treatments-xml/data/2B/FF/34/2BFF343250245B6EB7C5A338BDB4ED8A.xml

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<mods:title id="F2A0B00D2030F9633C371C8B561E8A1A">Island hoppers: Integrative taxonomic revision of Hogna wolf spiders (Araneae, Lycosidae) endemic to the Madeira islands with description of a new species</mods:title>
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<mods:namePart id="AA23096E0D0BE972475EF05431386BA3">Crespo, Luis C.</mods:namePart>
<mods:nameIdentifier id="E0A2093A9714AD751F367154A893DD50" type="ORCID">https://orcid.org/0000-0002-5388-5661</mods:nameIdentifier>
<mods:affiliation id="6237395739E65C196BF5F1B19782AE7B">Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), Universitat de Barcelona, Av. Diagonal 643, 08028 Barcelona, Spain &amp; Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History (LUOMUS), University of Helsinki, P. O. Box 17, 00014 Helsinki, Finland</mods:affiliation>
<mods:nameIdentifier id="8EDF5C6035D4876B15739A0D16F982B4" type="email">luiscarloscrespo@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5B49069842D2E916F6E74EE6046A9C3E">Silva, Isamberto</mods:namePart>
<mods:affiliation id="053416C5F873BB2E9C1FEB75D171E7B5">Instituto das Florestas e Conservacao da Natureza IP-RAM, Jardim Botanico da Madeira, Caminho do Meio, Bom Sucesso, 9064 - 512, Funchal, Portugal</mods:affiliation>
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<mods:namePart id="DA9E1F49C29752A2E89D0D72C7A41C58">Enguidanos, Alba</mods:namePart>
<mods:affiliation id="E1D7E13B831414229FAC67D528CFBCAE">Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), Universitat de Barcelona, Av. Diagonal 643, 08028 Barcelona, Spain</mods:affiliation>
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<mods:namePart id="31351F5B20D0962F0C9BD0F9BC585BC6">Cardoso, Pedro</mods:namePart>
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<mods:affiliation id="20DBDFF16C436B0E2C7540DC899461BA">Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History (LUOMUS), University of Helsinki, P. O. Box 17, 00014 Helsinki, Finland</mods:affiliation>
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<mods:namePart id="D49F034727B589BD60ED5EBAD62EFDD8">Arnedo, Miquel</mods:namePart>
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<mods:affiliation id="29E642B4DA2B7C040E54D7AA38C7944A">Department of Evolutionary Biology, Ecology and Environmental Sciences (Arthropods), Biodiversity Research Institute (IRBio), Universitat de Barcelona, Av. Diagonal 643, 08028 Barcelona, Spain</mods:affiliation>
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<mods:date id="E46989CE107968B61F01F9D03C5D16D1">2022</mods:date>
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<treatment id="2BFF343250245B6EB7C5A338BDB4ED8A" ID-GBIF-Taxon="193383534" LSID="urn:lsid:plazi:treatment:2BFF343250245B6EB7C5A338BDB4ED8A" httpUri="http://treatment.plazi.org/id/2BFF343250245B6EB7C5A338BDB4ED8A" lastPageNumber="84" pageId="0" pageNumber="84" scope_family="Lycosidae" scope_order="Araneae">
<subSubSection id="24535AD11344324A2CECAA11C20E1C20" pageId="0" pageNumber="84" type="nomenclature">
<paragraph id="C94AB9009D7B46F664235691005294E6" pageId="0" pageNumber="84">
<taxonomicName id="CED1202E2723722DB0BF9A219703292F" ID-CoL="3M6JG" LSID="2BFF3432-5024-5B6E-B7C5-A338BDB4ED8A" authority="(Thorell, 1875)" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">Hogna heeri (Thorell, 1875)</taxonomicName>
</paragraph>
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<subSubSection id="16D4C8E2A0F292D7681B583F128497A6" pageId="0" pageNumber="84" type="description">
<paragraph id="F42F704DE0FD2DD6D0BF7B253C78271C" pageId="0" pageNumber="84">
<figureCitation id="8BFBB7E69565C794012A493FA782F0C2" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">Figures 9</figureCitation>
<figureCitation id="44C4A4C54A9C388CE9FC9D69CC853601" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Photograph of H. heeri. Female specimen in captivity. Photograph credit Emidio Machado." figureDoi="10.3897/zookeys.1086.68015.figure10" httpUri="https://binary.pensoft.net/fig/646990" pageId="0" pageNumber="84">, 10</figureCitation>
<figureCitation id="6172D36531A68A040C46C42ADB5B60DD" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of H. heeri. Circles: present records; black triangles: revised records from literature; white triangles: unconfirmed records from literature." figureDoi="10.3897/zookeys.1086.68015.figure11" httpUri="https://binary.pensoft.net/fig/646991" pageId="0" pageNumber="84">, 11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F7FC76D96B5E2F4C483D41826A44A34F" pageId="0" pageNumber="84" type="reference_group">
<paragraph id="57CDAABD0755592571D2D02F3601739D" pageId="0" pageNumber="84">
<taxonomicName id="6D0FA47D2ED6D5ECAB5FA16EC9D6D952" authorityName="Thorell" authorityYear="1875" class="Arachnida" family="Lycosidae" genus="Trochosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochosa herii" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="herii">Trochosa herii</taxonomicName>
Thorell, 1875: 166 (Df).
</paragraph>
<paragraph id="894830E34FB0FEF8690233EB4A8270CD" pageId="0" pageNumber="84">
<taxonomicName id="E7CF23D02F3AF02A93A5F559BB3F2815" authorityName="Thorell" authorityYear="1875" class="Arachnida" family="Lycosidae" genus="Trochosa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trochosa herii" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="herii">Trochosa herii</taxonomicName>
Kulczynski, 1899: 433, pl. 9, fig. 188 (f).
</paragraph>
<paragraph id="9EEFDF50DDF2EC37EDDAF51A283F85DC" pageId="0" pageNumber="84">
<taxonomicName id="3AD76E874FF0BA2AC0BD521ACA659CA8" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">Hogna heeri</taxonomicName>
Roewer, 1955: 248.
</paragraph>
<paragraph id="BF5567124CF32F5CD77257DBAE90B3D8" pageId="0" pageNumber="84">
<taxonomicName id="9222863BC3E398A89B6D9EFFBE7AC4D8" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna herii" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="herii">Hogna herii</taxonomicName>
Roewer, 1959: 411, fig. 221a-d (f, Dm).
</paragraph>
<paragraph id="ADD5B118B3E4FC141AB03233780A2CD8" pageId="0" pageNumber="84">
<taxonomicName id="D8C42D5F9C050D494D6663591DA1849B" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">Hogna heeri</taxonomicName>
Wunderlich, 1992: 459, fig. 720, 720a (mf).
</paragraph>
</subSubSection>
<subSubSection id="003DA6F9A059EA126919377132A6666D" pageId="0" pageNumber="84" type="types">
<paragraph id="32DD3F4C3F8C36BCA854552AA74B3DCC" pageId="0" pageNumber="84">Types.</paragraph>
<paragraph id="2277D4CAE1B6F45938762BC30EE7234F" pageId="0" pageNumber="84">
<emphasis id="078EE6408F2E3CBD7E461BFA6293E62F" bold="true" italics="true" pageId="0" pageNumber="84">Syntypes</emphasis>
: Madeira • 2 ♀♀, leg. O. Heer, stored at NHRS, collection number JUST-000001113. Examined.
</paragraph>
</subSubSection>
<subSubSection id="588AD5452D3865CFFDEF84CCA79E5CE5" pageId="0" pageNumber="84" type="materials_examined">
<paragraph id="A6807634B4EB84484A94657E82585BFD" pageId="0" pageNumber="84">Material examined.</paragraph>
<paragraph id="7F6CE208297F7BE8862655D527D5D2F1" pageId="0" pageNumber="84">
<materialsCitation id="D2F497AFEA108EA9D885850BEEE9E4B2" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" latitude="32.41228" location="Bugio" longLatPrecision="1" longitude="-16.47466" municipality="Planalto Sul" specimenCount="1" specimenCount-female="1">
<location id="66E076A1B36B0918A384319F7499D4A2" LSID="urn:lsid:plazi:treatment:2BFF343250245B6EB7C5A338BDB4ED8A:66E076A1B36B0918A384319F7499D4A2" latitude="32.41228" longLatPrecision="1" longitude="-16.47466" municipality="Planalto Sul" name="Bugio">Bugio</location>
• 
<collectingMunicipality id="ACBF884708C2D68BF7C492A57046F7A8">Planalto Sul</collectingMunicipality>
, 
<geoCoordinate id="6742B5D4B21C270938CC3484E8C624E4" degrees="32.41228" direction="north" orientation="latitude" precision="1" value="32.41228">32.41228°N</geoCoordinate>
, 
<geoCoordinate id="0B3C4F3FFC82A8E4691D67DDA488A8FC" degrees="16.47466" direction="west" orientation="longitude" precision="1" value="-16.47466">16.47466°W</geoCoordinate>
, 
<specimenCount id="B4DB6D6E9DEB717EE0C1DE8AA2D306B8" type="female">1 ♀</specimenCount>
(LCPC)
</materialsCitation>
, 
<materialsCitation id="6384896B57348A4DB822AA41E8EF64F0" accessionNumber="SMF69107" collectingDate="2012-12-03" collectorName="I. Silva. Madeira, Freiras" location="Paul da Serra" specimenCount="3" specimenCount-female="3">
<collectingDate id="3BEB973DEBFE06F25BCB002240B018B7" value="2012-12-03">3.XII.2012</collectingDate>
, hand collecting, leg. 
<collectorName id="9EAA887B7A990DB116BA690F0074406F">I. Silva. Madeira</collectorName>
• 
<locationDeviation id="3F310E1D3D082FA3D41957566D3D5936" firstLocation="Eira do Serrado" secondLocation="Curral">between Eira do Serrado and Curral</locationDeviation>
das 
<collectorName id="38FC311ACF0FDD8ED960C0B7978F15AF">Freiras</collectorName>
, 
<specimenCount id="794D3F2F975B61EDD6E00F108CB1A533" type="female">1 ♀</specimenCount>
(
<accessionNumber id="A65C5DFE5183FDE135B5437FEA6F7A0D" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF69107">SMF69107</accessionNumber>
); 
<location id="CB0691C4166E65B96481B35E2DC833B5" LSID="urn:lsid:plazi:treatment:2BFF343250245B6EB7C5A338BDB4ED8A:CB0691C4166E65B96481B35E2DC833B5" name="Paul da Serra">
<normalizedToken id="2EC7C031CB96D77F74C11E358ADA9371" originalValue="Paúl">Paul</normalizedToken>
da Serra
</location>
, 
<specimenCount id="43FD88EC95A651B049AF684F9C94939F" type="female">2 ♀♀</specimenCount>
(MMUE G7572.874)
</materialsCitation>
, 
<date id="AD13313B700A1BD81B391148A2690B4A" value="1973-04-25">25.IV.1973</date>
, leg. J. Murphy, 
<specimenCount id="367F5534049BDC7025B4C2C88CC869AF" type="female">1 ♀</specimenCount>
(CRBALC0492: LC289), 
<geoCoordinate id="F88D0FCB5C044E53737E72E7EDCBBD3C" degrees="32.78182" direction="north" orientation="latitude" precision="1" value="32.78182">32.78182°N</geoCoordinate>
, 
<geoCoordinate id="F5527A0C12111824881D23479589D480" degrees="17.09978" direction="west" orientation="longitude" precision="1" value="-17.09978">17.09978°W</geoCoordinate>
, 
<date id="A942EF00C1A028DA3B439EF9D85F389C" value="2017-03-19">19.III.2017</date>
, hand collecting, leg. I. Silva, 
<specimenCount id="E69E8C187AF775BFA5C32F15A990E341" type="female">1 ♀</specimenCount>
(CRBALC0500: LC222) and 
<specimenCount id="A90103726D43BE5253C5B2CFC82344D4" type="juvenile">1 juvenile</specimenCount>
(CRBALC0494: LC291), 
<materialsCitation id="67F4887BCE3BE8DA58E5912CC92C5412" collectingDate="2017-03-28" collectorName="I. Silva" latitude="32.74036" location="Pico do Cidrao" longLatPrecision="1" longitude="-16.93877" specimenCount="1" specimenCount-female="1">
<collectingDate id="9DB32F786B0F34E58F6285B95CDA46BB" value="2017-03-28">28.III.2017</collectingDate>
, leg. 
<collectorName id="1439747FE8E307CAFA16CBF4F1898C58">I. Silva</collectorName>
; 
<location id="E9827648877E7094820E0F4BE5C67546" LSID="urn:lsid:plazi:treatment:2BFF343250245B6EB7C5A338BDB4ED8A:E9827648877E7094820E0F4BE5C67546" latitude="32.74036" longLatPrecision="1" longitude="-16.93877" name="Pico do Cidrao">
Pico do 
<normalizedToken id="8321063C561F44ED93FF1C2438FD3637" originalValue="Cidrão">Cidrao</normalizedToken>
</location>
, 
<geoCoordinate id="78646A20846C2E955FCFD9E3161B4299" degrees="32.74036" direction="north" orientation="latitude" precision="1" value="32.74036">32.74036°N</geoCoordinate>
, 
<geoCoordinate id="5A2B7CFF5CA5F870445F7B13F19EC03B" degrees="16.93877" direction="west" orientation="longitude" precision="1" value="-16.93877">16.93877°W</geoCoordinate>
, 
<specimenCount id="CDCB01414CF7B25D5DBE4C6CDA0A89CA" type="female">1 ♀</specimenCount>
(LCPC)
</materialsCitation>
, 
<date id="CC38527BC1A358742946E4D5F673A166" value="2003-06-24">24.VI.2003</date>
, pitfall trapping, leg. M. Freitas, 
<specimenCount id="E5C22AB4BCA3D7AA0A8D173F4FC012AD" type="female">2 ♀♀</specimenCount>
(CRBALC0490: LC287, CRBALC0288: LC288), 
<materialsCitation id="6903FF449D48D86FC46B1B7FE601DFDF" collectingDate="2017-03-27" collectorName="L. Crespo, I. Silva" county="Pessegueiros" latitude="32.78837" location="Paul da Serra" longLatPrecision="1" longitude="-17.09857" municipality="Montado" specimenCount="2" specimenCount-female="2">
<collectingDate id="CA4531470362BC28E5BED33D4ED131B0" value="2017-03-27">27.III.2017</collectingDate>
, hand collecting, leg. 
<collectorName id="9E6631A70FCD9897A681857B0ECFFC5D">L. Crespo</collectorName>
&amp; 
<collectorName id="0AD39C4D78EC6D4BAA823CB71896D25C">I. Silva</collectorName>
; trail from 
<location id="3076BDF318F01E2CC81176634042B398" LSID="urn:lsid:plazi:treatment:2BFF343250245B6EB7C5A338BDB4ED8A:3076BDF318F01E2CC81176634042B398" county="Pessegueiros" latitude="32.78837" longLatPrecision="1" longitude="-17.09857" municipality="Montado" name="Paul da Serra">
<normalizedToken id="5E39130CEC78E03BDFA4CA7AA898A356" originalValue="Paúl">Paul</normalizedToken>
da Serra
</location>
to 
<collectingMunicipality id="291F3DD902AEDA89CAFFED18FAF3AA8C">Montado</collectingMunicipality>
dos 
<collectingCounty id="99235641C17E8DBC35DABA886D5AF398">Pessegueiros</collectingCounty>
, 
<geoCoordinate id="CF3AFA404D07EABA300B12D71600D5C7" degrees="32.78837" direction="north" orientation="latitude" precision="1" value="32.78837">32.78837°N</geoCoordinate>
, 
<geoCoordinate id="2B7B1E3A65F2ECDF7BF9DD799FDAAC79" degrees="17.09857" direction="west" orientation="longitude" precision="1" value="-17.09857">17.09857°W</geoCoordinate>
, 
<specimenCount id="C290E19918350B3E3A54A40237BD56DD" type="female">2 ♀♀</specimenCount>
(CRBALC0270: LC184, CRBALC0501: LC223)
</materialsCitation>
and 
<specimenCount id="C923C14FAE85EFADC682FD98C94E5280" type="juvenile">1 juvenile</specimenCount>
(CRBALC0493: LC290), 
<materialsCitation id="D22C65FDB3922F402513F8660F7F7DD9" accessionNumber="SMF37575" collectingDate="2017-03-28" collectorName="L. Crespo, I. Silva" specimenCount="1" specimenCount-female="1">
<collectingDate id="C4003FED17DD73E1FDF5E5D5B7B46F71" value="2017-03-28">28.III.2017</collectingDate>
, hand collecting, leg. 
<collectorName id="DC4D38A35071BCFE9BF9438074D024D0">L. Crespo</collectorName>
&amp; 
<collectorName id="17F372A25C69B3C8D4A5240C330F0214">I. Silva</collectorName>
; 
<specimenCount id="595C4370AFB5467B45D65CECD4CC0698" type="female">1 ♀</specimenCount>
(
<accessionNumber id="918879402B659C4F38C1E2D2EAC47144" httpUri="http://www.ncbi.nlm.nih.gov/protein/SMF37575">SMF37575</accessionNumber>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="019D1E28FC26391377181CD66E5D32CE" pageId="0" pageNumber="84" type="diagnosis">
<paragraph id="13B1A157DE3A245207C3F3A7DFA3C32A" pageId="0" pageNumber="84">Diagnosis.</paragraph>
<paragraph id="F428B3BB583C49BF322F43756415BF53" pageId="0" pageNumber="84">
<taxonomicName id="BE673C3AEBF899A40C85C040C158EAF9" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="8DD36851A12E1303A66E3E8410F45D3C" italics="true" pageId="0" pageNumber="84">Hogna heeri</emphasis>
</taxonomicName>
can be diagnosed by the genitalia: the males, according to literature, by a straight embolus (
<bibRefCitation id="C99B85AB63AB9AD68C761A05E1C17E55" author="Wunderlich, J" journalOrPublisher="Beitraege zur Araneologie" pageId="0" pageNumber="84" pagination="1 - 619" refId="B76" refString="Wunderlich, J, 1992. Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Oekologie, Biogeographie und Evolution. Beitraege zur Araneologie 1: 1 - 619" title="Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Oekologie, Biogeographie und Evolution." volume="1" year="1992">Wunderlich 1992</bibRefCitation>
: 595, fig. 720); in females, by epigynal anterior pockets with widely divergent lateral border and median septum with a wide posterior transverse part (Fig. 
<figureCitation id="3E7661A778F2844C0EBE841A309708A5" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">9</figureCitation>
). Similar species include 
<taxonomicName id="C3A0BD4F7F880E79A0A8C843B078FFB7" class="Arachnida" kingdom="Animalia" lsidName="H. insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis id="39D324D0228B06BB5B675FF1EC67AE2F" italics="true" pageId="0" pageNumber="84">H. insularum</emphasis>
</taxonomicName>
and 
<taxonomicName id="02AA30F39B4C3A9512E355BBDB69AF0B" class="Arachnida" kingdom="Animalia" lsidName="H. isambertoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="isambertoi">
<emphasis id="FADE485EEDF903AF95064AA38B87CA71" italics="true" pageId="0" pageNumber="84">H. isambertoi</emphasis>
</taxonomicName>
sp. nov., from which it cannot be somatically differentiated.
</paragraph>
<caption id="0493075FFD64DE4D533FE592DD0E69D4" doi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84" start="Figure 9" startId="F9">
<paragraph id="9EE19C4D1EB9FB8316B611C600553521" pageId="0" pageNumber="84">
<emphasis id="5BAE55B7EB6375E43082B03AE4F3AAD1" bold="true" pageId="0" pageNumber="84">Figure 9.</emphasis>
<taxonomicName id="B05C3C7DEF80D0C6688788BDC2942D66" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="F994AF21065ECD751791EFB633CA1298" italics="true" pageId="0" pageNumber="84">Hogna heeri</emphasis>
</taxonomicName>
female (CRBALC0501): 
<emphasis id="E57AA61E5AADC58460745B64F308593D" bold="true" pageId="0" pageNumber="84">A</emphasis>
epigyne, ventral 
<emphasis id="C95F7F0A009E34F637264E94F5CB96D3" bold="true" pageId="0" pageNumber="84">B</emphasis>
vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="FF88BDA1B13AB9CBA993298BBB614965" pageId="0" pageNumber="84" type="redescription">
<paragraph id="44C4980ED77D6E8C158EDCC99756B15F" pageId="0" pageNumber="84">Redescription.</paragraph>
<paragraph id="F6E549DF072C940CE1576CA6890A1013" pageId="0" pageNumber="84">
<emphasis id="5778CE5A2A3D7A9F24F0FEAADDA69324" bold="true" pageId="0" pageNumber="84">Male</emphasis>
: We could not examine any male specimens.
</paragraph>
<paragraph id="CE2400A22F9602C629B2EE6EFEED8FF0" pageId="0" pageNumber="84">
<emphasis id="0FE8A3ED30B609B6867106EDFA880EFA" bold="true" pageId="0" pageNumber="84">Female</emphasis>
(
<emphasis id="C4A4155971FF875356B9F0695FB0EC8A" italics="true" pageId="0" pageNumber="84">CRBALC0500</emphasis>
): (Fig. 
<figureCitation id="40A5502297A57C708C03FE5B2B5780FA" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Photograph of H. blackwalli. Female specimen, recently dead, in captivity. Photograph credit Emidio Machado." figureDoi="10.3897/zookeys.1086.68015.figure7" httpUri="https://binary.pensoft.net/fig/646987" pageId="0" pageNumber="84">7</figureCitation>
corresponds to specimen CRBALC0501). Total length 13.54; carapace: 5.63 long, 4.4 wide.
</paragraph>
<paragraph id="2C237C52E78A117B9FE5CC0E9D30D86E" pageId="0" pageNumber="84">
<emphasis id="7EDC772076816EE5FEA181CEFA42FEF6" bold="true" italics="true" pageId="0" pageNumber="84">Colour</emphasis>
: carapace greyish brown, covered with short black setae, with a median cream longitudinal band, anteriorly broadened, covered with short white setae, with suffused greyish brown patches; two yellow marginal bands, with roughly round grey patches, covered with short white setae; four black striae well visible on each flank. Chelicerae dark brown, covered in black and yellow setae. Gnathocoxae and labium overall brown, with posterior margin blackish; sternum yellow, with a faint V-shaped grey patch and grey lateral borders. Legs yellow, with irregular grey suffused patches, except metatarsi and tarsi, brown. Pedipalps yellow except tibia, brown, tarsus, blackish brown. Abdomen with a pair of anterolateral black patches, extending laterally into grey flanks, mottled with yellowish patches covered with white setae; a median dark lanceolate patch is bordered by two yellowish longitudinal bands interconnected in anterior half, posteriorly by means of dark chevrons; venter yellowish, with a median dark grey longitudinal band, bordered by yellowish and grey small patches.
</paragraph>
<paragraph id="E94BD976AB7B1958B15167ACA3FCDF8E" pageId="0" pageNumber="84">
<emphasis id="8F7D76B39FFB48344FBB1E9652DF4D1B" bold="true" italics="true" pageId="0" pageNumber="84">Eyes</emphasis>
: MOQ: MW = 0.7 PW, MW = 1.1 LMP, MW = 1.1 AW; Cl = 0.9 DAME. Anterior eye row straight.
</paragraph>
<paragraph id="AB13C1B2888032539B1914F61274C39C" pageId="0" pageNumber="84">
<emphasis id="1B21AFCF2D06C59EE8B290F19FFB9772" bold="true" italics="true" pageId="0" pageNumber="84">Legs</emphasis>
: Measurements: Leg I: 13.0, TiI: 2.8; Leg IV: 16.10, TiIV: 3.22; TiIL/D: 3.7. Spination of Leg I: FeI: d1.1.1, p0.0.1; TiI: v2l.2l.2s; MtI: p0.0.1, r0.0.1, v2l.2l.1s. MtI with sparse scopulae in basal half and dense scopulae on distal half.
</paragraph>
<paragraph id="B9E4D69EFD65D9B3D77D7E66CCC09F9C" pageId="0" pageNumber="84">
<emphasis id="F5322D4843CD284271AECB9ECEB88009" bold="true" italics="true" pageId="0" pageNumber="84">Epigyne</emphasis>
: anterior pockets touching, short, with lateral borders widely divergent, converging solely at its posterior end (Fig. 
<figureCitation id="107267482E311DB18EBB03D6062B7E74" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">9A</figureCitation>
); anterior pocket cavities deep; median septum with wide posterior transverse part (Fig. 
<figureCitation id="EC32777E30A39390974DF52712059A3A" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">9A</figureCitation>
); spermathecae globular (Fig. 
<figureCitation id="A6E7809575B17E123695605831355BB1" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">9B</figureCitation>
); copulatory ducts basally with a laterally projected diverticulum (Fig. 
<figureCitation id="CC92ACA99C0901BE96F0233E4F693B7E" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">9B</figureCitation>
); fertilisation ducts emerging at the base of copulatory duct (Fig. 
<figureCitation id="A9E4CE1A3088CAF40A266C65CCF2F003" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Hogna heeri female (CRBALC 0501): A epigyne, ventral B vulva, dorsal. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.2 mm." figureDoi="10.3897/zookeys.1086.68015.figure9" httpUri="https://binary.pensoft.net/fig/646989" pageId="0" pageNumber="84">9B</figureCitation>
).
</paragraph>
<caption id="33FDCB1992B35FD0F4D4DB8E9FFBC4D2" doi="10.3897/zookeys.1086.68015.figure10" httpUri="https://binary.pensoft.net/fig/646990" pageId="0" pageNumber="84" start="Figure 10" startId="F10">
<paragraph id="00AA5EEED33E961C48D2CA0E5B998090" pageId="0" pageNumber="84">
<emphasis id="46CA8F74A484A4CF729AC17FEC911C02" bold="true" pageId="0" pageNumber="84">Figure 10.</emphasis>
Photograph of 
<taxonomicName id="11D212A90A0895364A857496E87D99C6" class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="EBC4666AC0877AEC155BCC5E2DBC5C7B" italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
. Female specimen in captivity. Photograph credit 
<normalizedToken id="82EB4C20BE7F1B763D66B243C274C3FB" originalValue="Emídio">Emidio</normalizedToken>
Machado.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DE1ECF8FDFE612ECD1F821E705CC7D5E" pageId="0" pageNumber="84" type="intraspecific variation">
<paragraph id="D724DE60797D72E67A75A9E9AD99C782" pageId="0" pageNumber="84">Intraspecific variation.</paragraph>
<paragraph id="A49D1DEA93B4E3AD5215A2398DF989F4" pageId="0" pageNumber="84">Carapace length, females: 5.6-5.8. In females, the ventral abdominal dark band may be entirely absent; the relative position of female epigynal anterior pockets may vary from touching to almost touching.</paragraph>
</subSubSection>
<subSubSection id="A4B0CB160D896A7E08BD7A154F2DA9AA" pageId="0" pageNumber="84" type="distribution">
<paragraph id="5133190533434EEF92C596789598AB5C" pageId="0" pageNumber="84">Distribution.</paragraph>
<paragraph id="5181AC3E74847400F7D9F40A6E992A1B" pageId="0" pageNumber="84">
This species is known from two distinct regions: high altitude localities in Madeira, always above 800 m, and the island of Bugio (Fig. 
<figureCitation id="FCF364BDD14BA9DB1340B6DD74085E54" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of H. heeri. Circles: present records; black triangles: revised records from literature; white triangles: unconfirmed records from literature." figureDoi="10.3897/zookeys.1086.68015.figure11" httpUri="https://binary.pensoft.net/fig/646991" pageId="0" pageNumber="84">11</figureCitation>
).
</paragraph>
<caption id="76A7E8B1CB9F59F6960A7FCC2070624E" doi="10.3897/zookeys.1086.68015.figure11" httpUri="https://binary.pensoft.net/fig/646991" pageId="0" pageNumber="84" start="Figure 11" startId="F11">
<paragraph id="EB8087A89FD2BF28CC4ABB792672CFCE" pageId="0" pageNumber="84">
<emphasis id="98A9EA76B434C9B815A325FCEFD169D9" bold="true" pageId="0" pageNumber="84">Figure 11.</emphasis>
Distribution of 
<taxonomicName id="AE4343FAD0027D6C15CFFFCDD644E7B7" class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="6CA70C68603714C8A6B5E630935558DF" italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
. Circles: present records; black triangles: revised records from literature; white triangles: unconfirmed records from literature.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EBE609AE3E958E91E185D6710A727B92" pageId="0" pageNumber="84" type="ecology">
<paragraph id="D364061C1FAE0404BAD92BD7F32EB6D0" pageId="0" pageNumber="84">Ecology.</paragraph>
<paragraph id="246D8320C5613939CB819561CF85451C" pageId="0" pageNumber="84">
<taxonomicName id="1F9E8C58620D6D1908680F4CF33ED361" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="1636C122F251625A8743BE6F21CF03F3" italics="true" pageId="0" pageNumber="84">Hogna heeri</emphasis>
</taxonomicName>
occurs in montane grasslands or 
<taxonomicName id="66345300E0D333DFB595803ADA07BDB5" class="Magnoliopsida" family="Ericaceae" genus="Erica" kingdom="Plantae" lsidName="Erica" order="Ericales" pageId="0" pageNumber="84" phylum="Tracheophyta" rank="genus">
<emphasis id="F55B91F855A12203B8C3637D7D7AEF96" italics="true" pageId="0" pageNumber="84">Erica</emphasis>
</taxonomicName>
shrubland in Madeira and the steep, semi-arid summit of Bugio.
</paragraph>
</subSubSection>
<subSubSection id="0D0010858D72DD55A19DAB2A5F6A95DC" pageId="0" pageNumber="84" type="conservation status">
<paragraph id="74AC6A02F5E195F8822360CE01E9F061" pageId="0" pageNumber="84">Conservation status.</paragraph>
<paragraph id="2530ABD8C3ABCADF0F9183F56DAC72E5" pageId="0" pageNumber="84">
<taxonomicName id="499EE6D2174B647F1A1834E55AED963C" baseAuthorityName="Thorell" baseAuthorityYear="1875" class="Arachnida" family="Lycosidae" genus="Hogna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hogna heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="9463F4808E3F0880C3E0AF635028BFF6" italics="true" pageId="0" pageNumber="84">Hogna heeri</emphasis>
</taxonomicName>
was assessed according to the IUCN Red List criteria, with the status of Least Concern (
<bibRefCitation id="381D7D1AE190355FF21316DF98456275" DOI="https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T58048559A58061002.en" author="Cardoso, P" journalOrPublisher="Molecular Ecology" pageId="0" pageNumber="84" refId="B16" refString="Cardoso, P, Crespo, LC, Silva, I, Borges, P, Boieiro, M, 2018b. Hogna heeri. The IUCN Red List of Threatened Species 2018: 8. https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T58048559A58061002.en" title="Hogna heeri. The IUCN Red List of Threatened Species 2018: 8." url="https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T58048559A58061002.en" year="2018 b">Cardoso et al 2018b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="60C783D1F2EF41A4BC0010289A793DB1" pageId="0" pageNumber="84" type="comments">
<paragraph id="4B1117A8BC50B25BC1399D4739729F29" pageId="0" pageNumber="84">Comments.</paragraph>
<paragraph id="5F77DA9A206CFBE4A185465CB39F87B3" pageId="0" pageNumber="84">
The specific epithet of 
<taxonomicName id="43619E78A65DB6B62B07E6049EFA1189" class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="BCCC52DDF2AEA12137DC5CA25FE1EDEE" italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
has been one of the names renamed by 
<bibRefCitation id="60137DEF49C05E98B503382BFCB0D1F5" author="Bonnet, P" journalOrPublisher="Douladoure, Toulouse" pageId="0" pageNumber="84" refId="B8" refString="Bonnet, P, 1959. Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome II. Systematique des araignees (Etude par ordre alphabetique) (5eme partie: T-Z). Douladoure, Toulouse, 4231-5058." title="Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome II. Systematique des araignees (Etude par ordre alphabetique) (5 eme partie: T-Z). Douladoure, Toulouse, 4231 - 5058." year="1959">Bonnet (1959)</bibRefCitation>
, who changed all previously described spider 
<normalizedToken id="C6C2DD234E40ECCEF679200B077270A4" originalValue="species’">species'</normalizedToken>
names which were patronyms ending in 
<normalizedToken id="E5C444B818AC6D99F8B8B027D35389F2" originalValue="“ii”">&quot;ii&quot;</normalizedToken>
to end in 
<normalizedToken id="F2354E96BDFD391BC4FEA01A16547CF7" originalValue="“i”">&quot;i&quot;</normalizedToken>
, as a way to correct spelling (
<bibRefCitation id="31480A79340AAB792CED2718ED448E0C" author="Bonnet, P" journalOrPublisher="Douladoure, Toulouse" pageId="0" pageNumber="84" refId="B7" refString="Bonnet, P, 1945. Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome I. Douladoure, Toulouse" title="Bibliographia araneorum. Analyse methodique de toute la litterature araneologique jusqu'en 1939. Tome I." year="1945">Bonnet 1945</bibRefCitation>
). Although the ICZN argues for the maintenance of the original spelling, common usage dictates that these modified spellings continue to be used. The disjunct distribution of 
<taxonomicName id="1139732A74D388BCFAC7D1BF1B66782F" class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="00CEC8D0A6A89D3009E51BEDC9022D08" italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
, with populations in Madeira and Bugio, is somewhat baffling. The only known specimens from Bugio previously reported (
<bibRefCitation id="085C228443B3D1B09657C2AA9E6DFF3D" author="Crespo, L" journalOrPublisher="Revista Iberica de Aracnologia" pageId="0" pageNumber="84" pagination="11 - 23" refId="B24" refString="Crespo, L, Silva, I, Borges, P, Cardoso, P, 2013. Rapid biodiversity assessment, faunistics and description of a new spider species (Araneae) from Desertas Islands and Madeira (Portugal). Revista Iberica de Aracnologia 23: 11 - 23" title="Rapid biodiversity assessment, faunistics and description of a new spider species (Araneae) from Desertas Islands and Madeira (Portugal)." volume="23" year="2013">Crespo et al. 2013</bibRefCitation>
) were examined: while the female matches 
<taxonomicName id="EB7DBD8EF90FA05DBC57CAA0E64F0FE5" class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="544F3B18936D1A8D00DC228D3F6E329B" italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
, the male pedipalp is the same as that of 
<taxonomicName id="873FAB7A711909107B1B50963F3EA60B" class="Arachnida" kingdom="Animalia" lsidName="H. isambertoi" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="isambertoi">
<emphasis id="042A842A4EC7C004F94D37A594D6401D" italics="true" pageId="0" pageNumber="84">H. isambertoi</emphasis>
</taxonomicName>
sp. nov., with the tip of the embolus slightly tilted anteriorly (Fig. 
<figureCitation id="8BD963DBBDC1843D73C0BE00B4289EE4" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Hogna isambertoi sp. nov. A-C male (SMF): A left male palp, ventral B detail of the median apophysis, anteroventral C SEM image, right male palp, ventral D, E female (SMF): D epigynum, ventral E vulva, dorsal. Abbreviations, male palp: C - cymbium, E - embolus, MA - median apophysis, P - palea, T - tegulum, TA - terminal apophysis. Abbreviations, female genitalia: D - diverticulum, H - epigynal hoods, MS - median septum, S - spermatheca. Scale bars: 0.5 mm (A); 0.2 mm (B-E)." figureDoi="10.3897/zookeys.1086.68015.figure18" httpUri="https://binary.pensoft.net/fig/646998" pageId="0" pageNumber="84">18A</figureCitation>
). We would like to remark that Wunderlich reported an apophysis at the base of the embolus (indicated with an arrow in his figure) as a diagnostic feature to identify males of 
<taxonomicName id="B99C43E36ED1BA5C220F67E65107D314" class="Arachnida" kingdom="Animalia" lsidName="H. heeri" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="heeri">
<emphasis id="39BD2A89C4348D77158E90BE2F666DAF" italics="true" pageId="0" pageNumber="84">H. heeri</emphasis>
</taxonomicName>
(
<bibRefCitation id="2B9FA7308A9402C151EC35A06A7D54D6" author="Wunderlich, J" journalOrPublisher="Beitraege zur Araneologie" pageId="0" pageNumber="84" pagination="1 - 619" refId="B76" refString="Wunderlich, J, 1992. Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Oekologie, Biogeographie und Evolution. Beitraege zur Araneologie 1: 1 - 619" title="Die Spinnen-Fauna der Makaronesischen Inseln: Taxonomie, Oekologie, Biogeographie und Evolution." volume="1" year="1992">Wunderlich 1992</bibRefCitation>
: fig. 720), which appears to be either inconspicuous or missing altogether. To us it seems the arrow is pointing to the pars pendula membrane connecting the terminal apophysis with the embolus. Unfortunately, we could not gather molecular information from the Bugio specimens due to their poor preservation. Lastly, while revising 
<normalizedToken id="C4C3AB6950A0D7BF042D545896846F2B" originalValue="Thorell’s">Thorell's</normalizedToken>
type series, we identified one of the three adult females in the original vial as 
<taxonomicName id="AD5D998124672060707E0AAA1813DF24" class="Arachnida" kingdom="Animalia" lsidName="H. insularum" order="Araneae" pageId="0" pageNumber="84" phylum="Arthropoda" rank="species" species="insularum">
<emphasis id="FB2A2A16E02B1C9AC0E2590E45AD2382" italics="true" pageId="0" pageNumber="84">H. insularum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>