172 lines
19 KiB
XML
172 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.379.6457" ID-GBIF-Dataset="11de03c0-97c4-47b9-812b-b8e99890bb88" ID-PMC="PMC3935226" ID-Pensoft-Pub="1313-2970-379-1" ID-PubMed="24574855" ID-ZBK="237EB0D412AF485689C55E2AA52C4CEA" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-379-1" ModsDocOrigin="ZooKeys 379" ModsDocTitle="Revision of the genus Reichardtiolus Kryzhanovskij, 1959 (Coleoptera, Histeridae, Saprininae)" checkinTime="1451246330986" checkinUser="pensoft" docAuthor="Lackner, Tomas" docDate="2014" docId="2B358B1D24F170F253D497EE2F223B11" docLanguage="en" docName="ZooKeys 379: 1-27" docOrigin="ZooKeys 379" docSource="http://dx.doi.org/10.3897/zookeys.379.6457" docTitle="Reichardtiolus perses Lackner, 2014, sp. n." docType="treatment" docUuid="9B800BDD-A4B9-4D0B-BCE1-85CE9859E039" docUuidSource="ZooBank" docVersion="4" lastPageNumber="14" masterDocId="FFC4FF952E63FFB76559FFBF62144D69" masterDocTitle="Revision of the genus Reichardtiolus Kryzhanovskij, 1959 (Coleoptera, Histeridae, Saprininae)" masterLastPageNumber="27" masterPageNumber="1" pageNumber="11" updateTime="1668157689324" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genus Reichardtiolus Kryzhanovskij, 1959 (Coleoptera, Histeridae, Saprininae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lackner, Tomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>379</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>27</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.379.6457</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.379.6457</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-379-1</mods:identifier>
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<mods:identifier type="ZBK">237EB0D412AF485689C55E2AA52C4CEA</mods:identifier>
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<mods:identifier type="ZooBank">237EB0D412AF485689C55E2AA52C4CEA</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152050886" LSID="urn:lsid:zoobank.org:act:9B800BDD-A4B9-4D0B-BCE1-85CE9859E039" httpUri="http://treatment.plazi.org/id/2B358B1D24F170F253D497EE2F223B11" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
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<subSubSection pageId="10" pageNumber="11" type="nomenclature">
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName LSID="http://zoobank.org/9B800BDD-A4B9-4D0B-BCE1-85CE9859E039" class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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<taxonomicNameLabel pageId="10" pageNumber="11">sp. n.</taxonomicNameLabel>
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Figs 48-61
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="type locality">
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<paragraph pageId="10" pageNumber="11">Type locality.</paragraph>
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<paragraph pageId="10" pageNumber="11">Iran, Kerman, Talab.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="type material examined">
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<paragraph pageId="10" pageNumber="11">Type material examined.</paragraph>
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<paragraph pageId="10" pageNumber="11">
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Holotype, male, side-mounted on triangular point with male genitalia extracted and glued to the same triangular point as the specimen, left protarsus and left mid-leg missing, piece of left elytron from the elytral flank along the elytral base towards the fourth elytral stria chipped out; with the following labels:
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<normalizedToken originalValue="“♂”">"♂"</normalizedToken>
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(printed); followed by: "Kerman: str. Talab / 19-20.i.[19]01 / N. Zarudny" (printed-written label in Russian); followed by: "Coll. Semenov-Tian-Shansky" (printed); followed by: "ZOOLOGICAL / INSTITUTE RAS / ST. PETERSBURG" (yellow label, printed); followed by: "
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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/ sp.nov. HOLOTYPE / Det. T. Lackner 2013" (red label, printed) (ZIN). Paratypes: 1 ♀, ibid (sputter coated with gold) (ZIN); 1 ♀, ibid, but 20.i.[19]01, with an additional written-printed label: "
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<taxonomicName class="Insecta" family="Histeridae" genus="Exaesiopus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exaesiopus" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">Exaesiopus</taxonomicName>
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/ duriculus Rtt. / Reichardt det." (TLAN).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnostic description">
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<paragraph pageId="10" pageNumber="11">Diagnostic description.</paragraph>
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<paragraph lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
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Body size: PEL: 2.50-3.75 mm; APW: 0.75-1.15 mm; PPW: 1.90-2.75 mm; EW: 2.00-3.00 mm; EL: 1.75-2.50 mm. Body in general (except for
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus pavlovskii" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pavlovskii">
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<pageBreakToken pageId="11" pageNumber="12" start="start">Reichardtiolus</pageBreakToken>
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pavlovskii
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</taxonomicName>
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) larger than the rest of congeners, cuticle similar to that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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; legs, antennae and mouthparts chestnut brown. Mouthparts similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, mentum on anterior margin deeply emarginated medially (Fig. 49). Clypeus and frons (Fig. 48) coarsely and densely punctate; frontal stria weakened medially; frontal disk with low protuberances and shallow depressions, very coarsely and densely punctate, especially medially; clypeus margined laterally. Pronotum as in
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, punctuation medially sparser, punctures weak and separated by several times their diameter. Elytra generally similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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; punctuation of pygydium generally denser than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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(compare Figs 12 and 51). Prosternal process flattened to slightly concave, compressed laterally; carinal prosternal striae approximate, complete; prosternal foveae small. Mesoventrite sub-quadrate, marginal stria anteriorly complete; punctuation sparser than that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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, punctures separated by several times their diameter; meso-metaventral stria absent, in case of one specimen substituted by a string of punctures. Metaventrite, metepisternum and abdominal ventrites similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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. Legs similar to those of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus sphingis" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sphingis">Reichardtiolus sphingis</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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. Male genitalia: 8th sternite (Figs 52-53) strongly sclerotized, apically with dense row of short setae and setose velum; 8th tergite apically with deep emargination, on basal half with numerous pores and pseudopores (Fig. 53). Sclerotization of 9th tergite divided medially (as in
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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), on apical half with pores and pseudopores; 10th tergite inwardly arcuate on its basal margin. 9th tergite on apical third with faint, weakly sclerotized bisinuate line, visible only from lateral view (Fig. 59). Spiculum gastrale (Fig. 54) on apical end inwardly arcuate (although not as deeply as with
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus sphingis" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sphingis">Reichardtiolus sphingis</taxonomicName>
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or
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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), with a unique sclerotized ring medially; basal end of spiculum gastrale outwardly arcuate. Aedeagus generally most similar to that of
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus sphingis" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="sphingis">Reichardtiolus sphingis</taxonomicName>
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, but blunt apically (compare Figs 31 and 58).
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</paragraph>
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<caption pageId="11" pageNumber="12">
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<paragraph pageId="11" pageNumber="12">
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Figures 48-51. 48
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., head, dorsal view 49
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., mentum, ventral view 50
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., prosternum 51
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., pygidium.
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</paragraph>
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</caption>
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<caption pageId="11" pageNumber="12">
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<paragraph pageId="11" pageNumber="12">
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Figures 52-61. 52
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., 8th sternite and tergite, ventral view 53 ditto, dorsal view 54
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., spiculum gastrale, ventral view 55 ditto, lateral view 56
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., 8th sternite and tergite, lateral view 57
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., 9th + 10th tergites, dorsal view 58
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., apex of aedeagus, frontal view 59
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., 9th + 10th tergites, lateral view 60
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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sp. n., aedeagus, dorsal view 61 ditto, lateral view.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="12" pageNumber="13" type="differential diagnosis">
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<paragraph pageId="12" pageNumber="13">
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<pageBreakToken pageId="12" pageNumber="13" start="start">Differential</pageBreakToken>
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diagnosis.
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</paragraph>
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<paragraph pageId="12" pageNumber="13">
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus perses" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="perses">Reichardtiolus perses</taxonomicName>
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is the second largest species of the genus (after
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus pavlovskii" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pavlovskii">Reichardtiolus pavlovskii</taxonomicName>
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) and externally very similar to
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus duriculus" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="duriculus">Reichardtiolus duriculus</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus aldhaferi" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="aldhaferi">Reichardtiolus aldhaferi</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus sphingis" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="sphingis">Reichardtiolus sphingis</taxonomicName>
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, differing from them mainly by the structure of male terminalia. From the largest species of the genus,
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<taxonomicName class="Insecta" family="Histeridae" genus="Reichardtiolus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Reichardtiolus pavlovskii" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="pavlovskii">Reichardtiolus pavlovskii</taxonomicName>
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it differs by the same characteristics as the preceding three species.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="biology">
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<paragraph pageId="13" pageNumber="14">
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<pageBreakToken pageId="13" pageNumber="14" start="start">Biology</pageBreakToken>
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.
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</paragraph>
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<paragraph pageId="13" pageNumber="14">Unknown, presumably similar to its congeners.</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="distribution">
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<paragraph pageId="13" pageNumber="14">Distribution.</paragraph>
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<paragraph pageId="13" pageNumber="14">Iran, environs of Kerman (Fig. 72).</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="14" type="etymology">
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<paragraph pageId="13" pageNumber="14">Etymology.</paragraph>
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<paragraph pageId="13" pageNumber="14">
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The name of this new species means
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<normalizedToken originalValue="“Persian”">"Persian"</normalizedToken>
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. It is a noun in apposition in the nominative singular form.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |