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<document id="E45B9143A56029EA69172088AB3AF9C1" ID-DOI="10.11646/zootaxa.5338.1.1" ID-GBIF-Dataset="e7148a02-ac1a-4172-9071-606bd1e703fc" ID-ISSN="1175­5334" ID-Zenodo-Dep="8283738" ID-ZooBank="CAA37212-3008-43F4-98DE-94D7A3FFD3B6" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="karina" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1692955869775" checkinUser="plazi" docAuthor="Mu, Yannan &amp; Zhang, Feng" docDate="2023" docId="364E87DE8656FFA15AA6FE5DDAC12443" docLanguage="en" docName="zootaxa.5338.1.1.pdf" docOrigin="Zootaxa 5338 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5338.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Otacilia impleta, sp. nov." docType="treatment" docVersion="7" lastPageNumber="41" masterDocId="CA77FFA68673FF895A31FFEBD84D203E" masterDocTitle="Further additions to the guardstone spider fauna from China (Araneae: Phrurolithidae)" masterLastPageNumber="104" masterPageNumber="1" pageNumber="38" updateTime="1720090975140" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="79E98009D1B6EBF007D33BD453C66CB9">Further additions to the guardstone spider fauna from China (Araneae: Phrurolithidae)</mods:title>
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<mods:namePart id="100AAB227D6FDF499D1E5E64EF436FF9">Mu, Yannan</mods:namePart>
<mods:affiliation id="022B596655AEFC8104FC81BC6112116F">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Developed, Hebei University, Baoding, Hebei 071002, P. R. China &amp; 15188605531 @ 163. com; https: // orcid. org / 0000 - 0002 - 2504 - 673 X</mods:affiliation>
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<mods:namePart id="89FA7BBA9FF57CF08633735239697F9E">Zhang, Feng</mods:namePart>
<mods:affiliation id="0C7971728D81B4ECE9B40F390CD9D42F">The Key Laboratory of Zoological Systematics and Application, Institute of Life Science and Green Developed, Hebei University, Baoding, Hebei 071002, P. R. China</mods:affiliation>
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<treatment id="364E87DE8656FFA15AA6FE5DDAC12443" ID-DOI="http://doi.org/10.5281/zenodo.8283847" ID-GBIF-Taxon="212459940" ID-Zenodo-Dep="8283847" LSID="urn:lsid:plazi:treatment:364E87DE8656FFA15AA6FE5DDAC12443" httpUri="http://treatment.plazi.org/id/364E87DE8656FFA15AA6FE5DDAC12443" lastPageId="40" lastPageNumber="41" pageId="37" pageNumber="38">
<subSubSection id="F6FD65438656FFAC5AA6FE5DDA0321EE" box="[151,590,438,465]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="BE5836C88656FFAC5AA6FE5DDA0321EE" blockId="37.[151,590,438,501]" box="[151,590,438,465]" pageId="37" pageNumber="38">
<heading id="E51081A48656FFAC5AA6FE5DDA0321EE" bold="true" box="[151,590,438,465]" fontSize="11" level="1" pageId="37" pageNumber="38" reason="1">
<emphasis id="8C93EADA8656FFAC5AA6FE5DDA0321EE" bold="true" box="[151,590,438,465]" pageId="37" pageNumber="38">
<taxonomicName id="79E74D4B8656FFAC5AA6FE5DD91421EF" ID-CoL="C7XLC" authority="Mu &amp; Zhang, 2023" box="[151,345,438,465]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="impleta" status="sp. nov.">
<emphasis id="8C93EADA8656FFAC5AA6FE5DD91421EF" bold="true" box="[151,345,438,465]" italics="true" pageId="37" pageNumber="38">Otacilia impleta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97A057A18656FFAC5B51FE5CD9F021EF" box="[352,445,439,465]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
(ffiŴṚff)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6FD65438656FFAC5AA6FE30D95821CB" box="[151,277,475,501]" pageId="37" pageNumber="38" type="description">
<paragraph id="BE5836C88656FFAC5AA6FE30D95821CB" blockId="37.[151,590,438,501]" box="[151,277,475,501]" pageId="37" pageNumber="38">
<figureCitation id="26DC2A4D8656FFAC5AA6FE30D95821CB" box="[151,277,475,501]" captionStart-0="FIGURE 28" captionStart-1="FIGURE 29" captionStart-2="FIGURE 30" captionStartId-0="37.[151,250,1882,1906]" captionStartId-1="38.[151,250,1850,1874]" captionStartId-2="39.[151,250,1514,1538]" captionTargetBox-0="[161,1432,633,1854]" captionTargetBox-1="[151,1436,188,1815]" captionTargetBox-2="[156,1435,186,1485]" captionTargetId-0="figure-127@37.[151,1436,633,1858]" captionTargetId-1="figure-21@38.[151,1436,181,1826]" captionTargetId-2="figure-17@39.[151,1435,181,1490]" captionTargetPageId-0="37" captionTargetPageId-1="38" captionTargetPageId-2="39" captionText-0="FIGURE 28. Habitus of Otacilia impleta sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." captionText-1="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." captionText-2="FIGURE 30. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, ventral view; B. Same, retrolateral view; C. Same, dorsal view; D. Epigyne, ventral view; E. Same, dorsal view. Scale bars: 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8283835" figureDoi-1="http://doi.org/10.5281/zenodo.8283839" figureDoi-2="http://doi.org/10.5281/zenodo.8283843" httpUri-0="https://zenodo.org/record/8283835/files/figure.png" httpUri-1="https://zenodo.org/record/8283839/files/figure.png" httpUri-2="https://zenodo.org/record/8283843/files/figure.png" pageId="37" pageNumber="38">Figs 2830</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F6FD65438656FFAE5AA6FDC8DAF026AC" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="BE5836C88656FFAC5AA6FDC8DD28225F" blockId="37.[151,1437,546,611]" pageId="37" pageNumber="38">
<emphasis id="8C93EADA8656FFAC5AA6FDC8D9092202" bold="true" box="[151,324,546,573]" pageId="37" pageNumber="38">Type material.</emphasis>
<materialsCitation id="0E8F3C958656FFAC5B7BFDC8DD28225F" ID-GBIF-Occurrence="4400675348" collectingDate="2021-07-26" collectorName="Yannan Mu." country="China" county="Nanjian Yi Autonomous County" elevation="2294" latitude="24.892597" location="Fenghuang Mountain" longLatPrecision="1" longitude="100.32852" municipality="Dali Bai Autonomous Prefecture" pageId="37" pageNumber="38" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="8C93EADA8656FFAC5B7BFDC8D9FA2203" bold="true" box="[330,439,547,573]" pageId="37" pageNumber="38">
<typeStatus id="615C886A8656FFAC5B7BFDC8D9FA2203" box="[330,439,547,573]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
</emphasis>
♁,
<emphasis id="8C93EADA8656FFAC5BEEFDC9DA0B2202" bold="true" box="[479,582,546,572]" pageId="37" pageNumber="38">
<collectingCountry id="C6F076588656FFAC5BEEFDC9DA0C2202" box="[479,577,546,572]" name="China" pageId="37" pageNumber="38">CHINA</collectingCountry>
:
</emphasis>
<emphasis id="8C93EADA8656FFAC587DFDC8DB442202" box="[588,777,547,572]" italics="true" pageId="37" pageNumber="38">
<collectingRegion id="7C23F82A8656FFAC587DFDC8DB442202" box="[588,777,547,572]" country="China" name="Yunnan" pageId="37" pageNumber="38">Yunnan Province</collectingRegion>
</emphasis>
:
<collectingMunicipality id="5E3CACB28656FFAC5926FDC8DCC82203" box="[791,1157,547,573]" pageId="37" pageNumber="38">Dali Bai Autonomous Prefecture</collectingMunicipality>
,
<collectingCounty id="57394E448656FFAC5EBEFDC8D8A6225F" pageId="37" pageNumber="38">Nanjian Yi Autonomous County</collectingCounty>
,
<location id="BB3860138656FFAC5AC7FDACD9AB225F" LSID="urn:lsid:plazi:treatment:364E87DE8656FFA15AA6FE5DDAC12443:BB3860138656FFAC5AC7FDACD9AB225F" box="[246,486,583,609]" country="China" county="Nanjian Yi Autonomous County" latitude="24.892597" longLatPrecision="1" longitude="100.32852" municipality="Dali Bai Autonomous Prefecture" name="Fenghuang Mountain" pageId="37" pageNumber="38" stateProvince="Yunnan">Fenghuang Mountain</location>
(
<geoCoordinate id="DBD3500F8656FFAC5BC4FDA3DAEF225D" box="[501,674,584,611]" degrees="24" direction="north" minutes="53" orientation="latitude" pageId="37" pageNumber="38" precision="1" seconds="33.35" value="24.892597">24°5333.35″N</geoCoordinate>
,
<geoCoordinate id="DBD3500F8656FFAC589CFDA3DB2F225D" box="[685,866,584,611]" degrees="100" direction="east" minutes="19" orientation="longitude" pageId="37" pageNumber="38" precision="1" seconds="42.67" value="100.32852">100°1942.67″E</geoCoordinate>
,
<elevation id="35CAD1FB8656FFAC595FFDACDBB3225F" box="[878,1022,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.294" pageId="37" pageNumber="38" unit="m" value="2294.0">
<quantity id="791F9B2D8656FFAC595FFDACDB8E225F" box="[878,963,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.294" pageId="37" pageNumber="38" unit="m" value="2294.0">2294 m</quantity>
a.s.l.
</elevation>
),
<date id="CA5910088656FFAC5E24FDACDCEF225F" box="[1045,1186,583,609]" pageId="37" pageNumber="38" value="2021-07-26">
<collectingDate id="DA1DE9E08656FFAC5E24FDACDCEF225F" box="[1045,1186,583,609]" pageId="37" pageNumber="38" value="2021-07-26">26 Jul. 2021</collectingDate>
</date>
, leg.
<collectorName id="1312531E8656FFAC5EEDFDACDD28225F" box="[1244,1381,583,609]" pageId="37" pageNumber="38">Yannan Mu.</collectorName>
</materialsCitation>
</paragraph>
<caption id="EA9866408656FFAC5AA6F8B1DA4F27A8" ID-DOI="http://doi.org/10.5281/zenodo.8283835" ID-Zenodo-Dep="8283835" httpUri="https://zenodo.org/record/8283835/files/figure.png" pageId="37" pageNumber="38" startId="37.[151,250,1882,1906]" targetBox="[161,1432,633,1854]" targetPageId="37" targetType="figure">
<paragraph id="BE5836C88656FFAC5AA6F8B1DA4F27A8" blockId="37.[151,1437,1882,1942]" pageId="37" pageNumber="38">
<emphasis id="8C93EADA8656FFAC5AA6F8B1D96D274C" bold="true" box="[151,288,1882,1906]" pageId="37" pageNumber="38">FIGURE 28.</emphasis>
Habitus of
<taxonomicName id="79E74D4B8656FFAC5BA7F8B1DA71274C" box="[406,572,1882,1906]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="impleta" status="sp. nov.">
<emphasis id="8C93EADA8656FFAC5BA7F8B1DA71274C" box="[406,572,1882,1906]" italics="true" pageId="37" pageNumber="38">Otacilia impleta</emphasis>
</taxonomicName>
<emphasis id="8C93EADA8656FFAC5873F8B1DADF274C" bold="true" box="[578,658,1882,1906]" pageId="37" pageNumber="38">
<taxonomicNameLabel id="97A057A18656FFAC5873F8B1DADF274C" box="[578,658,1882,1906]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="8C93EADA8656FFAC5891F8B1DAF4274C" bold="true" box="[672,697,1882,1906]" pageId="37" pageNumber="38">A.</emphasis>
Male holotype, dorsal view;
<emphasis id="8C93EADA8656FFAC59D0F8B1DBB4274C" bold="true" box="[993,1017,1882,1906]" pageId="37" pageNumber="38">B.</emphasis>
Same, ventral view;
<emphasis id="8C93EADA8656FFAC5EFEF8B1DCAA274C" bold="true" box="[1231,1255,1882,1906]" pageId="37" pageNumber="38">C.</emphasis>
Female paratype, dorsal view;
<emphasis id="8C93EADA8656FFAC5B2BF895D97E27A8" bold="true" box="[282,307,1918,1942]" pageId="37" pageNumber="38">D.</emphasis>
Same, ventral view.
</paragraph>
</caption>
<caption id="EA9866408655FFAF5AA6F8D1DC4627DD" ID-DOI="http://doi.org/10.5281/zenodo.8283839" ID-Zenodo-Dep="8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="38" pageNumber="39" startId="38.[151,250,1850,1874]" targetBox="[151,1436,188,1815]" targetPageId="38" targetType="figure">
<paragraph id="BE5836C88655FFAF5AA6F8D1DC4627DD" blockId="38.[151,1437,1850,2019]" pageId="38" pageNumber="39">
<emphasis id="8C93EADA8655FFAF5AA6F8D1D96F276C" bold="true" box="[151,290,1850,1874]" pageId="38" pageNumber="39">FIGURE 29.</emphasis>
Copulatory organs of
<taxonomicName id="79E74D4B8655FFAF583DF8D0DAFE276D" box="[524,691,1851,1875]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="impleta" status="sp. nov.">
<emphasis id="8C93EADA8655FFAF583DF8D0DAFE276D" box="[524,691,1851,1875]" italics="true" pageId="38" pageNumber="39">Otacilia impleta</emphasis>
</taxonomicName>
<emphasis id="8C93EADA8655FFAF588AF8D0DB40276D" bold="true" box="[699,781,1851,1875]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="97A057A18655FFAF588AF8D0DB40276D" box="[699,781,1851,1875]" pageId="38" pageNumber="39" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="8C93EADA8655FFAF592CF8D1DB78276C" bold="true" box="[797,821,1850,1874]" pageId="38" pageNumber="39">A.</emphasis>
Male left palp, prolateral view;
<emphasis id="8C93EADA8655FFAF5EB6F8D0DCD3276D" bold="true" box="[1159,1182,1851,1875]" pageId="38" pageNumber="39">B.</emphasis>
Same, retrolateral view;
<emphasis id="8C93EADA8655FFAF5AA6F8B5D8FD2748" bold="true" box="[151,176,1886,1910]" pageId="38" pageNumber="39">C.</emphasis>
Same, ventral view;
<emphasis id="8C93EADA8655FFAF5BBEF8B4D9E52749" bold="true" box="[399,424,1887,1911]" pageId="38" pageNumber="39">D.</emphasis>
Same, dorsal view;
<emphasis id="8C93EADA8655FFAF584FF8B4DADB2749" bold="true" box="[638,662,1887,1911]" pageId="38" pageNumber="39">E.</emphasis>
Epigyne, ventral view;
<emphasis id="8C93EADA8655FFAF59A0F8B4DBE92749" bold="true" box="[913,932,1887,1911]" pageId="38" pageNumber="39">F.</emphasis>
Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae.
</paragraph>
</caption>
<caption id="EA9866408654FFAE5AA6FA01DC5A2618" ID-DOI="http://doi.org/10.5281/zenodo.8283843" ID-Zenodo-Dep="8283843" httpUri="https://zenodo.org/record/8283843/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,250,1514,1538]" targetBox="[156,1435,186,1485]" targetPageId="39" targetType="figure">
<paragraph id="BE5836C88654FFAE5AA6FA01DC5A2618" blockId="39.[151,1436,1514,1574]" pageId="39" pageNumber="40">
<emphasis id="8C93EADA8654FFAE5AA6FA01D96C263C" bold="true" box="[151,289,1514,1538]" pageId="39" pageNumber="40">FIGURE 30.</emphasis>
Copulatory organs of
<taxonomicName id="79E74D4B8654FFAE583BFA01DAFC263C" box="[522,689,1514,1538]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="impleta" status="sp. nov.">
<emphasis id="8C93EADA8654FFAE583BFA01DAFC263C" box="[522,689,1514,1538]" italics="true" pageId="39" pageNumber="40">Otacilia impleta</emphasis>
</taxonomicName>
<emphasis id="8C93EADA8654FFAE5888FA01DB46263C" bold="true" box="[697,779,1514,1538]" pageId="39" pageNumber="40">
<taxonomicNameLabel id="97A057A18654FFAE5888FA01DB46263C" box="[697,779,1514,1538]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
:
<emphasis id="8C93EADA8654FFAE592BFA01DB7E263C" bold="true" box="[794,819,1514,1538]" pageId="39" pageNumber="40">A.</emphasis>
Male left palp, ventral view;
<emphasis id="8C93EADA8654FFAE5E56FA01DC32263C" bold="true" box="[1127,1151,1514,1538]" pageId="39" pageNumber="40">B.</emphasis>
Same, retrolateral view;
<emphasis id="8C93EADA8654FFAE5FB5FA01DDD1263C" bold="true" box="[1412,1436,1514,1538]" pageId="39" pageNumber="40">C.</emphasis>
Same, dorsal view;
<emphasis id="8C93EADA8654FFAE5B6EF9E5D9352618" bold="true" box="[351,376,1550,1574]" pageId="39" pageNumber="40">D.</emphasis>
Epigyne, ventral view;
<emphasis id="8C93EADA8654FFAE585BF9E5DACF2618" bold="true" box="[618,642,1550,1574]" pageId="39" pageNumber="40">E.</emphasis>
Same, dorsal view. Scale bars: 0.2 mm.
</paragraph>
</caption>
<paragraph id="BE5836C88654FFAE5AF6F993DAF026AC" blockId="39.[151,1437,1656,2043]" box="[199,701,1656,1683]" pageId="39" pageNumber="40">
<materialsCitation id="0E8F3C958654FFAE5AF6F993DAF426AC" ID-GBIF-Occurrence="4400675304" box="[199,697,1656,1683]" collectingDate="2021-07-26" collectorName="Yannan Mu." country="China" county="Nanjian Yi Autonomous County" elevation="2294" latitude="24.892597" location="Fenghuang Mountain" longLatPrecision="1" longitude="100.32852" municipality="Dali Bai Autonomous Prefecture" pageId="39" pageNumber="40" specimenCount="6" specimenCount-female="6" stateProvince="Yunnan" typeStatus="paratype">
<emphasis id="8C93EADA8654FFAE5AF6F993D97226AC" bold="true" box="[199,319,1656,1682]" pageId="39" pageNumber="40">
<typeStatus id="615C886A8654FFAE5AF6F993D97226AC" box="[199,319,1656,1682]" pageId="39" pageNumber="40" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="A8E1FD418654FFAE5B7FF993D93526AD" box="[334,376,1656,1683]" pageId="39" pageNumber="40" type="female">6 ♀</specimenCount>
, with same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F6FD65438654FFAE5AF6F977DA6126E4" pageId="39" pageNumber="40" type="etymology">
<paragraph id="BE5836C88654FFAE5AF6F977DA6126E4" blockId="39.[151,1437,1656,2043]" pageId="39" pageNumber="40">
<emphasis id="8C93EADA8654FFAE5AF6F977D9032688" bold="true" box="[199,334,1692,1718]" pageId="39" pageNumber="40">Etymology.</emphasis>
The specific name is derived from the Latin word “
<emphasis id="8C93EADA8654FFAE59A6F977DBBB2688" box="[919,1014,1692,1718]" italics="true" pageId="39" pageNumber="40">impletus</emphasis>
”, referring to the satisfaction derived from naming this species; adjective.
</paragraph>
</subSubSection>
<subSubSection id="F6FD65438654FFAE5AF6F90FDA3D278C" pageId="39" pageNumber="40" type="diagnosis">
<paragraph id="BE5836C88654FFAE5AF6F90FDA3D278C" blockId="39.[151,1437,1656,2043]" pageId="39" pageNumber="40">
<emphasis id="8C93EADA8654FFAE5AF6F90FD90F26C0" bold="true" box="[199,322,1764,1790]" pageId="39" pageNumber="40">Diagnosis.</emphasis>
This new species resembles
<taxonomicName id="79E74D4B8654FFAE5844F90EDA9526C0" baseAuthorityName="Fu, Chen &amp; Zhang" baseAuthorityYear="2016" box="[629,728,1764,1791]" class="Arachnida" family="Phrurolithidae" genus="Otacilia" kingdom="Animalia" order="Araneae" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="celata">
<emphasis id="8C93EADA8654FFAE5844F90EDA9526C0" box="[629,728,1764,1791]" italics="true" pageId="39" pageNumber="40">O.celata</emphasis>
</taxonomicName>
in having round copulatory openings and globular spermathecae, but can be recognized by: 1) the curved, arc-shaped retrolateral tibial apophysis (vs not curved, cf.
<figureCitation id="26DC2A4D8654FFAE5ED1F8E3DD0C271C" box="[1248,1345,1800,1826]" captionStart="FIGURE 29" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[151,1436,188,1815]" captionTargetId="figure-21@38.[151,1436,181,1826]" captionTargetPageId="38" captionText="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="39" pageNumber="40">Fig. 29C</figureCitation>
and fig. 2B in Fu
<emphasis id="8C93EADA8654FFAE5ACCF8C6D9792778" box="[253,308,1836,1862]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2016), 2) the longer, upwards-extending embolus curved towards the retrolateral side of the cymbium (vs short, not curved, cf.
<figureCitation id="26DC2A4D8654FFAE5B81F8BBDA5F2754" box="[432,530,1872,1898]" captionStart="FIGURE 29" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[151,1436,188,1815]" captionTargetId="figure-21@38.[151,1436,181,1826]" captionTargetPageId="38" captionText="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="39" pageNumber="40">Fig. 29C</figureCitation>
and fig. 2B in Fu
<emphasis id="8C93EADA8654FFAE58D1F8BADB552754" box="[736,792,1872,1898]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2016), 3) the smaller, ridge-shaped tegular apophysis (vs large, triangular, cf.
<figureCitation id="26DC2A4D8654FFAE5B4BF89FD99127B0" box="[378,476,1908,1934]" captionStart="FIGURE 29" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[151,1436,188,1815]" captionTargetId="figure-21@38.[151,1436,181,1826]" captionTargetPageId="38" captionText="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="39" pageNumber="40">Fig. 29C</figureCitation>
and fig. 1D in Fu
<emphasis id="8C93EADA8654FFAE589BF89EDAAE27B0" box="[682,739,1908,1934]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2016), 4) thinner, longer connecting tubes (vs thick and short, cf.
<figureCitation id="26DC2A4D8654FFAE5A8AF873D957278C" box="[187,282,1944,1970]" captionStart="FIGURE 29" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[151,1436,188,1815]" captionTargetId="figure-21@38.[151,1436,181,1826]" captionTargetPageId="38" captionText="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="39" pageNumber="40">Fig. 29F</figureCitation>
and fig. 1G in Fu
<emphasis id="8C93EADA8654FFAE5BD9F872DA6D278C" box="[488,544,1944,1970]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
2016).
</paragraph>
</subSubSection>
<subSubSection id="F6FD65438654FFA15AF6F857DDDB2467" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="BE5836C88654FFA15AF6F857DD5F217F" blockId="39.[151,1437,1656,2043]" lastBlockId="40.[151,1437,150,1149]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="8C93EADA8654FFAE5AF6F857D91427E8" bold="true" box="[199,345,1980,2006]" pageId="39" pageNumber="40">Description.</emphasis>
Male (
<typeStatus id="615C886A8654FFAE5B96F857DA5B27E8" box="[423,534,1980,2006]" pageId="39" pageNumber="40" type="holotype">Holotype</typeStatus>
): total length 2.40, carapace 1.13 long, 1.11 wide; abdomen 1.27 long, 0.84 wide. Eye sizes and interdistances:AME 0.06, ALE 0.08, PME 0.05, PLE 0.08; AMEAME 0.03, AMEALE 0.01, ALE ALE 0.16, PMEPME 0.05, PMEPLE 0.05, PLEPLE 0.24, ALEPLE 0.05. EAW 0.38, CRW 0.55, EAW/CRW 0.69, CRW/CW 0.50. MOA 0.19 long, anterior width 0.15, posterior width 0.16.
<collectingCountry id="C6F07658865BFFA15E14FF50DC0120EB" box="[1061,1100,187,213]" name="Switzerland" pageId="40" pageNumber="41">CH</collectingCountry>
0.11,
<collectingCountry id="C6F07658865BFFA15EA3FF50DCF520EB" box="[1170,1208,187,213]" name="Switzerland" pageId="40" pageNumber="41">CH</collectingCountry>
/AME 1.83. Labium 0.06 long, 0.18 wide. Sternum 0.57 long, 0.53 wide. Leg measurements: 4.24 (1.18, 0.47, 1.08, 0.95, 0.56), II 3.66 (1.02, 0.42, 0.84, 0.80, 0.58), III 3.25 (0.87, 0.40, 0.64, 0.80, 0.54), IV 4.95 (1.31, 0.45, 1.13, 1.30, 0.76). Spination: femora IIV d 1, femur I pl 3, femur II pl 1, tibia pv 6 rv 6, tibia II pv 6 rv 5, metatarsi II pv 4 rv 3.
</paragraph>
<paragraph id="BE5836C8865BFFA15AF6FEA0DCFC2193" blockId="40.[151,1437,150,1149]" pageId="40" pageNumber="41">
Coloration. Carapace dark brown, with several shapes resembling flowing water droplets beside fovea.Abdomen black, dorsum nearly entirely covered by scutum, with pair of oval yellow markings near middle and two W-shaped stripes posteriorly (
<figureCitation id="26DC2A4D865BFFA15B41FE78D9B52193" box="[368,504,403,429]" captionStart="FIGURE 28" captionStartId="37.[151,250,1882,1906]" captionTargetBox="[161,1432,633,1854]" captionTargetId="figure-127@37.[151,1436,633,1858]" captionTargetPageId="37" captionText="FIGURE 28. Habitus of Otacilia impleta sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8283835" httpUri="https://zenodo.org/record/8283835/files/figure.png" pageId="40" pageNumber="41">Fig. 28A, B</figureCitation>
). Legs yellow, with black annuli near tip of femora and tibiae.
</paragraph>
<paragraph id="BE5836C8865BFFA15AF6FE5CDB2822BB" blockId="40.[151,1437,150,1149]" pageId="40" pageNumber="41">
Palp as in
<figureCitation id="26DC2A4D865BFFA15B7AFE5CD99121EF" box="[331,476,439,465]" captionStart="FIGURE 29" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[151,1436,188,1815]" captionTargetId="figure-21@38.[151,1436,181,1826]" captionTargetPageId="38" captionText="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="40" pageNumber="41">Figs 29AD</figureCitation>
,
<figureCitation id="26DC2A4D865BFFA15BDCFE5CDA0F21EF" box="[493,578,439,465]" captionStart="FIGURE 30" captionStartId="39.[151,250,1514,1538]" captionTargetBox="[156,1435,186,1485]" captionTargetId="figure-17@39.[151,1435,181,1490]" captionTargetPageId="39" captionText="FIGURE 30. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, ventral view; B. Same, retrolateral view; C. Same, dorsal view; D. Epigyne, ventral view; E. Same, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8283843" httpUri="https://zenodo.org/record/8283843/files/figure.png" pageId="40" pageNumber="41">30AC</figureCitation>
. Femoral apophysis protruding slightly, round, at middle parts of femur. Retrolateral tibial apophysis strongly curved, arc-shaped in ventral view; dorsal tibial apophysis smaller than retrolateral tibial apophysis, with broad base and thin tip, leaf-shaped in dorsal view. Tegulum slightly wider than cymbium, oval; tegular apophysis small, triangular, located at retrolateral tip of tegulum. Sperm duct distinct, tapering from retrolateral side of tegulum to embolus. Embolus short, base wide, triangular, tip curved towards retrolateral side of cymbium tip. Conductor narrow, transparent.
</paragraph>
<paragraph id="BE5836C8865BFFA15AF6FD64DA2823F7" blockId="40.[151,1437,150,1149]" pageId="40" pageNumber="41">
Female (One
<typeStatus id="615C886A865BFFA15B53FD64D9852297" box="[354,456,655,681]" pageId="40" pageNumber="41" type="paratype">paratype</typeStatus>
): total length 2.68, carapace 1.29 long, 1.12 wide; abdomen 1.39 long, 1.10 wide. Eye sizes and interdistances: AME 0.06, ALE 0.10, PME 0.06, PLE 0.08; AMEAME 0.04, AMEALE 0.01, ALE ALE 0.18, PMEPME 0.07, PMEPLE 0.06, PLEPLE 0.31, ALEPLE 0.06. EAW 0.42, CRW 0.60, EAW/CRW 0.70, CRW/CW 0.57. MOA 0.420 long, anterior width 0.17, posterior width 0.19.
<collectingCountry id="C6F07658865BFFA15E19FD10DC02232B" box="[1064,1103,763,789]" name="Switzerland" pageId="40" pageNumber="41">CH</collectingCountry>
0.10,
<collectingCountry id="C6F07658865BFFA15EA2FD10DCF4232B" box="[1171,1209,763,789]" name="Switzerland" pageId="40" pageNumber="41">CH</collectingCountry>
/AME 1.67. Labium 0.14 long, 0.19 wide. Sternum 0.82 long, 0.69 wide. Leg measurements: 4.64 (1.51, 0.47, 1.15, 0.97, 0.54), II 3.67 (1.01, 0.42, 0.88, 0.83, 0.53), III 3.25 (0.86, 0.38, 0.68, 0.80, 0.53), IV 4.82 (1.29, 0.46, 1.09, 1.21, 0.77). Spination: femora IIV d 1, femur I pl 3, femur II pl 1, tibia pv 6 rv 6, tibia II pv 6 rv 5, metatarsi II pv 4 rv 3. Carapace color and pattern same as male, abdomen black, without dorsal scutum, covered by fluff, several white spots and stripes faint in dorsal view (
<figureCitation id="26DC2A4D865BFFA15BE1FC44DA1423F7" box="[464,601,943,969]" captionStart="FIGURE 28" captionStartId="37.[151,250,1882,1906]" captionTargetBox="[161,1432,633,1854]" captionTargetId="figure-127@37.[151,1436,633,1858]" captionTargetPageId="37" captionText="FIGURE 28. Habitus of Otacilia impleta sp. nov.: A. Male holotype, dorsal view; B. Same, ventral view; C. Female paratype, dorsal view; D. Same, ventral view." figureDoi="http://doi.org/10.5281/zenodo.8283835" httpUri="https://zenodo.org/record/8283835/files/figure.png" pageId="40" pageNumber="41">Fig. 28C, D</figureCitation>
).
</paragraph>
<paragraph id="BE5836C8865BFFA15AF6FC38DDDB2467" blockId="40.[151,1437,150,1149]" pageId="40" pageNumber="41">
Epigyne as in
<figureCitation id="26DC2A4D865BFFA15B58FC38D9A023D3" box="[361,493,979,1005]" captionStart="FIGURE 29" captionStartId="38.[151,250,1850,1874]" captionTargetBox="[151,1436,188,1815]" captionTargetId="figure-21@38.[151,1436,181,1826]" captionTargetPageId="38" captionText="FIGURE 29. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, prolateral view; B. Same, retrolateral view; C. Same, ventral view; D. Same, dorsal view; E. Epigyne, ventral view; F. Same, dorsal view. Abbreviations: E—embolus; C—conductor; DTA—dorsal tibial apophysis; FA—femoral apophysis; PTA—prolateral tibial apophysis; RTA—retrolateral tibial apophysis; SD—sperm duct; TA—tegular apophysis; B—bursa; CO—copulatory opening; CD—copulatory duct; CT— connecting tube; FD—fertilization duct; GA—glandular appendage; S—spermathecae." figureDoi="http://doi.org/10.5281/zenodo.8283839" httpUri="https://zenodo.org/record/8283839/files/figure.png" pageId="40" pageNumber="41">Figs 29EF</figureCitation>
,
<figureCitation id="26DC2A4D865BFFA15BC8FC38DA0123D3" box="[505,588,979,1005]" captionStart="FIGURE 30" captionStartId="39.[151,250,1514,1538]" captionTargetBox="[156,1435,186,1485]" captionTargetId="figure-17@39.[151,1435,181,1490]" captionTargetPageId="39" captionText="FIGURE 30. Copulatory organs of Otacilia impleta sp. nov.: A. Male left palp, ventral view; B. Same, retrolateral view; C. Same, dorsal view; D. Epigyne, ventral view; E. Same, dorsal view. Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8283843" httpUri="https://zenodo.org/record/8283843/files/figure.png" pageId="40" pageNumber="41">30DE</figureCitation>
. Epigynal plate weakly sclerotized, posterior edge straight. Median septum absent. Copulatory openings round, located at middle of plate. Copulatory ducts not visible. Glandular appendages short, digitiform. Connecting tubes V-shaped, shorter than fertilization duct. Bursa large, transparent, balloon-shaped. Spermathecae globular, close to each other. Fertilization ducts located at anterior margin of spermathecae.
</paragraph>
</subSubSection>
<subSubSection id="F6FD6543865BFFA15AF6FB89DAC12443" box="[199,652,1122,1149]" pageId="40" pageNumber="41" type="distribution">
<paragraph id="BE5836C8865BFFA15AF6FB89DAC12443" blockId="40.[151,1437,150,1149]" box="[199,652,1122,1149]" pageId="40" pageNumber="41">
<emphasis id="8C93EADA865BFFA15AF6FB89D92C2442" bold="true" box="[199,353,1122,1148]" pageId="40" pageNumber="41">Distribution.</emphasis>
<collectingCountry id="C6F07658865BFFA15B56FB88D9E72443" box="[359,426,1123,1149]" name="China" pageId="40" pageNumber="41">China</collectingCountry>
(
<collectingRegion id="7C23F82A865BFFA15B88FB88DACD2443" box="[441,640,1123,1149]" country="China" name="Yunnan" pageId="40" pageNumber="41">Yunnan Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>