139 lines
13 KiB
XML
139 lines
13 KiB
XML
<document id="43941E44911E683F28A46B71F27B61D7" ID-CLB-Dataset="32897" ID-DOI="http://dx.doi.org/10.3897/zookeys.678.12886" ID-GBIF-Dataset="5ccea4ea-e00a-42ee-ac09-7fd965d6c9af" ID-PMC="PMC5523363" ID-Pensoft-Pub="1313-2970-678-73" ID-PubMed="28769697" ID-ZooBank="69702921167246A9AE5E033DD0E41B8C" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-678-73" ModsDocOrigin="ZooKeys 678" ModsDocTitle="Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae)" checkinTime="1496801215117" checkinUser="pensoft" docAuthor="Bergsten, Johannes, Weingartner, Elisabeth & Hajek, Jiri" docDate="2017" docId="2A5ECDD7849177F8C1EC6A3C83D17A90" docLanguage="en" docName="ZooKeys 678: 73-96" docOrigin="ZooKeys 678" docSource="http://dx.doi.org/10.3897/zookeys.678.12886" docTitle="Hyphydrus sanctus Sharp 1882" docType="treatment" docVersion="6" lastPageNumber="76" masterDocId="FFB20778FFF04C67F159FB77FFB2FFBC" masterDocTitle="Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae)" masterLastPageNumber="96" masterPageNumber="73" pageNumber="76" updateTime="1701374349769" updateUser="ExternalLinkService">
|
||
<mods:mods id="BCBDC4E1951867367E6CCC27C0F1CE77" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CDBA1EF56D45967D0980465384748483">
|
||
<mods:title id="0D56DC797397E7E6D1E2569E380A288C">Species delimitation of the Hyphydrusovatus complex in western Palaearctic with an update of species distributions (Coleoptera, Dytiscidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="42ED4EF9320586E58368D616D90616F8" type="personal">
|
||
<mods:role id="A8013A50A0B89CC19B139C8DFA6F84FB">
|
||
<mods:roleTerm id="21285A9B2B57655A69C6DE546188BFAC">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E5A64CADF558DF1E3F4D35B578149BFD">Bergsten, Johannes</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="DE551EA228835319B5F824E7A8170083" type="personal">
|
||
<mods:role id="BDAB2E549414EC372437016CA41ED06F">
|
||
<mods:roleTerm id="73FE23CC86209726AEFDA60F365D651E">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2C65C60394276E6E54E0FCE870253212">Weingartner, Elisabeth</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6FC42081FB0908A58F2D6F28FA8A2553" type="personal">
|
||
<mods:role id="64E8F0AB59B518EFD36B8C94D82BD473">
|
||
<mods:roleTerm id="65C099D5CD4AF40C550FC8E01F848CC3">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4B23EF6CE092B6BDE44B016A143DEB72">Hajek, Jiri</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="E1A2FCC353FAFC3FAC4199EF8D48B896">text</mods:typeOfResource>
|
||
<mods:relatedItem id="F355925B7E794E1BE00E634BC4D95E71" type="host">
|
||
<mods:titleInfo id="33B2C8B5EB0DB8911B6A2B5586817C5B">
|
||
<mods:title id="23CFD904E1B6395E58C8C9622EF6C329">ZooKeys</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="B6A0A44BBE1A10A4E450B71918517F0B">
|
||
<mods:date id="161601BB51435626A005C984DCCDF11C">2017</mods:date>
|
||
<mods:detail id="6B2C756A0F13973542F56A955151E133" type="volume">
|
||
<mods:number id="F44C40A67363ABF1FA56C799384F9EB0">678</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="0697D589DDF464C0F8D0C53874C3FD41" unit="page">
