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<document id="A100E4427A7576E226E1FB6AAACF40C3" ID-DOI="10.3853/j.0067-1975.55.2003.1383" ID-ISSN="2201-4349" ID-Zenodo-Dep="10092797" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629577880548" checkinUser="felipe" docAuthor="Zhen, Yong Yi, Percival, Ian G. &amp; Webby, Barry D." docDate="2003" docId="37439A20CB00FF16FF5BFEC8FAEBFE09" docLanguage="en" docName="RecAustMus.55.2.169-220.pdf" docOrigin="Records of the Australian Museum 55 (2)" docSource="https://journals.australian.museum/zhen-et-al-2003-rec-aust-mus-552-169220/" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Scolopodus multicostatus Barnes &amp; Tuke 1970" docType="treatment" docVersion="5" lastPageNumber="212" masterDocId="CB7AE258CB2BFF3DFFC5FFC8FFB8FFDC" masterDocTitle="Early Ordovician Conodonts from Far Western New South Wales, Australia" masterLastPageNumber="220" masterPageNumber="169" pageNumber="212" updateTime="1699559098628" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="C3FB6BFF68D1401E6F9A7D0C45F9A148">Early Ordovician Conodonts from Far Western New South Wales, Australia</mods:title>
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<mods:namePart id="D81DD82011C77CD56D46BC817C2E5BE5">Zhen, Yong Yi</mods:namePart>
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<mods:namePart id="1F8BB43E5C3ECA4535AA3AED866BBFEA">Percival, Ian G.</mods:namePart>
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<mods:namePart id="783B0A20FA8C0C990B86B4F685CB2B7E">Webby, Barry D.</mods:namePart>
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<mods:typeOfResource id="E40A14AAA719746FCAC5A6C306D3FC49">text</mods:typeOfResource>
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<mods:title id="BA1D05CC696B203BB6CFA6CCC7347E31">Records of the Australian Museum</mods:title>
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<mods:date id="B4B0D0A83F6DF3BFCE2899E48EE43646">2003</mods:date>
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<mods:number id="4DDB721F598FF8FE858DB867F40AC0CC">2003-08-13</mods:number>
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<mods:number id="EC281DE23A03799A65917E1E658DFB56">55</mods:number>
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<mods:number id="4CA08A694A389E4665C3CEBB83DF7A1A">2</mods:number>
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<mods:url id="C5B46D61ECA23337171C6070DF11560C">https://journals.australian.museum/zhen-et-al-2003-rec-aust-mus-552-169220/</mods:url>
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<mods:classification id="793D505A0CA07E2B1539357EBCC7659F">journal article</mods:classification>
<mods:identifier id="E327528A03455B20914AB8F399D7FC72" type="DOI">10.3853/j.0067-1975.55.2003.1383</mods:identifier>
<mods:identifier id="DFDF1AF72932C1CF7CC6B8BE22E299BC" type="ISSN">2201-4349</mods:identifier>
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<treatment id="37439A20CB00FF16FF5BFEC8FAEBFE09" LSID="urn:lsid:plazi:treatment:37439A20CB00FF16FF5BFEC8FAEBFE09" httpUri="http://treatment.plazi.org/id/37439A20CB00FF16FF5BFEC8FAEBFE09" lastPageNumber="212" pageId="43" pageNumber="212">
<subSubSection id="F7F078BDCB00FF16FF5BFEC8FD7FFEC4" box="[158,711,256,280]" pageId="43" pageNumber="212" type="nomenclature">
<paragraph id="BF552B36CB00FF16FF5BFEC8FD7FFEC4" blockId="43.[158,711,256,280]" box="[158,711,256,280]" pageId="43" pageNumber="212">
<heading id="E41D9C5ACB00FF16FF5BFEC8FD7FFEC4" box="[158,711,256,280]" centered="true" fontSize="11" level="2" pageId="43" pageNumber="212" reason="2">
<taxonomicName id="78EA50B5CB00FF16FF5BFEC8FD7FFEC4" authority="Barnes &amp; Tuke, 1970" authorityName="Barnes &amp; Tuke" authorityYear="1970" box="[158,711,256,280]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FF5BFEC8FD7FFEC4" bold="true" box="[158,711,256,280]" pageId="43" pageNumber="212">
<emphasis id="8D9EF724CB00FF16FF5BFEC8FE7DFEC4" bold="true" box="[158,453,256,280]" italics="true" pageId="43" pageNumber="212">Scolopodus multicostatus</emphasis>
Barnes &amp; Tuke, 1970
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FEB4FEE5FE5FFE98" box="[369,487,301,324]" pageId="43" pageNumber="212" type="description">
<paragraph id="BF552B36CB00FF16FEB4FEE5FE5FFE98" blockId="43.[369,487,301,324]" box="[369,487,301,324]" pageId="43" pageNumber="212">
<figureCitation id="27D137B3CB00FF16FEB4FEE5FE5FFE98" box="[369,487,301,324]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26A–R</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FFBCFE92FEB4FE62" pageId="43" pageNumber="212" type="reference_group">
<paragraph id="BF552B36CB00FF16FFBCFE92FE33FE55" blockId="43.[121,749,345,500]" pageId="43" pageNumber="212">
<taxonomicName id="78EA50B5CB00FF16FFBCFE92FDCAFEB2" authority="Barnes &amp; Tuke, 1970: 92" authorityName="Barnes &amp; Tuke" authorityPageNumber="92" authorityYear="1970" box="[121,626,345,367]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FFBCFE92FECAFEB3" box="[121,370,346,367]" italics="true" pageId="43" pageNumber="212">Scolopodus multicostatus</emphasis>
Barnes &amp; Tuke, 1970: 92
</taxonomicName>
, pl. 18, figs. 5, 9, 15, 16, text-fig. 6D.
</paragraph>
<paragraph id="BF552B36CB00FF16FFBCFE47FEB4FE62" blockId="43.[121,749,345,500]" pageId="43" pageNumber="212">
<taxonomicName id="78EA50B5CB00FF16FFBCFE47FEC9FE78" authorityName="Barnes &amp; Tuke" authorityYear="1970" box="[121,369,399,420]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FFBCFE47FEC9FE78" box="[121,369,399,420]" italics="true" pageId="43" pageNumber="212">Scolopodus multicostatus</emphasis>
</taxonomicName>
.–Ethington &amp; Clark, 1982: 101, pl. 11, figs. 19, 20.
</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FFBCFE0CFF40FE28" pageId="43" pageNumber="212" type="description">
<paragraph id="BF552B36CB00FF16FFBCFE0CFD6CFE05" blockId="43.[121,749,345,500]" box="[121,724,452,473]" pageId="43" pageNumber="212">
<taxonomicName id="78EA50B5CB00FF16FFBCFE0CFEC9FE05" authorityName="Barnes &amp; Tuke" authorityYear="1970" box="[121,369,452,473]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FFBCFE0CFEC9FE05" box="[121,369,452,473]" italics="true" pageId="43" pageNumber="212">Scolopodus multicostatus</emphasis>
</taxonomicName>
.–Stait &amp; Druce, 1993: 310, figs. 13H–
</paragraph>
<paragraph id="BF552B36CB00FF16FF52FE17FF40FE28" blockId="43.[121,749,345,500]" box="[151,248,479,500]" pageId="43" pageNumber="212">I, 19F–J,L.</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FFBCFDC0FD03FDA4" pageId="43" pageNumber="212" type="materials_examined">
<paragraph id="BF552B36CB00FF16FFBCFDC0FD03FDA4" blockId="43.[121,749,520,632]" pageId="43" pageNumber="212">
<emphasis id="8D9EF724CB00FF16FFBCFDC0FF61FDFC" bold="true" box="[121,217,520,544]" pageId="43" pageNumber="212">Material</emphasis>
.
<materialsCitation id="0F82216BCB00FF16FF20FDC1FD75FDE1" location="Tabita Formation" municipality="Mount Arrowsmith" pageId="43" pageNumber="212" specimenCount="85">
<specimenCount id="A9ECE0BFCB00FF16FF20FDC1FEC9FDFC" box="[229,369,521,544]" pageId="43" pageNumber="212" type="generic">85 specimens</specimenCount>
(8 Pa, 14 Pb, 17 Sa, 17 Sb, 17 Sc, 12 Sd) from the
<location id="BA357DEDCB00FF16FEC4FDEEFE0CFDE1" LSID="urn:lsid:plazi:treatment:37439A20CB00FF16FF5BFEC8FAEBFE09:BA357DEDCB00FF16FEC4FDEEFE0CFDE1" box="[257,436,550,573]" municipality="Mount Arrowsmith" name="Tabita Formation" pageId="43" pageNumber="212">Tabita Formation</location>
at
<collectingMunicipality id="5F31B14CCB00FF16FE16FDEEFD25FDE1" box="[467,669,550,573]" pageId="43" pageNumber="212">Mount Arrowsmith</collectingMunicipality>
, and
</materialsCitation>
<materialsCitation id="0F82216BCB00FF16FD17FDEEFD00FDA4" location="Koonenberry Gap" pageId="43" pageNumber="212" specimenCount="48">
<specimenCount id="A9ECE0BFCB00FF16FD17FDEEFF5DFD86" pageId="43" pageNumber="212" type="generic">48 specimens</specimenCount>
(3 Pa, 7 Pb, 6 Sa, 15 Sb, 12 Sc, 5 Sd) from the unnamed dolomitic limestone unit at
<location id="BA357DEDCB00FF16FE3FFDA9FD00FDA4" LSID="urn:lsid:plazi:treatment:37439A20CB00FF16FF5BFEC8FAEBFE09:BA357DEDCB00FF16FE3FFDA9FD00FDA4" box="[506,696,609,632]" name="Koonenberry Gap" pageId="43" pageNumber="212">Koonenberry Gap</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FFBCFD45FE87FC88" pageId="43" pageNumber="212" type="diagnosis">
<paragraph id="BF552B36CB00FF16FFBCFD45FE87FC88" blockId="43.[121,749,653,852]" pageId="43" pageNumber="212">
<emphasis id="8D9EF724CB00FF16FFBCFD45FF48FD79" bold="true" box="[121,240,653,677]" pageId="43" pageNumber="212">Diagnosis</emphasis>
. A species of
<taxonomicName id="78EA50B5CB00FF16FE63FD45FD94FD78" authorityName="Pander" authorityYear="1856" box="[422,556,653,676]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="genus">
<emphasis id="8D9EF724CB00FF16FE63FD45FD94FD78" box="[422,556,653,676]" italics="true" pageId="43" pageNumber="212">Scolopodus</emphasis>
</taxonomicName>
consisting of a seximembrate apparatus including two laterally compressed scandodiform
<taxonomicName id="78EA50B5CB00FF16FED5FD00FE39FD03" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[272,385,712,735]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">P elements</taxonomicName>
with a short base, and multicostate
<taxonomicName id="78EA50B5CB00FF16FFBCFD2DFF56FD20" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[121,238,741,764]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">S elements</taxonomicName>
with a longer base and a broad anterior margin; symmetrical Sa and
<taxonomicName id="78EA50B5CB00FF16FE9FFCCAFE66FCC5" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[346,478,770,793]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">Sd elements</taxonomicName>
more or less rounded in cross section, asymmetrical Sb and
<taxonomicName id="78EA50B5CB00FF16FDC2FCE8FD33FCEB" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[519,651,800,823]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">Sc elements</taxonomicName>
laterally more compressed.
</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FFBCFCADFDA3F949" pageId="43" pageNumber="212" type="description">
<paragraph id="BF552B36CB00FF16FFBCFCADFDA3F949" blockId="43.[121,749,869,1685]" pageId="43" pageNumber="212">
<emphasis id="8D9EF724CB00FF16FFBCFCADFF46FCA1" bold="true" box="[121,254,869,893]" pageId="43" pageNumber="212">Description</emphasis>
.
<taxonomicName id="78EA50B5CB00FF16FECFFCAEFEC6FCA1" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[266,382,870,893]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">P elements</taxonomicName>
scandodiform with erect cusp;
<taxonomicName id="78EA50B5CB00FF16FD0AFCAEFF6AFC46" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">
<emphasis id="8D9EF724CB00FF16FD0AFCAEFF6AFC46" italics="true" pageId="43" pageNumber="212">Pa element</emphasis>
</taxonomicName>
with a smooth outer lateral face; inner face multicostate with a median costa, an anterolateral costa, a posterior costa and a number of interposed weaker costae (
<figureCitation id="27D137B3CB00FF16FF44FC13FF43FC2E" box="[129,251,987,1010]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26A,B</figureCitation>
).
<emphasis id="8D9EF724CB00FF16FED4FC13FE3EFC2E" box="[273,390,987,1010]" italics="true" pageId="43" pageNumber="212">
Pb
<taxonomicName id="78EA50B5CB00FF16FEF6FC13FE3EFC2E" box="[307,390,987,1010]" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">element</taxonomicName>
</emphasis>
has a stronger, more or less bladelike antero-inner lateral costa, and a costa along the posterior margin (
<figureCitation id="27D137B3CB00FF16FF15FBDEFEF8FBF1" box="[208,320,1046,1069]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26C–E</figureCitation>
); outer lateral face smooth or with a few weak, short costae near the base.
<taxonomicName id="78EA50B5CB00FF16FE2FFBFBFDE5FB96" box="[490,605,1075,1098]" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">
<emphasis id="8D9EF724CB00FF16FE2FFBFBFDE5FB96" box="[490,605,1075,1098]" italics="true" pageId="43" pageNumber="212">Sa element</emphasis>
</taxonomicName>
symmetrical, with a reclined cusp which is more or less rounded in cross section, a broad smooth anterior face, and bearing three or more costae on each posterolateral face (
<figureCitation id="27D137B3CB00FF16FDF9FB43FD12FB7E" box="[572,682,1163,1186]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26F–I</figureCitation>
).
<taxonomicName id="78EA50B5CB00FF16FD17FB43FF75FB63" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">
<emphasis id="8D9EF724CB00FF16FD17FB43FF75FB63" italics="true" pageId="43" pageNumber="212">Sb element</emphasis>
</taxonomicName>
slightly asymmetrical, laterally compressed; cusp proclined with a broad anterior face and sharp posterior margin; the inner lateral face with two to four stronger costa and several finer ones in between; outer lateral face with a stronger anterolateral costa and several finer ones posterior to it (
<figureCitation id="27D137B3CB00FF16FF77FAF3FED9FA8E" box="[178,353,1339,1362]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26J,K,M,N</figureCitation>
).
<taxonomicName id="78EA50B5CB00FF16FEBFFAF3FE56FA8E" box="[378,494,1339,1362]" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">
<emphasis id="8D9EF724CB00FF16FEBFFAF3FE56FA8E" box="[378,494,1339,1362]" italics="true" pageId="43" pageNumber="212">Sc element</emphasis>
</taxonomicName>
asymmetrical, laterally compressed; cusp suberect with a sharp posterior costa along the posterior margin and a sharp anterolateral costa on the inner side (
<figureCitation id="27D137B3CB00FF16FF35FA5BFEE8FA76" box="[240,336,1427,1450]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26L</figureCitation>
), as well as a number of weaker costae (typically three or four) on each lateral face; costae on the outer lateral face weaker than those on the inner lateral face (
<figureCitation id="27D137B3CB00FF16FF44FA23FF53F9DE" box="[129,235,1515,1538]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26 O</figureCitation>
).
<taxonomicName id="78EA50B5CB00FF16FEC2FA23FEC6F9DE" box="[263,382,1515,1538]" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">
<emphasis id="8D9EF724CB00FF16FEC2FA23FEC6F9DE" box="[263,382,1515,1538]" italics="true" pageId="43" pageNumber="212">Sd element</emphasis>
</taxonomicName>
nearly symmetrical, less laterally compressed in comparison with the Sb and
<taxonomicName id="78EA50B5CB00FF16FDA4F9C0FD50F9C3" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[609,744,1544,1567]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">Sc elements</taxonomicName>
; basal cavity opening rounded (often flared); broad anterior face, a costa along the posterior margin, with several costae on each of the lateral faces; the anterolateral costa on each side stronger than others (
<figureCitation id="27D137B3CB00FF16FE54F9B6FDB7F949" box="[401,527,1662,1685]" captionStart="Fig" captionStartId="44.[166,202,1884,1905]" captionTargetBox="[166,1466,136,1867]" captionTargetId="figure-9@44.[166,1467,136,1867]" captionTargetPageId="44" captionText="Fig. 26. Scolopodus multicostatus Barnes &amp; Tuke, 1970: A,B, Pa element, AMF120460, Y4–2, A, inner lateral view, B, outer lateral view; C,D, Pb element, AMF120461, M/A11-2, C, outer lateral view, D, postero-inner lateral view; E, Pb element, AMF120462, M/ A11-2, posterior view; F,?Sa element, AMF120463, TAB1/8.1, basal view; G–I,?Sa element, AMF120464, C1613, G,H, anterolateral views, I, anterior view; J,K, Sb element, AMF120465, C1613, J, inner lateral view, K, outer lateral view; L, Sc element, AMF120468, M/A11-1, inner lateral view; M,N, Sb element, AMF120466, C1613, M, inner lateral view, N, outer lateral view; O,P, Sc element, AMF120467, C1613, O, outer lateral view, P, inner lateral view; Q,R, Sd element,AMF120469, C1613, lateral views. Scale bars 100 µm." figureDoi="http://doi.org/10.5281/zenodo.10092844" httpUri="https://zenodo.org/record/10092844/files/figure.png" pageId="43" pageNumber="212">Fig. 26Q,R</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="F7F078BDCB00FF16FFBCF96EFAEBFE09" pageId="43" pageNumber="212" type="discussion">
<paragraph id="BF552B36CB00FF16FFBCF96EFAEBFE09" blockId="43.[121,749,1702,2018]" lastBlockId="43.[794,1422,153,469]" pageId="43" pageNumber="212">
<emphasis id="8D9EF724CB00FF16FFBCF96EFF58F962" bold="true" box="[121,224,1702,1726]" pageId="43" pageNumber="212">Remarks</emphasis>
. Two species of
<taxonomicName id="78EA50B5CB00FF16FE4EF96FFDBDF962" authorityName="Pander" authorityYear="1856" box="[395,517,1703,1726]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="genus">
<emphasis id="8D9EF724CB00FF16FE4EF96FFDBDF962" box="[395,517,1703,1726]" italics="true" pageId="43" pageNumber="212">Scolopodus</emphasis>
</taxonomicName>
are present in western
<collectingRegion id="7D2EE5D4CB00FF16FFBCF90BFEF2F906" box="[121,330,1731,1754]" country="Australia" name="New South Wales" pageId="43" pageNumber="212">New South Wales</collectingRegion>
;
<taxonomicName id="78EA50B5CB00FF16FE9CF90CFDB5F907" authorityName="Barnes &amp; Tuke" authorityYear="1970" box="[345,525,1732,1755]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FE9CF90CFDB5F907" box="[345,525,1732,1755]" italics="true" pageId="43" pageNumber="212">S. multicostatus</emphasis>
</taxonomicName>
is distinguished in generally having fewer costae, which are also much weaker and not as sharp-edged as in the co-occurring
<taxonomicName id="78EA50B5CB00FF16FD9AF937FD5DF8CA" authorityName="Pander" authorityYear="1856" box="[607,741,1791,1814]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="quadratus">
<emphasis id="8D9EF724CB00FF16FD9AF937FD5DF8CA" box="[607,741,1791,1814]" italics="true" pageId="43" pageNumber="212">S. quadratus</emphasis>
</taxonomicName>
. The Newfoundland
<typeStatus id="60519594CB00FF16FE9BF8D3FE35F8EE" box="[350,397,1819,1842]" pageId="43" pageNumber="212">type</typeStatus>
specimens of
<taxonomicName id="78EA50B5CB00FF16FDFDF8D4FD55F8EF" authorityName="Barnes &amp; Tuke" authorityYear="1970" box="[568,749,1820,1843]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FDFDF8D4FD55F8EF" box="[568,749,1820,1843]" italics="true" pageId="43" pageNumber="212">S. multicostatus</emphasis>
</taxonomicName>
figured by Barnes &amp; Tuke (1970), which are all slightly asymmetrical with a short base and erect cusp, are identical to the
<collectingRegion id="7D2EE5D4CB00FF16FF7FF8BBFECFF856" box="[186,375,1907,1930]" country="Australia" name="New South Wales" pageId="43" pageNumber="212">New South Wales</collectingRegion>
<taxonomicName id="78EA50B5CB00FF16FEB8F8BBFDBAF856" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[381,514,1907,1930]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">Sb elements</taxonomicName>
. Stait &amp; Druce (1993) recognized a seximembrate apparatus for
<taxonomicName id="78EA50B5CB00FF16FD85F859FD54F874" authorityName="Barnes &amp; Tuke" authorityYear="1970" box="[576,748,1937,1960]" class="Conodonta" family="Strachanognathidae" genus="Scolopodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Protopanderodontida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="multicostatus">
<emphasis id="8D9EF724CB00FF16FD85F859FD54F874" box="[576,748,1937,1960]" italics="true" pageId="43" pageNumber="212">S. multicostatus</emphasis>
</taxonomicName>
from the Coolibah Formation of the Georgina Basin, including scandodiform (Pa), posteriorly keeled scandodiform (Pb), acontiodiform (Sa), planoconvex (Sb), laterally compressed paltodiform (Sc) and equidimensional paltodiform (Sd) elements. The Coolibah Formation material generally exhibits more numerous, strongly developed costae than do the specimens in western
<collectingRegion id="7D2EE5D4CB00FF16FA9EFEC6FC1FFE9E" country="Australia" name="New South Wales" pageId="43" pageNumber="212">New South Wales</collectingRegion>
; however,
<taxonomicName id="78EA50B5CB00FF16FBE7FEE3FB1FFE9E" baseAuthorityName="Repetski" baseAuthorityYear="1982" box="[1058,1191,299,322]" class="Conodonta" family="Acanthodontidae" genus="Scalpellodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="elements">Pa elements</taxonomicName>
from both areas are identical. Specimens referred to the symmetrical
<taxonomicName id="78EA50B5CB00FF16FADFFE81FA35FEBC" box="[1306,1421,329,352]" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">Sa element</taxonomicName>
in our collections have weakly developed costae, a broader anterior face, and a more strongly reclined cusp in comparison with the
<taxonomicName id="78EA50B5CB00FF16FBDCFE69FB23FE64" box="[1049,1179,417,440]" class="Conodonta" family="Protopanderodontidae" genus="Protopanderodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Conodontophorida" pageId="43" pageNumber="212" phylum="Chordata" rank="species" species="element">Sa element</taxonomicName>
illustrated from the Coolibah Formation (Stait &amp; Druce, 1993, fig. 19L).
</paragraph>
</subSubSection>
</treatment>
</document>
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