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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.788.21222" ID-GBIF-Dataset="f062cb75-2d34-4171-a3f5-72fb9771df04" ID-PMC="PMC6189229" ID-Pensoft-Pub="1313-2970-788-87" ID-PubMed="30337826" ID-ZBK="FCC76C6C5445497F9C6717A4F1A51464" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1313-2970-788-87" ModsDocOrigin="ZooKeys 788" ModsDocTitle="A review of Leucosigma Druce, 1908: a newly discovered case of fern-feeding and descriptions of three new species (Lepidoptera, Noctuidae)" checkinTime="1539050140006" checkinUser="pensoft" docAuthor="Goldstein, Paul Z., Janzen, Daniel H., Proshek, Benjamin, Dapkey, Tanya &amp; Hallwachs, Winnie" docDate="2018" docId="29F8B224D3827AE8AA0E4CAC72EC1FC0" docLanguage="en" docName="ZooKeys 788: 87-133" docOrigin="ZooKeys 788" docSource="http://dx.doi.org/10.3897/zookeys.788.21222" docTitle="Leucosigma viridipicta Dognin 1910, comb. n." docType="treatment" docVersion="4" lastPageNumber="100" masterDocId="FFC1FD22FF872E407E7FFF8E7B5FFFF9" masterDocTitle="A review of Leucosigma Druce, 1908: a newly discovered case of fern-feeding and descriptions of three new species (Lepidoptera, Noctuidae)" masterLastPageNumber="133" masterPageNumber="87" pageNumber="100" updateTime="1668166274258" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of Leucosigma Druce, 1908: a newly discovered case of fern-feeding and descriptions of three new species (Lepidoptera, Noctuidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Goldstein, Paul Z.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Janzen, Daniel H.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Proshek, Benjamin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dapkey, Tanya</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hallwachs, Winnie</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>788</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>87</mods:start>
<mods:end>133</mods:end>
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</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.788.21222</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.788.21222</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-788-87</mods:identifier>
<mods:identifier type="ZBK">FCC76C6C5445497F9C6717A4F1A51464</mods:identifier>
<mods:identifier type="ZooBank">FCC76C6C5445497F9C6717A4F1A51464</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="148657023" LSID="urn:lsid:plazi:treatment:29F8B224D3827AE8AA0E4CAC72EC1FC0" httpUri="http://treatment.plazi.org/id/29F8B224D3827AE8AA0E4CAC72EC1FC0" lastPageNumber="100" pageId="13" pageNumber="100">
<subSubSection pageId="13" pageNumber="100" type="nomenclature">
<paragraph pageId="13" pageNumber="100">
<taxonomicName authority="Dognin, 1910" authorityName="Dognin" authorityYear="1910" class="Insecta" family="Noctuidae" genus="Leucosigma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Leucosigma viridipicta" order="Lepidoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="viridipicta">Leucosigma viridipicta (Dognin, 1910)</taxonomicName>
<taxonomicNameLabel pageId="13" pageNumber="100">comb. n.</taxonomicNameLabel>
Figs 41, 42, 51, 52; Male genitalia: Figs 65, 88, 113, 114) 127, 128
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="reference_group">
<paragraph pageId="13" pageNumber="100">
<taxonomicName class="Insecta" family="Noctuidae" genus="Gonodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonodes viridipicta" order="Lepidoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="viridipicta">Gonodes viridipicta</taxonomicName>
Dognin, 1910: 13. Type locality: French Guiana: St. Laurent du Maroni.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="materials_examined">
<paragraph pageId="13" pageNumber="100">Material examined.</paragraph>
<paragraph pageId="13" pageNumber="100">
Type material. FRENCH GUIANA: HOLOTYPE ♂; S. -Laurent de Maroni Guy Franc; Dognin collection,
<taxonomicName class="Insecta" family="Noctuidae" genus="Gonodes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gonodes viridipicta" order="Lepidoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="species" species="viridipicta">Gonodes viridipicta</taxonomicName>
1/10 Type ♂ Dognin not in USNM [illeg.], Type No. 32413 U.S.N.M., ♂ USNM Dissection 148176, USNMENT00973419. Type at USNM.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="materials_examined">
<paragraph pageId="13" pageNumber="100">Other material examined.</paragraph>
<paragraph pageId="13" pageNumber="100">(1♂). PERU: Huacamayo, Carabaya, dry seas., 3100 ft, June 04. (G. Ockenden), Rothschild Bequest B.M. 1939-1, NHMUK01606202.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="diagnosis">
<paragraph pageId="13" pageNumber="100">Diagnosis.</paragraph>
<paragraph pageId="13" pageNumber="100">
Smaller than
<taxonomicName lsidName="L. albimixta" pageId="13" pageNumber="100" rank="species" species="albimixta">L. albimixta</taxonomicName>
, with fused orbicular and reniform spots similarly swollen, but with the basal, antemedial, and postmedial lines less conspicuously highlighted in black and white. Cucullus with medial flange directed basad, its ventral edge precisely complementing the dorsal edge of clasper from which it appears to have been separated during development; apices of the costal lobes swollen.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="description">
<paragraph pageId="13" pageNumber="100">Re-description.</paragraph>
<paragraph pageId="13" pageNumber="100">Head. Antennae setose-ciliate, bifasciculate in males, scaling above uniformly grayish brown. Vertex and labial palpi with scales predominantly grayish brown; frons and inner face of palpus with paler scaling. Eyes smooth.</paragraph>
<paragraph pageId="13" pageNumber="100">
Thorax. Prothoracic scales grayish brown, concolorous with vertex. Wings. Forewing length 10.9 mm (holotype, male), average 10.8 mm (males, n = 2). Apical patch dominantly green on forewing upperside and underside, and on underside of hind wing.
<normalizedToken originalValue="Legs">Legs-</normalizedToken>
Scaling predominantly tannish brown, more or less concolorous with thoracic vestiture; a single pair of striped mid-tibial spurs, two pairs on hind-tibiae; three rows of tibial spines on all legs.
</paragraph>
<paragraph pageId="13" pageNumber="100">Abdomen. Dorsum covered in uniformly tannish-brown scales and hairs; ventral side more darkly scaled, especially at terminal tuft; some pinkish scaling ventrally.</paragraph>
<paragraph pageId="13" pageNumber="100">Male genitalia. Uncus robust and densely setose, almost bottlebrush-like for distal 3/4, the setae blonde, concolorous with neighboring setal tufts, and arranged in clusters sharing a single socket and shingled, appearing scale-like in situ. Tegumen raised at base of uncus. Vinculum laterally concave. Saccus blunt. Juxta rhomboid, without a dorsal projection. Sacculus (1) wide, barely tapered. Cucullus (2) bent backward (ventro-cephalad) and bears basally directed flange marking separation from dorsal edge of clasper, evidently ruptured during development of especially robust dorsal processes (3); each dorsal process swollen apically, resembling a ball-headed Native American war club, heavily setose, and with a conspicuous tuft of ventro-medially directed spine-like setae. Clasper (4) anvil shaped, its dorsal edge complementing the ventral edge of the cucullar flange. Aedeagus weakly sclerotized, granular appearance continuing to base of vesica; subbasal diverticulum asymmetrically bulbous, dumbbell-shaped.</paragraph>
<paragraph pageId="13" pageNumber="100">Female genitalia. Unknown.</paragraph>
<paragraph pageId="13" pageNumber="100">Immature stages. Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="biology_ecology">
<paragraph pageId="13" pageNumber="100">Biology.</paragraph>
<paragraph pageId="13" pageNumber="100">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="distribution">
<paragraph pageId="13" pageNumber="100">Distribution.</paragraph>
<paragraph pageId="13" pageNumber="100">French Guiana, Peru.</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="100" type="remarks">
<paragraph pageId="13" pageNumber="100">Remarks.</paragraph>
<paragraph pageId="13" pageNumber="100">
Female specimens at MNHN (Paris) were not available for study. This species is noteworthy in that the uncus is covered in shingled, scale-like clusters of setae (Figs 65, 127, 128), reminiscent of other fern-feeding species in the genus
<taxonomicName class="Insecta" family="Noctuidae" genus="Lophomyra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lophomyra" order="Lepidoptera" pageId="13" pageNumber="100" phylum="Arthropoda" rank="genus">Lophomyra</taxonomicName>
. The relationship between these genera requires more thorough sampling and analysis of both taxa and genes.
</paragraph>
</subSubSection>
</treatment>
</document>