|
||
<mods:start id="CFF4978E1F9F2B1B0D742E09F455B8F0">73</mods:start>
|
||
<mods:end id="D0F97B481664331551137CDA9EF712EF">96</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="0F0C877C014B654055F0D82C597A66EA">
|
||
<mods:url id="F05B7248A847BACB0B0B50898FE8EEDD">http://dx.doi.org/10.3897/zookeys.678.12886</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="8C75950D0F6F6001B17C37B7CBB4E9B8">journal article</mods:classification>
|
||
<mods:identifier id="FDB03A48304E9391D79372E19105DF47" type="DOI">http://dx.doi.org/10.3897/zookeys.678.12886</mods:identifier>
|
||
<mods:identifier id="E8AFD8951F2BE32A4AC7FE3F1D9ECA2A" type="Pensoft-Pub">1313-2970-678-73</mods:identifier>
|
||
<mods:identifier id="009B2684156E1870569874E4AB3C86DE" type="ZooBank">69702921167246A9AE5E033DD0E41B8C</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="2A5ECDD7849177F8C1EC6A3C83D17A90" ID-GBIF-Taxon="131812398" LSID="urn:lsid:plazi:treatment:2A5ECDD7849177F8C1EC6A3C83D17A90" httpUri="http://treatment.plazi.org/id/2A5ECDD7849177F8C1EC6A3C83D17A90" lastPageNumber="76" pageId="3" pageNumber="76">
|
||
<subSubSection id="4BFFE7A985FA4C323C5A242540BE5F7D" pageId="3" pageNumber="76" type="nomenclature">
|
||
<paragraph id="5188AC1C6D66344FFF849E9EE90C7257" pageId="3" pageNumber="76">
|
||
<taxonomicName id="CC8139BC681C957F76EF755306DE2DD7" ID-CoL="3NSSY" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Hyphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphydrus sanctus" order="Coleoptera" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="sanctus">
|
||
<pageBreakToken id="4261B86FA0A01636E818914B3500A1A4" pageId="3" pageNumber="76" start="start">Hyphydrus</pageBreakToken>
|
||
sanctus Sharp, 1882
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="BD0CA7AB19BFF14DDECC423358956BB7" pageId="3" pageNumber="76" type="reference_group">
|
||
<paragraph id="FCD19A4ECF58D03368E76F2829051933" pageId="3" pageNumber="76">
|
||
<taxonomicName id="9F02814C0197A31315D6DA66B68244F1" class="Insecta" family="Dytiscidae" genus="Hyphydrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hyphydrus sanctus" order="Coleoptera" pageId="3" pageNumber="76" phylum="Arthropoda" rank="species" species="sanctus">Hyphydrus sanctus</taxonomicName>
|
||
Sharp, 1882: 380.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2432F6D5D920CA4A761E7F43B7295B76" pageId="3" pageNumber="76" type="published records">
|
||
<paragraph id="B3A8866F23D3EACC9E9B6786156F0CAA" pageId="3" pageNumber="76">Published records.</paragraph>
|
||
<paragraph id="57047F9A2FD8F9985B85C6A07E511676" pageId="3" pageNumber="76">
|
||
Israel:
|
||
<bibRefCitation id="B82E24C692B1BB2467A74F5C4F292DB2" author="Sharp, D" journalOrPublisher="Scientific Transactions of the Royal Dublin Society" pageId="8" pageNumber="81" pagination="179 - 1003" title="On aquatic carnivorous Coleoptera or Dytiscidae" volume="2" year="1882">Sharp (1882</bibRefCitation>
|
||
: 380);
|
||
<bibRefCitation id="E79E57E73142DB2B66AF1F216DF7126A" author="Bistroem, O" journalOrPublisher="Acta Zoologica Fennica" pageId="7" pageNumber="80" pagination="1 - 121" title="A revision of the genus Hyphydrus Illiger (Coleoptera, Dytiscidae)." volume="165" year="1982">
|
||
<normalizedToken id="34A7D758D2974FEF3E995092CD609D61" originalValue="Biström">Bistroem</normalizedToken>
|
||
(1982
|
||
</bibRefCitation>
|
||
: 39);
|
||
<bibRefCitation id="BF4A8CF18FEFE8A6A848C5A9924B7AA5" author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="8" pageNumber="81" pagination="129 - 140" title="Neue und bemerkenswerte Schwimmkaefer aus dem Nahen Osten." volume="57" year="1984">Wewalka (1984</bibRefCitation>
|
||
: 131). Jordan:
|
||
<bibRefCitation id="212B940AA38C70AB254BE6EAD57C86AA" author="Bistroem, O" journalOrPublisher="Acta Zoologica Fennica" pageId="7" pageNumber="80" pagination="1 - 121" title="A revision of the genus Hyphydrus Illiger (Coleoptera, Dytiscidae)." volume="165" year="1982">
|
||
<normalizedToken id="329DB4655DC16EC779503F045E51D4F4" originalValue="Biström">Bistroem</normalizedToken>
|
||
(1982
|
||
</bibRefCitation>
|
||
: 39);
|
||
<bibRefCitation id="1E56906D0024DD4763FD6C35073A03AE" author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="8" pageNumber="81" pagination="129 - 140" title="Neue und bemerkenswerte Schwimmkaefer aus dem Nahen Osten." volume="57" year="1984">Wewalka (1984</bibRefCitation>
|
||
: 131). Syria:
|
||
<bibRefCitation id="53A4B1008290FD5030697988836E0201" author="Bistroem, O" journalOrPublisher="Acta Zoologica Fennica" pageId="7" pageNumber="80" pagination="1 - 121" title="A revision of the genus Hyphydrus Illiger (Coleoptera, Dytiscidae)." volume="165" year="1982">
|
||
<normalizedToken id="A93575BD6FD1C594B63E92E593961198" originalValue="Biström">Bistroem</normalizedToken>
|
||
(1982
|
||
</bibRefCitation>
|
||
: 39);
|
||
<bibRefCitation id="08F57FA7D1E35A2055C79C15867283A3" author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="8" pageNumber="81" pagination="129 - 140" title="Neue und bemerkenswerte Schwimmkaefer aus dem Nahen Osten." volume="57" year="1984">Wewalka (1984</bibRefCitation>
|
||
: 131).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E004DC3D55ADC04EC5F9E6FA3C9D4859" pageId="3" pageNumber="76" type="materials_examined">
|
||
<paragraph id="1FCDF84B7E0FA058DC1FF3340A20FC2B" pageId="3" pageNumber="76">Material examined.</paragraph>
|
||
<paragraph id="E988E8747AA42C3DA73BF7BCC83B4D61" pageId="3" pageNumber="76">
|
||
Israel: 2♂♂, 1♀, Hula reserve, 21.iii.1985; 4♂♂, 8♀♀, same locality, but 13.iv.1986; 1♂, 3♀♀, Talme Elazar, 21.iv.1986; 1♂, 6♀♀, Magan Michael, 21.iv.1986, all M.
|
||
<normalizedToken id="3D20007037AA3DC7E09D22F1948D91D5" originalValue="Jäch">Jaech</normalizedToken>
|
||
leg. (NHMW, NMPC). Turkey: 3♂♂, 13♀♀,
|
||
<normalizedToken id="7C4F661C9D8F15BB25A3D3050E46C020" originalValue="Muğla">Mugla</normalizedToken>
|
||
vil. [=province],
|
||
<normalizedToken id="90F977A75095CC1C76368E8F103B07A0" originalValue="Köyçeğiz">Koeycegiz</normalizedToken>
|
||
, 27.v.1991, S.
|
||
<normalizedToken id="FD2FA49A51EBEF430FADF229CD5F975B" originalValue="Schödl">Schoedl</normalizedToken>
|
||
leg. (NHMW, NMPC); 3♂♂, 2♀♀, same data, but M.
|
||
<normalizedToken id="AC0CBC3F1AEDB57DF8A3D96BFE8298E4" originalValue="Jäch">Jaech</normalizedToken>
|
||
leg. (NHMW, NMPC).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E43DA08BBD3C571269EF6EE0F4C0AEE9" pageId="3" pageNumber="76" type="diagnosis">
|
||
<paragraph id="2BE56F67E5701B84BC44FDCE10F2CC3A" pageId="3" pageNumber="76">Diagnosis.</paragraph>
|
||
<paragraph id="91204F68F205AE54FC919350ACB1F3A7" pageId="3" pageNumber="76">Habitus as depicted in Figs 4d, 5d. Clypeus with anterior margin medially nearly straight (Fig. 6c). Reticulation of dorsal surface confined to head and more distinct and impressed anteriorly (Fig. 6c), and to sides of pronotum. Punctation of head fine, visible on whole surface (Fig. 6c); punctures dense, distance between them smaller than their diameter (Fig. 6c). Punctation of pronotum double, fine, distance between larger punctures bigger than their diameter. Punctation of elytra double, diameter of small punctures less than half of diameter of large punctures; distance between large punctures bigger than their diameter. Epipleura reticulated with very fine punctures. Metatibia with outer margin nearly straight.</paragraph>
|
||
<paragraph id="7684A0ACD52381C3FC6FF2C1DAFFFC22" pageId="3" pageNumber="76">
|
||
Male. Longer metatibial spur long, nearly as long as metatarsomere I-II combined (Fig. 7c); spur broad and straight in basal two thirds with small but distinct serration, attenuated and curved apically (Fig. 7c). Male genitalia as in Fig. 8
|
||
<normalizedToken id="E574656DBB1883539EE345F6E3079F71" originalValue="o–r">o-r</normalizedToken>
|
||
, median lobe in ventral view slightly narrowing from base to apex.
|
||
</paragraph>
|
||
<paragraph id="D87B21FEC8653EF216C420310CD56B7D" pageId="3" pageNumber="76">
|
||
Female. Only matt females of
|
||
<taxonomicName id="4CBA3D78FDB91A3C1BDD62C08F4DE218" lsidName="H. sanctus" pageId="3" pageNumber="76" rank="species" species="sanctus">H. sanctus</taxonomicName>
|
||
are known so far. Whole surface densely reticulated, meshes on elytra somewhat elongate. Large punctures well visible, small punctures indistinct among reticulation. Longer tibial spur similar to that of male, but almost straight in apical third. Female genitalia as in Fig. 8
|
||
<normalizedToken id="F16562048AC5C4B9036654A61CEF546E" originalValue="s–u">s-u</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="080D000BA7DEE6110CB6B17BEE4FA569" pageId="3" pageNumber="76" type="habitat">
|
||
<paragraph id="F144B853404FB68DECB3DECAE74F1E0E" pageId="3" pageNumber="76">Habitat.</paragraph>
|
||
<paragraph id="D9CE59D9F8016F78A1C06D600751FABD" pageId="3" pageNumber="76">
|
||
Similarly to the other two species,
|
||
<taxonomicName id="F1F14106E9A00ED395795988DE23F7E5" lsidName="H. sanctus" pageId="3" pageNumber="76" rank="species" species="sanctus">H. sanctus</taxonomicName>
|
||
inhabits various types of standing and slowly flowing water bodies with at least some vegetation.
|
||
<bibRefCitation id="7440615FEC8D3AC68689FBADB5F14C44" author="Wewalka, G" journalOrPublisher="Koleopterologische Rundschau" pageId="8" pageNumber="81" pagination="129 - 140" title="Neue und bemerkenswerte Schwimmkaefer aus dem Nahen Osten." volume="57" year="1984">Wewalka (1984)</bibRefCitation>
|
||
reported several specimens from a densely vegetated pool and single occurrences from an artificial pool with clear water, an irrigation ditch and from a stream.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="07E0ADB0465598B507F6E14D5EB8F999" pageId="3" pageNumber="76" type="distribution">
|
||
<paragraph id="D53C89DB4FC8DB14295A5DDB8F79F204" pageId="3" pageNumber="76">Distribution.</paragraph>
|
||
<paragraph id="8E28BD56E1A04ABDC2803FB0D4664DDA" pageId="3" pageNumber="76">A species distributed in the Levant region of the Near East. So far recorded from several localities in Israel, Jordan and Syria (Fig. 9). First record from Turkey.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |