="[1094,1225,1346,1371]" italics="true" pageId="32" pageNumber="33">H. venningi</emphasis>
|
||
</taxonomicName>
|
||
; (3) venter largely pale, with only tips of ventral scales dark vs. venter pale mesially, at least on the anterior part of the body, clouded with darker hues of brown on the outer parts of ventrals in
|
||
<taxonomicName id="779326D4FFAFFFCAFCC1FA7AFC4CCDAA" authorityName="Wall" authorityYear="1910" box="[828,959,1420,1445]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="venningi">
|
||
<emphasis id="82E78145FFAFFFCAFCC1FA7AFC4CCDAA" box="[828,959,1420,1445]" italics="true" pageId="32" pageNumber="33">H. venningi</emphasis>
|
||
</taxonomicName>
|
||
; (4) dorsal pattern made of isolate, bright orange or rusty-red blotches, usually horizontally elongate or vertically elongate as crossbars anteriorly vs. dorsal surface distinctly chequered by the presence on sides and upper part of the body of diffuse blackish-brown or very dark grey blotches, and with a dorsolateral series of irregular blotches, yellow-ochre or yellowish-brown (bright yellowish-ochre in life), enlarged and forming a chain on the first quarter to third of the body; (5) the distribution range,
|
||
<collectingCountry id="C8841DC7FFAFFFCAFF1EF9B5FEB1CE52" box="[227,322,1603,1629]" name="Vietnam" pageId="32" pageNumber="33">Vietnam</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="C8841DC7FFAFFFCAFE22F9B2FDE6CE51" box="[479,533,1604,1630]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
|
||
vs. Northeast
|
||
<collectingCountry id="C8841DC7FFAFFFCAFD45F9B5FD01CE52" box="[696,754,1603,1629]" name="India" pageId="32" pageNumber="33">India</collectingCountry>
|
||
and North
|
||
<collectingCountry id="C8841DC7FFAFFFCAFC8DF9B2FC2ECE51" box="[880,989,1604,1630]" name="Myanmar" pageId="32" pageNumber="33">Myanmar</collectingCountry>
|
||
for
|
||
<taxonomicName id="779326D4FFAFFFCAFBF6F9B2FB7DCE52" authorityName="Wall" authorityYear="1910" box="[1035,1166,1604,1629]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="33" phylum="Chordata" rank="species" species="venningi">
|
||
<emphasis id="82E78145FFAFFFCAFBF6F9B2FB7DCE52" box="[1035,1166,1604,1629]" italics="true" pageId="32" pageNumber="33">H. venningi</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8890EDCFFAFFFC8FF3AF99EFAA8CBE2" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33" type="description">
|
||
<paragraph id="B02C5D57FFAFFFCAFF3AF99EFDABCF35" blockId="32.[151,1437,1309,2034]" pageId="32" pageNumber="33">
|
||
<emphasis id="82E78145FFAFFFCAFF3AF99EFEA2CE8D" bold="true" box="[199,337,1640,1666]" pageId="32" pageNumber="33">Description</emphasis>
|
||
.—Body moderately robust but elongate, somewhat thinner in males, and cylindrical; head oval, rather short, moderately distinct from the thick neck, flattened anteriorly; snout long, flat, blunt or slightly round seen from above, subrectangular seen in profile, 2.0–2.2 times as long as horizontal diameter of eye; no canthus rostralis; nostril directed dorsolaterally, small, round, piercing in the middle of the divided nasal; eye moderately sized or small, its diameter about 1.2–1.4 times the distance between the lower margins of eye and of lip, with a round pupil; tail long, thin and tapering.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAFFFCAFF3AF8B3FB60CF8B" blockId="32.[151,1437,1309,2034]" pageId="32" pageNumber="33">
|
||
The maximum known length is
|
||
<quantity id="776BF0B2FFAFFFCAFDCDF8B3FD7DCF50" box="[560,654,1861,1887]" metricMagnitude="-1" metricUnit="m" metricValue="7.959999999999999" pageId="32" pageNumber="33" unit="mm" value="796.0">796 mm</quantity>
|
||
for a male (SVL
|
||
<quantity id="776BF0B2FFAFFFCAFCA8F8B3FC46CF50" box="[853,949,1861,1888]" metricMagnitude="-1" metricUnit="m" metricValue="5.279999999999999" pageId="32" pageNumber="33" unit="mm" value="528.0">528 mm</quantity>
|
||
, TaL
|
||
<quantity id="776BF0B2FFAFFFCAFC09F8B3FBA6CF50" box="[1012,1109,1861,1887]" metricMagnitude="-1" metricUnit="m" metricValue="2.68" pageId="32" pageNumber="33" unit="mm" value="268.0">268 mm</quantity>
|
||
; specimen ZISP NLO 3257). The longest known female is
|
||
<quantity id="776BF0B2FFAFFFCAFE1FF89FFDCCCF8B" box="[482,575,1897,1924]" metricMagnitude="-1" metricUnit="m" metricValue="6.85" pageId="32" pageNumber="33" unit="mm" value="685.0">685 mm</quantity>
|
||
long (SVL
|
||
<quantity id="776BF0B2FFAFFFCAFD3FF89FFCD2CF8B" box="[706,801,1897,1924]" metricMagnitude="-1" metricUnit="m" metricValue="4.65" pageId="32" pageNumber="33" unit="mm" value="465.0">465 mm</quantity>
|
||
, TaL
|
||
<quantity id="776BF0B2FFAFFFCAFCA2F89CFC4CCF8B" box="[863,959,1898,1924]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="32" pageNumber="33" unit="mm" value="220.0">220 mm</quantity>
|
||
; ZISP NLO 2831).
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAFFFCAFF3AF878FA6ACFA6" blockId="32.[151,1437,1309,2034]" box="[199,1433,1934,1961]" pageId="32" pageNumber="33">
|
||
Ratio TaL/TL
|
||
<geoCoordinate id="D5A73B90FFAFFFCAFE9AF879FE54CFA6" box="[359,423,1935,1961]" degrees="0.297" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="55" value="0.297">0.297</geoCoordinate>
|
||
<geoCoordinate id="D5A73B90FFAFFFCAFE5AF879FE05CFA6" box="[423,502,1935,1961]" degrees="0.337" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-0.337">–0.337</geoCoordinate>
|
||
, without clear sexual dimorphism (
|
||
<specimenCount id="A69596DEFFAFFFCAFC7DF878FC2FCFA7" box="[896,988,1934,1960]" pageId="32" pageNumber="33" type="male">6 males</specimenCount>
|
||
:
|
||
<geoCoordinate id="D5A73B90FFAFFFCAFC1BF879FBD6CFA6" box="[998,1061,1935,1961]" degrees="0.301" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="55" value="0.301">0.301</geoCoordinate>
|
||
<geoCoordinate id="D5A73B90FFAFFFCAFBDBF879FB85CFA6" box="[1062,1142,1935,1961]" degrees="0.337" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-0.337">–0.337</geoCoordinate>
|
||
;
|
||
<specimenCount id="A69596DEFFAFFFCAFB7CF879FB01CFA7" box="[1153,1266,1934,1961]" pageId="32" pageNumber="33" type="female">3 females</specimenCount>
|
||
:
|
||
<geoCoordinate id="D5A73B90FFAFFFCAFB01F879FACDCFA6" box="[1276,1342,1935,1961]" degrees="0.297" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="55" value="0.297">0.297</geoCoordinate>
|
||
<geoCoordinate id="D5A73B90FFAFFFCAFAC3F879FA7ECFA6" box="[1342,1421,1935,1961]" degrees="0.321" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="55" value="-0.321">–0.321</geoCoordinate>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAFFFCAFF3AF842FE41CFFD" blockId="32.[151,1437,1309,2034]" pageId="32" pageNumber="33">
|
||
<emphasis id="82E78145FFAFFFCAFF3AF842FEC2CFC2" box="[199,305,1972,1997]" italics="true" pageId="32" pageNumber="33">Dentition</emphasis>
|
||
. 28–34 maxillary teeth, gradually enlarging in a continuous series, the last 2 teeth moderately enlarged, without diastema.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFF61FB95C8F5" blockId="33.[151,1437,150,2038]" pageId="33" pageNumber="34">
|
||
<emphasis id="82E78145FFAEFFCBFF3AFF61FE81C8BE" box="[199,370,151,177]" italics="true" pageId="33" pageNumber="34">Body scalation</emphasis>
|
||
. DSR: 17–19-15–17-15–17 rows; scales rhomboedric, normal in shape, distinctly keeled on 2
|
||
<superScript id="47E6F01FFFAEFFCBFF58FF4DFF36C8D9" attach="none" box="[165,197,187,214]" fontSize="6" pageId="33" pageNumber="34">nd–</superScript>
|
||
10
|
||
<superScript id="47E6F01FFFAEFFCBFF1DFF4DFF1EC8C6" attach="none" box="[224,237,187,201]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
, moderately on the anterior part of the body, strongly posteriorly, very strongly in the region of the vent and along the base of the tail; scales on 1
|
||
<superScript id="47E6F01FFFAEFFCBFDCEFF16FDCDC8E1" attach="left" box="[563,574,224,238]" fontSize="6" pageId="33" pageNumber="34">st</superScript>
|
||
row smooth anteriorly, feebly keeled posteriorly.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFEF3FEC5C966" blockId="33.[151,1437,150,2038]" pageId="33" pageNumber="34">
|
||
The number of dorsal scale rows is quite variable in this species. In our sample of
|
||
<specimenCount id="A69596DEFFAEFFCBFB7EFEF3FAD3C910" box="[1155,1312,261,288]" pageId="33" pageNumber="34" type="generic">15 specimens</specimenCount>
|
||
, the dorsal scale row formulas are: 18–15–15 (
|
||
<specimenCount id="A69596DEFFAEFFCBFDCFFEDCFD46C94A" box="[562,693,298,325]" pageId="33" pageNumber="34" type="generic">1 specimen</specimenCount>
|
||
), 19–15–15 (4), 17–17–17 (1), 18–17–15 (2), 18–17–17 (2), and 19–17–17 (5).
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFE82FD8CC9BD" blockId="33.[151,1437,150,2038]" pageId="33" pageNumber="34">VEN: 158–172 (plus 1 or 2 preventrals), with a sexual dimorphism (see below); SC: 116–146, all paired, with a sexual dimorphism; cloacal plate divided.</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFE4BFE1FC9D8" blockId="33.[151,1437,150,2038]" box="[199,492,445,471]" pageId="33" pageNumber="34">Ratio VEN/SC 1.18–1.44.</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFE14FEFCCA4A" blockId="33.[151,1437,150,2038]" pageId="33" pageNumber="34">Position of the reduction to 6 scale rows around the tail: SC 4–16, with a strong sexual dimorphism (see below); ratio: length of the portion of tail with 6 dorsal scale rows/length of the portion of tail with 4 dorsal scale rows: 0.10–0.64.</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFDA6FEA4CD46" blockId="33.[151,1437,150,2038]" pageId="33" pageNumber="34">
|
||
<emphasis id="82E78145FFAEFFCBFF3AFDA6FE81CA65" box="[199,370,592,618]" italics="true" pageId="33" pageNumber="34">Head scalation</emphasis>
|
||
. Arrangement of upper head scales complete but prefrontals modified, including 2 internasals, 1 or 2 prefrontals (see below), 2 supraoculars, 1 frontal, and 2 parietals. Rostral hexagonal, wider than high, barely visible from above; nasal subrectangular, elongate, vertically divided by a furrow, with the fore and hind parts of similar size, altogether about 1.3–1.5 times longer than high; internasals distinctly subtriangular, elongate, in broad contact with each other, 1.2–1.4 times longer than wide, distinctly narrowing anteriorly with anterior margin about 0.4–0.5 times the width of the posterior margin; 1 prefrontal (in 12/
|
||
<specimenCount id="A69596DEFFAEFFCBFC6CFCFEFBDFCB2C" box="[913,1068,776,803]" pageId="33" pageNumber="34" type="generic">15 specimens</specimenCount>
|
||
), shield-like and broad, or 2 prefrontals (3/15), subrectangular, in both cases broader than long and 0.9–1.2 times as long as internasals, in broad contact with loreal; frontal hexagonal, shield-like, rather wide but short, its apex directed posteriorly, 1.0–1.2 times longer than wide, 1.7–2.0 times longer than prefrontals; on each side 1 entire, subtriangular supraocular, rather broad, 1.8–2.2 times longer than wide, slightly narrower than internasals, about 0.5 times as wide as frontal; parietals large, long and wide, 1.7–2.1 times longer than frontal and in contact for a length 1.2 times as long as the frontal length; large, rectangular loreal scale, elongate, 1.5–1.8 times longer than high, in broad contact with the nasal; 2 subequal preoculars on each side in all examined specimens; 2 or rarely 3 small postoculars, the upper one largest; 8 or 9 supralabials, 1
|
||
<superScript id="47E6F01FFFAEFFCBFE7FFBD9FE7ECC32" attach="left" box="[386,397,1071,1085]" fontSize="6" pageId="33" pageNumber="34">st</superScript>
|
||
as long as high, 2
|
||
<superScript id="47E6F01FFFAEFFCBFDAAFBD8FD85CC46" attach="none" box="[599,630,1070,1097]" fontSize="6" pageId="33" pageNumber="34">nd–</superScript>
|
||
5
|
||
<superScript id="47E6F01FFFAEFFCBFD7EFBD8FD63CC33" attach="left" box="[643,656,1070,1084]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
longer than high, other supralabials as long as high, 1
|
||
<superScript id="47E6F01FFFAEFFCBFB13FBD9FAFBCC46" attach="none" box="[1262,1288,1071,1097]" fontSize="6" pageId="33" pageNumber="34">st–</superScript>
|
||
2
|
||
<superScript id="47E6F01FFFAEFFCBFAE8FBD8FAD6CC33" attach="left" box="[1301,1317,1070,1084]" fontSize="6" pageId="33" pageNumber="34">nd</superScript>
|
||
supralabials, small and short or, exceptionally 1
|
||
<superScript id="47E6F01FFFAEFFCBFDB9FBA2FDACCC61" attach="none" box="[580,607,1108,1134]" fontSize="6" pageId="33" pageNumber="34">st–</superScript>
|
||
3
|
||
<superScript id="47E6F01FFFAEFFCBFD96FBA5FD8ACC6E" attach="left" box="[619,633,1107,1121]" fontSize="6" pageId="33" pageNumber="34">rd</superScript>
|
||
supralabials, in contact with nasal, 2
|
||
<superScript id="47E6F01FFFAEFFCBFBEEFBA5FBC0CC61" attach="none" box="[1043,1075,1107,1134]" fontSize="6" pageId="33" pageNumber="34">nd–</superScript>
|
||
3
|
||
<superScript id="47E6F01FFFAEFFCBFBC2FBA5FBBECC6E" attach="left" box="[1087,1101,1107,1121]" fontSize="6" pageId="33" pageNumber="34">rd</superScript>
|
||
or 3
|
||
<superScript id="47E6F01FFFAEFFCBFB82FBA5FB6FCC61" attach="none" box="[1151,1180,1107,1134]" fontSize="6" pageId="33" pageNumber="34">rd–</superScript>
|
||
4
|
||
<superScript id="47E6F01FFFAEFFCBFB54FBA5FB45CC6E" attach="left" box="[1193,1206,1107,1121]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
supralabials in large contact with the loreal, 4
|
||
<superScript id="47E6F01FFFAEFFCBFE57FB8EFE35CC9C" attach="none" box="[426,454,1144,1171]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
5
|
||
<superScript id="47E6F01FFFAEFFCBFE2EFB8EFE13CC89" attach="left" box="[467,480,1144,1158]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
supralabials, or exceptionally solely 4
|
||
<superScript id="47E6F01FFFAEFFCBFC71FB8EFC6ACC89" attach="none" box="[908,921,1144,1158]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
, solely 5
|
||
<superScript id="47E6F01FFFAEFFCBFC00FB8EFBF9CC89" attach="none" box="[1021,1034,1144,1158]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
, 4
|
||
<superScript id="47E6F01FFFAEFFCBFBD8FB8EFBB2CC9D" attach="none" box="[1061,1089,1144,1170]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
6
|
||
<superScript id="47E6F01FFFAEFFCBFBB3FB8EFBA8CC89" attach="left" box="[1102,1115,1144,1158]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
or 5
|
||
<superScript id="47E6F01FFFAEFFCBFB70FB8EFB5BCC9D" attach="none" box="[1165,1192,1144,1170]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
6
|
||
<superScript id="47E6F01FFFAEFFCBFB48FB8EFB31CC89" attach="left" box="[1205,1218,1144,1158]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
supralabials, entering orbit; 6
|
||
<superScript id="47E6F01FFFAEFFCBFEE5FB6BFEC0CCB7" attach="none" box="[280,307,1181,1208]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
7
|
||
<superScript id="47E6F01FFFAEFFCBFEBDFB6BFEBECCA4" attach="left" box="[320,333,1181,1195]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
or 7
|
||
<superScript id="47E6F01FFFAEFFCBFE79FB6BFE52CCB8" attach="none" box="[388,417,1181,1207]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
8
|
||
<superScript id="47E6F01FFFAEFFCBFE50FB6BFE49CCA4" attach="left" box="[429,442,1181,1195]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
supralabials largest; 1 anterior temporal, narrow and elongate, in all known specimens with complete temporal formulas as 1+1+2 or rarely 1+2+2 or 1+3 (this latter condition only in the
|
||
<typeStatus id="6F28E3F5FFAEFFCBFB15FB34FAEECCD3" box="[1256,1309,1218,1244]" pageId="33" pageNumber="34">type</typeStatus>
|
||
); 10 (rarely 8 or 9) infralabials, 1
|
||
<superScript id="47E6F01FFFAEFFCBFE79FB11FE7CCCFA" attach="left" box="[388,399,1255,1269]" fontSize="6" pageId="33" pageNumber="34">st</superScript>
|
||
pair in contact with each other behind the mental, 1
|
||
<superScript id="47E6F01FFFAEFFCBFC2AFB11FC02CD0E" attach="none" box="[983,1009,1254,1281]" fontSize="6" pageId="33" pageNumber="34">st–</superScript>
|
||
5
|
||
<superScript id="47E6F01FFFAEFFCBFC03FB10FBF8CCFB" attach="left" box="[1022,1035,1254,1268]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
infralabials in contact with anterior chin shields, 5
|
||
<superScript id="47E6F01FFFAEFFCBFEC6FAFDFEBBCD16" attach="left" box="[315,328,1291,1305]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
infralabial largest; mental triangular; anterior pair of chin shields equal to or slightly longer than posterior shields.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFCBFF3AFAA5FCD6CF12" blockId="33.[151,1437,150,2038]" pageId="33" pageNumber="34">
|
||
<emphasis id="82E78145FFAEFFCBFF3AFAA5FE3ECD62" box="[199,461,1363,1389]" italics="true" pageId="33" pageNumber="34">Coloration and pattern</emphasis>
|
||
. In preservative, the dorsal and lateral surfaces of body are dark chestnut brown, dark brown or dark greyish-brown, darker above than on the sides, either nearly uniform or, more usually, profusely variegated with diffuse, darker brown blotches, larger on the upper parts of the sides than on the vertebral region; scales of the lower lateral scale rows usually spotted or marked with diffuse, yellowish-grey blotches or streaks; in 3 out of
|
||
<specimenCount id="A69596DEFFAEFFCBFF4BFA12FEA2CDF2" box="[182,337,1507,1534]" pageId="33" pageNumber="34" type="generic">13 specimens</specimenCount>
|
||
, a faint dorsolateral stripe is present on 5
|
||
<superScript id="47E6F01FFFAEFFCBFCE3FA15FCCACDF1" attach="none" box="[798,825,1507,1534]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
7
|
||
<superScript id="47E6F01FFFAEFFCBFCBBFA15FCA0CDFE" attach="left" box="[838,851,1507,1521]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
dorsal rows, barely paler than the background color and poorly distinct; on each side, a series of about 50 to 70 distinct dorsolateral blotches, cream, ochre, yellowishbrown, pinkish-brown or orange-brown on 5
|
||
<superScript id="47E6F01FFFAEFFCBFD79F9DDFD53CE49" attach="none" box="[644,672,1579,1606]" fontSize="6" pageId="33" pageNumber="34">th–</superScript>
|
||
7
|
||
<superScript id="47E6F01FFFAEFFCBFD50F9DDFD49CE36" attach="left" box="[685,698,1579,1593]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
dorsal rows, separated from each other by 2–3 dorsal scales, forming a discontinuous dorsolateral stripe; the first few blotches are usually vertically elongate and form a few vertical crossbars extending outwards and reaching downwards the level of the 3
|
||
<superScript id="47E6F01FFFAEFFCBFC3EF985FC22CE8E" attach="left" box="[963,977,1651,1665]" fontSize="6" pageId="33" pageNumber="34">rd</superScript>
|
||
scale row on the anteriormost part of the body, quickly followed by thick and horizontally elongate blotches extending up to the first anterior quarter to third of the body; progressively these blotches are reduced to irregular or broken spots or even disappear posteriorly; a short, cream ventrolateral stripe on the anterior part of the body, up to 20
|
||
<superScript id="47E6F01FFFAEFFCBFC39F929FC22CEE2" attach="left" box="[964,977,1759,1773]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
to 30
|
||
<superScript id="47E6F01FFFAEFFCBFBEDF929FBEECEE2" attach="left" box="[1040,1053,1759,1773]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
ventrals. The upper surface of tail is as the body, either uniform or with only a few pale dots.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFAEFFC8FF3AF8D1FF37C8F6" blockId="33.[151,1437,150,2038]" lastBlockId="34.[151,1437,151,1617]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
|
||
The head is dark chestnut-brown, more or less variegated above with ochre yellow or pale yellowish-brown, often with numerous minute scattered dark dots; a ochre yellow or pale yellowish-brown sagittal line present, but faint, or absent; last three posterior supralabials irregularly and faintly edged with ochre-yellow anteriorly; a short, narrow or relatively broad, oblique, more or less distinct postocular streak on anterior and lower posterior temporals, and on top of 8
|
||
<superScript id="47E6F01FFFAEFFCBFEBAF841FEA7CFCA" attach="left" box="[327,340,1975,1989]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
and 9
|
||
<superScript id="47E6F01FFFAEFFCBFE66F841FE5BCFCA" attach="left" box="[411,424,1975,1989]" fontSize="6" pageId="33" pageNumber="34">th</superScript>
|
||
supralabials, reaching the neck behind the corner of the mouth and extending as a large, cream or yellowish-ochre oblique, elongate blotch on the nape, connecting the posterior end of the postocular streak with the first dorsolateral blotch. The throat, chin and infralabials are cream or pale yellowish-ochre, more or less heavily spotted with dark brown spots; infralabials heavily marbled or spotted with dark brown. Eye deep black in life.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFEF5FB86C986" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">The venter is cream, pale yellowish-ochre or pale brown on the inner half to two thirds of its width, with tips of ventrals dark brown or dark greyish-brown, this dark surface progressively widening posteriorly; often some irregular dark spots scattered on the pale part. The under surface of tail is as the venter but the pale central part is narrower than the dark, outer parts anteriorly, the tail becoming progressively entirely dark below.</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFE65FE6ACAC2" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">In life, the background coloration is chestnut-brown, dark olive-brown or dark brown, more or less iridescent; the upper dorsal dark blotches are blackish-brown or very dark grey whereas the lower lateral scale rows are marked with yellowish-ochre blotches or streaks; when present, the dorsal stripe is rusty-brown; the main dorsolateral blotches are yellowish-ochre, orange or reddish-brown, a colour often still seen in recently preserved specimens; the short ventrolateral stripe is pale yellowish-ochre. Pale markings on the head are yellowish-ochre or dirty, greyish-yellow; the elongate blotch on the temporal region and on the hinder part of the head is yellowish-ochre or sometimes reddish-ochre; behind the head, the streak obliquely directed upwards and backwards on the neck is of the same colour or turns to bright orange or yellowish-red. The background colour of the throat and venter is cream or pale yellowish-grey.</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFD20FEE9CB1A" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">
|
||
<emphasis id="82E78145FFADFFC8FF3AFD20FEB8CAFF" bold="true" box="[199,331,726,752]" pageId="34" pageNumber="35">Hemipenes</emphasis>
|
||
.—In situ, the organ is single, short and subcylindrical, not forked, reaching the 10
|
||
<superScript id="47E6F01FFFADFFC8FAFDFD20FAFECAEB" attach="left" box="[1280,1293,726,740]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
|
||
SC, spinose throughout.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFCE8FCA3CB36" blockId="34.[151,1437,151,1617]" box="[199,848,798,825]" pageId="34" pageNumber="35">
|
||
<emphasis id="82E78145FFADFFC8FF3AFCE8FE43CB37" bold="true" box="[199,432,798,824]" pageId="34" pageNumber="35">Sexual dimorphism</emphasis>
|
||
.
|
||
<emphasis id="82E78145FFADFFC8FE4AFCE9FE20CB36" bold="true" box="[439,467,799,825]" pageId="34" pageNumber="35">—</emphasis>
|
||
It is expressed in three characters:
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFCB5FE4ECB8E" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">(1) by the difference in the number of ventrals: males: 168–172 (mean = 167.7, s = 1.7); females: 158–167 (mean = 164.2, SD = 3.4).</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFC7DFE4FCBC6" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">(2) by the difference in the number of subcaudals: males: 126–146 (mean = 138.7, s = 6.9); females: 116–126 (mean = 119.7, SD = 5.5).</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFC25FAA8CBE2" blockId="34.[151,1437,151,1617]" box="[199,1371,978,1005]" pageId="34" pageNumber="35">
|
||
(3) by the position of the reduction to 6 scale rows around the tail: males: 12–16
|
||
<superScript id="47E6F01FFFADFFC8FBB4FC24FBA5CBEF" attach="left" box="[1097,1110,978,992]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
|
||
SC; females: 4–7
|
||
<superScript id="47E6F01FFFADFFC8FAE3FC24FAD8CBEF" attach="left" box="[1310,1323,978,992]" fontSize="6" pageId="34" pageNumber="35">th</superScript>
|
||
SC.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8890EDCFFADFFC8FF3AFC00FA9FCCCA" pageId="34" pageNumber="35" type="distribution">
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFC00FA9FCCCA" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">
|
||
<emphasis id="82E78145FFADFFC8FF3AFC00FE47CC1F" bold="true" box="[199,436,1014,1040]" pageId="34" pageNumber="35">
|
||
Distribution (
|
||
<figureCitation id="28A841D2FFADFFC8FE94FC00FE58CC1F" box="[361,427,1014,1040]" captionStart="FIGURE 3" captionStartId="12.[151,260,1218,1244]" captionTargetBox="[226,1360,246,1184]" captionTargetId="figure-55@12.[219,1368,234,1194]" captionTargetPageId="12" captionText="FIGURE 3. Distribution map of species with 15 or 17 dorsal scale rows at midbody. Circles: Hebius annamensis Bourret. Diamonds: Hebius venningi Wall. Inverted triangles: Hebius taronensis Smith. Squares: Hebius nigriventer Wall. Triangles: Hebius chapaensis Bourret." figureDoi="http://doi.org/10.5281/zenodo.4463746" httpUri="https://zenodo.org/record/4463746/files/figure.png" pageId="34" pageNumber="35">Fig. 3</figureCitation>
|
||
)
|
||
</emphasis>
|
||
.—This species has been recorded only from central
|
||
<collectingCountry id="C8841DC7FFADFFC8FC04FC01FBABCC1E" box="[1017,1112,1015,1041]" name="Vietnam" pageId="34" pageNumber="35">Vietnam</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="C8841DC7FFADFFC8FB0FFC01FADFCC1E" box="[1266,1324,1015,1041]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
|
||
.
|
||
<emphasis id="82E78145FFADFFC8FACDFC00FA65CC1F" bold="true" box="[1328,1430,1014,1040]" pageId="34" pageNumber="35">
|
||
<collectingCountry id="C8841DC7FFADFFC8FACDFC00FA65CC1F" box="[1328,1430,1014,1040]" name="Vietnam" pageId="34" pageNumber="35">Vietnam</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<collectingRegion id="725793B5FFADFFC8FF6AFBEDFF08CC3A" box="[151,251,1051,1077]" country="Vietnam" name="Da Nang" pageId="34" pageNumber="35">Da Nang</collectingRegion>
|
||
City. Ba Na Nature Reserve.
|
||
<collectingRegion id="725793B5FFADFFC8FDC2FBEDFCF5CC3A" box="[575,774,1051,1077]" country="Vietnam" name="Ha Tinh" pageId="34" pageNumber="35">Ha Tinh Province</collectingRegion>
|
||
. Huong Son District.
|
||
<collectingRegion id="725793B5FFADFFC8FC0EFBEDFB39CC3A" box="[1011,1226,1051,1077]" country="Vietnam" name="Kon Tum" pageId="34" pageNumber="35">Kon Tum Province</collectingRegion>
|
||
. Ngoc Linh Nature Reserve, Kon Plong District.
|
||
<collectingRegion id="725793B5FFADFFC8FE15FBC9FD11CC56" box="[488,738,1087,1113]" country="Vietnam" name="Quang Binh" pageId="34" pageNumber="35">Quang Binh Province</collectingRegion>
|
||
. Dong Chau Forest, Lê Thuy District; Phong Nha - Ke Bang National Park, Minh Hoa District.
|
||
<collectingRegion id="725793B5FFADFFC8FDDFFB95FCF6CC72" box="[546,773,1123,1149]" country="Vietnam" name="Quang Tri" pageId="34" pageNumber="35">Quang Tri Province</collectingRegion>
|
||
. Huong Hoa District.
|
||
<collectingRegion id="725793B5FFADFFC8FC02FB95FADACC72" box="[1023,1321,1123,1149]" country="Vietnam" name="Thua Thien-Hue" pageId="34" pageNumber="35">Thua Thien-Hue Province</collectingRegion>
|
||
. Bach Ma National Park—
|
||
<emphasis id="82E78145FFADFFC8FEACFB71FE78CCAE" bold="true" box="[337,395,1159,1185]" pageId="34" pageNumber="35">
|
||
<collectingCountry id="C8841DC7FFADFFC8FEACFB71FE78CCAE" box="[337,395,1159,1185]" name="Laos" pageId="34" pageNumber="35">Laos</collectingCountry>
|
||
</emphasis>
|
||
.
|
||
<collectingRegion id="725793B5FFADFFC8FE61FB71FD63CCAE" box="[412,656,1159,1185]" country="Laos" name="Champasak" pageId="34" pageNumber="35">Champasak Province</collectingRegion>
|
||
. Dong Hua Sao National Biodiversity Conservation Area, Pakxong District.
|
||
<collectingRegion id="725793B5FFADFFC8FF04FB5DFE33CCCA" box="[249,448,1195,1221]" country="Laos" name="Xekong" pageId="34" pageNumber="35">Xékong Province</collectingRegion>
|
||
. Sap National Biodiversity Conservation Area, Kaleum District; Dakchung District.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F8890EDCFFADFFC8FF3AFB38FDB4CE5E" pageId="34" pageNumber="35" type="biology_ecology">
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFB38FB1CCD5A" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">
|
||
<emphasis id="82E78145FFADFFC8FF3AFB38FED2CCE7" bold="true" box="[199,289,1230,1256]" pageId="34" pageNumber="35">Biology</emphasis>
|
||
.—This species inhabits regions covered with tropical wet evergreen forest and subtropical montane evergreen forests between about 300 and
|
||
<quantity id="776BF0B2FFADFFC8FD81FB04FD28CD02" box="[636,731,1266,1293]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="34" pageNumber="35" unit="m" value="1500.0">1,500 m</quantity>
|
||
a.s.l., although most specimens for which elevation data are available were collected between 1,200 and
|
||
<quantity id="776BF0B2FFADFFC8FD75FAE1FD17CD3E" box="[648,740,1303,1329]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="34" pageNumber="35" unit="m" value="1400.0">1,400 m</quantity>
|
||
a.s.l. (see also
|
||
<bibRefCitation id="D40220A6FFADFFC8FC6DFAE1FB53CD3E" author="Stuart, B. L. & Nguyen, T. Q." box="[912,1184,1303,1329]" pageId="34" pageNumber="35" pagination="2040117" refId="ref51137" refString="Stuart, B. L. & Nguyen, T. Q. (2012 a) Parahelicops annamensis. The IUCN Red List of Threatened Species, 2012, e. T 192102 A 2040117." type="journal article" year="2012">Stuart & Nguyen 2012a</bibRefCitation>
|
||
). All were collected in primary montane evergreen forests and in close association with fast-running, rocky forest streams.
|
||
</paragraph>
|
||
<paragraph id="B02C5D57FFADFFC8FF3AFAA9FDB4CE5E" blockId="34.[151,1437,151,1617]" pageId="34" pageNumber="35">
|
||
This species seems to be mainly nocturnal and aquatic, although one specimen was found on the ground. According to
|
||
<bibRefCitation id="D40220A6FFADFFC8FEECFA75FE5BCD91" author="Stuart, B. L." box="[273,424,1411,1438]" pageId="34" pageNumber="35" pagination="167 - 171" refId="ref51006" refString="Stuart, B. L. (2006) A second specimen of Parahelicops annamensis Bourret, 1934 (Serpentes: Colubridae / Natricinae). Hamadryad, 30 (1 - 2), 167 - 171." type="journal article" year="2006">Stuart (2006)</bibRefCitation>
|
||
, specimen FMNH 258637 was collected at night (20.30 h) in steep terrain covered by wet evergreen forest between
|
||
<quantity id="776BF0B2FFADFFC8FE48FA50FDADCDCE" box="[437,606,1446,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.39" metricValueMax="1.5" metricValueMin="1.28" pageId="34" pageNumber="35" unit="m" value="1390.0" valueMax="1500.0" valueMin="1280.0">
|
||
<geoCoordinate id="D5A73B90FFADFFC8FE48FA50FE07CDCE" box="[437,500,1446,1473]" degrees="1,280" direction="north" orientation="latitude" pageId="34" pageNumber="35" precision="55" value="1.28">1,280</geoCoordinate>
|
||
<geoCoordinate id="D5A73B90FFADFFC8FE09FA50FDB1CDCE" box="[500,578,1446,1473]" degrees="1,500" direction="west" orientation="longitude" pageId="34" pageNumber="35" precision="55" value="-1.5">–1,500</geoCoordinate>
|
||
m
|
||
</quantity>
|
||
elevation. The snake was first observed on land
|
||
<quantity id="776BF0B2FFADFFC8FB84FA50FB4ECDCE" box="[1145,1213,1446,1473]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="35" unit="cm" value="50.0">50 cm</quantity>
|
||
from a small, swift, rocky stream, then it dove into the water and swam under a rock on the stream bottom. Specimen ZFMK 86457 was captured at night (20.00–21.00h) in the leaf litter of a forest stream at
|
||
<quantity id="776BF0B2FFADFFC8FC67FA18FC13CE06" box="[922,992,1518,1545]" metricMagnitude="2" metricUnit="m" metricValue="5.1" pageId="34" pageNumber="35" unit="m" value="510.0">510 m</quantity>
|
||
a.s.l., with only the snout tip protruding from the surface of the water (
|
||
<bibRefCitation id="D40220A6FFADFFC8FE15F9E5FD4CCE22" author="David, P. & Pauwels, O. S. G. & Nguyen, T. Q. & Vogel, G." box="[488,703,1554,1581]" pageId="34" pageNumber="35" pagination="203 - 217" refId="ref47072" refString="David, P., Pauwels, O. S. G., Nguyen, T. Q. & Vogel, G. (2015 b) On the taxonomic status of the Thai endemic freshwater snake Parahelicops boonsongi, with the erection of a new genus (Squamata: Natricidae). Zootaxa, 3948 (2), 203 - 217. https: // doi. org / 10.11646 / zootaxa. 3948.2.3" type="journal article" year="2015">
|
||
David
|
||
<emphasis id="82E78145FFADFFC8FDC8F9E5FD95CE22" box="[565,614,1555,1581]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
|
||
. 2015b
|
||
</bibRefCitation>
|
||
). Other biological data of
|
||
<taxonomicName id="779326D4FFADFFC8FC1FF9E5FB4FCE23" box="[994,1212,1555,1581]" class="Squamata" family="Colubridae" genus="Hebius" higherTaxonomySource="CoL" kingdom="Animalia" pageId="34" pageNumber="35" phylum="Chordata" rank="species" species="annamensis">
|
||
<emphasis id="82E78145FFADFFC8FC1FF9E5FB4FCE23" box="[994,1212,1555,1581]" italics="true" pageId="34" pageNumber="35">Hebius annamensis</emphasis>
|
||
</taxonomicName>
|
||
, such as its diet and reproductive habits, are still unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |