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<document id="AEDAE390E0D6E4CB37EE508D61713D75" ID-CLB-Dataset="20208" ID-DOI="10.11646/zootaxa.5115.4.1" ID-GBIF-Dataset="b99d2dc5-e0a6-4f73-b2c6-b21b6350c69f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6361552" ID-ZooBank="320C2E00-8035-431A-8AD7-D483D81923F6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1647419732391" checkinUser="plazi" docAuthor="Ullah, Mishkat, Naeem, Muhammad, Mahmood, Khalid &amp; Garner, Beulah" docDate="2022" docId="291CD3215313DC79FF6DFBB38673F82F" docLanguage="en" docName="zootaxa.5115.4.1.pdf" docOrigin="Zootaxa 5115 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pristomachaerus coarctatus" docType="treatment" docVersion="7" lastPageNumber="455" masterDocId="D525AB595317DC7DFFFAFFFA836DFF91" masterDocTitle="Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan" masterLastPageNumber="486" masterPageNumber="451" pageNumber="455" updateTime="1699318106451" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="75141D4F6242922711457EB42A550DCF">Addition to the knowledge of Tribe Chlaeniini Brullé, 1834 (Coleoptera: Carabidae) from Pakistan</mods:title>
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<mods:namePart id="B4A6D29CE29425201ADC9795F0021D56">Ullah, Mishkat</mods:namePart>
<mods:affiliation id="CF590F7B93103E568BC585035BC137C0">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom.</mods:affiliation>
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<mods:namePart id="D5DFC8193E1D4B8CEFDC356CDDEDCB87">Naeem, Muhammad</mods:namePart>
<mods:nameIdentifier id="C7B3B5709D3A90296209490FE87940ED" type="ORCID">0000-0002-2025-2024</mods:nameIdentifier>
<mods:affiliation id="09B4C75B55821B99599EFA62480E2BEB">muhammad. naeem @ uaar. edu. pk; https: // orcid. org / 0000 - 0002 - 2025 - 2024</mods:affiliation>
<mods:nameIdentifier id="A1EEA47DE3454837D612DF08F99052A5" type="email">muhammad.naeem@uaar.edu.pk</mods:nameIdentifier>
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<mods:namePart id="035E7EBD3A7B540964627689E654B8F2">Mahmood, Khalid</mods:namePart>
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<mods:affiliation id="08BF4AAEB6D785AC08DE8B0F8504230F">dkmahmood @ msn. com; https: // orcid. org / 0000 - 0001 - 7162 - 3430</mods:affiliation>
<mods:nameIdentifier id="BB5721B02058853B36F094B83A9C2CE7" type="email">dkmahmood@msn.com</mods:nameIdentifier>
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<mods:namePart id="3DCF129581926A7C741124BBE9150805">Garner, Beulah</mods:namePart>
<mods:nameIdentifier id="DB2B147A757CF1ADF1826BFA2A0561A9" type="ORCID">0000-0002-5229-2450</mods:nameIdentifier>
<mods:affiliation id="A0A14149157221D9974C438859E4D477">Insects Division, Department of Life Sciences, Natural History Museum, Cromwell Road, London SW 7 5 BD, United Kingdom. &amp; b. garner @ nhm. ac. uk; https: // orcid. org / 0000 - 0002 - 5229 - 2450</mods:affiliation>
<mods:nameIdentifier id="428413EDD6272D988F225B426D387556" type="email">b.garner@nhm.ac.uk</mods:nameIdentifier>
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<mods:title id="089901993E1D05036670CD5DA09908AD">Zootaxa</mods:title>
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<mods:date id="56A368C43558993D5CF411422B4C6D60">2022</mods:date>
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<treatment id="291CD3215313DC79FF6DFBB38673F82F" ID-DOI="http://doi.org/10.5281/zenodo.6361454" ID-GBIF-Taxon="194051248" ID-Zenodo-Dep="6361454" LSID="urn:lsid:plazi:treatment:291CD3215313DC79FF6DFBB38673F82F" httpUri="http://treatment.plazi.org/id/291CD3215313DC79FF6DFBB38673F82F" lastPageNumber="455" pageId="4" pageNumber="455">
<subSubSection id="E9AF31BC5313DC79FF6DFBB38059FBF2" box="[151,820,1096,1124]" pageId="4" pageNumber="455" type="nomenclature">
<paragraph id="A10A62375313DC79FF6DFBB38059FBF2" blockId="4.[151,820,1096,1159]" box="[151,820,1096,1124]" pageId="4" pageNumber="455">
<heading id="FA42D55B5313DC79FF6DFBB38059FBF2" bold="true" box="[151,820,1096,1124]" fontSize="11" level="1" pageId="4" pageNumber="455" reason="1">
<taxonomicName id="66B519B45313DC79FF6DFBB38059FBF2" authority="(LaFerte-Senectere, 1851)" baseAuthorityName="LaFerte-Senectere" baseAuthorityYear="1851" box="[151,820,1096,1124]" class="Insecta" family="Carabidae" genus="Pristomachaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="coarctatus">
<emphasis id="93C1BE255313DC79FF6DFBB38059FBF2" bold="true" box="[151,820,1096,1124]" pageId="4" pageNumber="455">
<emphasis id="93C1BE255313DC79FF6DFBB38284FBF5" bold="true" box="[151,489,1097,1124]" italics="true" pageId="4" pageNumber="455">Pristomachaeruscoarctatus</emphasis>
(
<bibRefCitation id="C5241FC65313DC79FE02FBB28041FBF2" author="LaFerte-Senectere F. T. de" box="[504,812,1096,1123]" pageId="4" pageNumber="455" pagination="209 - 294" refId="ref27124" refString="LaFerte-Senectere F. T. de (1851) Revision de la tribu des patellimanes de Dejean, coleopteres pentameres de la famille des carabiques. Annales de la Societe Entomologique de France, 9 (2), 209 - 294." type="journal article" year="1851">LaFerté-Sénectère, 1851</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF6DFB97839AFB16" box="[151,247,1133,1159]" pageId="4" pageNumber="455" type="description">
<paragraph id="A10A62375313DC79FF6DFB97839AFB16" blockId="4.[151,820,1096,1159]" box="[151,247,1133,1159]" pageId="4" pageNumber="455">
(
<figureCitation id="398E7EB25313DC79FF5AFB978382FB16" box="[160,239,1133,1159]" captionStart="FIGURE 2" captionStartId="5.[151,250,1231,1255]" captionTargetBox="[151,1436,181,1206]" captionTargetId="figure-18@5.[151,1436,181,1207]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal habitus (a) Pristomachaerus coarctatus (LaFerté-Sénectère, 1851) imaged from NHMUK Collection; (b) Pristomachaerus littoralis littoralis (Motschulsky, 1859) comb. nov. imaged from NHMUK collection." figureDoi="http://doi.org/10.5281/zenodo.6361559" httpUri="https://zenodo.org/record/6361559/files/figure.png" pageId="4" pageNumber="455">Fig. 2a</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF6DFB4887EFFB5B" box="[151,1154,1201,1226]" pageId="4" pageNumber="455" type="reference_group">
<paragraph id="A10A62375313DC79FF6DFB4887EFFB5B" blockId="4.[151,1154,1201,1226]" box="[151,1154,1201,1226]" pageId="4" pageNumber="455">
<treatmentCitationGroup id="81A545195313DC79FF6DFB4887EFFB5B" box="[151,1154,1201,1226]" pageId="4" pageNumber="455">
<treatmentCitation id="201444265313DC79FF6DFB4881FAFB5B" author="LaFerte-Senectere F. T. de" box="[151,663,1201,1226]" page="230" pageId="4" pageNumber="455" year="1851">
<taxonomicName id="66B519B45313DC79FF6DFB4881FAFB5B" ID-CoL="68VLD" authority="LaFerte-Senectere, 1851: 230" authorityName="LaFerte-Senectere" authorityPageNumber="230" authorityYear="1851" box="[151,663,1201,1226]" class="Insecta" family="Carabidae" genus="Callistus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="coarctatus">
<emphasis id="93C1BE255313DC79FF6DFB48820FFB58" box="[151,354,1202,1226]" italics="true" pageId="4" pageNumber="455">Callistus coarctatus</emphasis>
<bibRefCitation id="C5241FC65313DC79FE92FB4881FAFB5B" author="LaFerte-Senectere F. T. de" box="[360,663,1201,1226]" pageId="4" pageNumber="455" pagination="209 - 294" refId="ref27124" refString="LaFerte-Senectere F. T. de (1851) Revision de la tribu des patellimanes de Dejean, coleopteres pentameres de la famille des carabiques. Annales de la Societe Entomologique de France, 9 (2), 209 - 294." type="journal article" year="1851">LaFerté-Sénectère, 1851: 230</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<taxonomicName id="66B519B45313DC79FD58FB488713FB5B" ID-CoL="68VJW" authority="Chaudoir, 1972: 383" authorityName="Chaudoir" authorityPageNumber="383" authorityYear="1972" box="[674,1150,1202,1226]" class="Insecta" family="Carabidae" genus="Callistomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="coarctatus">
<emphasis id="93C1BE255313DC79FD58FB4880C9FB58" box="[674,932,1202,1226]" italics="true" pageId="4" pageNumber="455">Callistomimus coarctatus</emphasis>
Chaudoir, 1972: 383
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF6DFB028245FAA7" pageId="4" pageNumber="455" type="materials_examined">
<paragraph id="A10A62375313DC79FF6DFB028245FAA7" blockId="4.[151,1437,1272,1983]" pageId="4" pageNumber="455">
<emphasis id="93C1BE255313DC79FF6DFB0282ECFA83" bold="true" box="[151,385,1272,1298]" pageId="4" pageNumber="455">Material examined.</emphasis>
<materialsCitation id="11DD686A5313DC79FE7DFB028249FAA7" ID-GBIF-Occurrence="3709719325" collectingDate="2015-08-27" collectionCode="PMNH" collectorName="Mishkat" country="Pakistan" location="PAKISTAN" pageId="4" pageNumber="455" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<collectingCountry id="D9A222A75313DC79FE7DFB028160FA83" box="[391,525,1272,1298]" name="Pakistan" pageId="4" pageNumber="455">PAKISTAN</collectingCountry>
<specimenCount id="B7B3A9BE5313DC79FDDFFB02813DFA83" box="[549,592,1272,1298]" pageId="4" pageNumber="455" type="male">1 ♂</specimenCount>
,
<specimenCount id="B7B3A9BE5313DC79FDA6FB0281EAFA83" box="[604,647,1272,1298]" pageId="4" pageNumber="455" type="female">1 ♀</specimenCount>
; Islamabad, Qauid-e-Azam University, Rumli;
<date id="D50B44F75313DC79FB60FB028658FA82" box="[1178,1333,1272,1299]" pageId="4" pageNumber="455" value="2015-08-27">
<collectingDate id="C54FBD1F5313DC79FB60FB028658FA82" box="[1178,1333,1272,1299]" pageId="4" pageNumber="455" value="2015-08-27">27 Aug. 2015</collectingDate>
</date>
;
<collectorName id="0C4007E15313DC79FABAFB0286F1FA83" box="[1344,1436,1272,1298]" pageId="4" pageNumber="455">Mishkat</collectorName>
leg.;
<collectionCode id="C7A4FAF25313DC79FF35FAE68249FAA7" box="[207,292,1308,1334]" country="USA" httpUri="http://grbio.org/cool/3s7c-0bcs" name="Peabody Museum of Natural History" pageId="4" pageNumber="455" type="Museum">PMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF3DFABA8251FA57" pageId="4" pageNumber="455" type="description">
<paragraph id="A10A62375313DC79FF3DFABA8251FA57" blockId="4.[151,1437,1272,1983]" pageId="4" pageNumber="455">
<emphasis id="93C1BE255313DC79FF3DFABA8213FACB" bold="true" box="[199,382,1344,1370]" pageId="4" pageNumber="455">Measurements.</emphasis>
Male (n=10): TL=
<quantity id="664DCFD25313DC79FDA5FABA81AAFACB" box="[607,711,1344,1371]" metricMagnitude="-3" metricUnit="m" metricValue="5.26" pageId="4" pageNumber="455" unit="mm" value="5.26">5.26 mm</quantity>
, HW=
<quantity id="664DCFD25313DC79FCE1FABA80EEFACB" box="[795,899,1344,1370]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="4" pageNumber="455" unit="mm" value="1.26">1.26 mm</quantity>
, PL=
<quantity id="664DCFD25313DC79FC33FABA875CFACB" box="[969,1073,1344,1370]" metricMagnitude="-3" metricUnit="m" metricValue="1.13" pageId="4" pageNumber="455" unit="mm" value="1.13">1.13 mm</quantity>
, PW=
<quantity id="664DCFD25313DC79FB7BFABA8785FACB" box="[1153,1256,1344,1370]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="4" pageNumber="455" unit="mm" value="1.49">1.49 mm</quantity>
, EL=
<quantity id="664DCFD25313DC79FACAFABA86F5FACB" box="[1328,1432,1344,1371]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" pageId="4" pageNumber="455" unit="mm" value="3.35">3.35 mm</quantity>
, EW=
<quantity id="664DCFD25313DC79FF27FA9E822AFAEF" box="[221,327,1380,1407]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="455" unit="mm" value="1.55">1.55 mm</quantity>
. Ratios PL/PW= 0.75, HW/PW= 0.84 and EL/EW= 2.16. Female (n=10): TL=
<quantity id="664DCFD25313DC79FB1CFA9E8622FAEF" box="[1254,1359,1380,1407]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" pageId="4" pageNumber="455" unit="mm" value="5.95">5.95 mm</quantity>
, HW=
<quantity id="664DCFD25313DC79FF6DFA728393FA33" box="[151,254,1416,1442]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="4" pageNumber="455" unit="mm" value="1.34">1.34 mm</quantity>
, PL=
<quantity id="664DCFD25313DC79FEB8FA7282C4FA33" box="[322,425,1416,1442]" metricMagnitude="-3" metricUnit="m" metricValue="1.26" pageId="4" pageNumber="455" unit="mm" value="1.26">1.26 mm</quantity>
, PW=
<quantity id="664DCFD25313DC79FE0CFA728130FA33" box="[502,605,1416,1442]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="4" pageNumber="455" unit="mm" value="1.62">1.62 mm</quantity>
, EL=
<quantity id="664DCFD25313DC79FD58FA728064FA33" box="[674,777,1416,1442]" metricMagnitude="-3" metricUnit="m" metricValue="3.67" pageId="4" pageNumber="455" unit="mm" value="3.67">3.67 mm</quantity>
, EW=
<quantity id="664DCFD25313DC79FCA2FA7280ADFA33" box="[856,960,1416,1442]" metricMagnitude="-3" metricUnit="m" metricValue="1.27" pageId="4" pageNumber="455" unit="mm" value="1.27">1.27 mm</quantity>
. Ratios PL/PW= 0.77, HW/PW= 0.82 and EL/EW= 2.88.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF3DFA2A80A6F977" pageId="4" pageNumber="455" type="diagnosis">
<paragraph id="A10A62375313DC79FF3DFA2A80A6F977" blockId="4.[151,1437,1272,1983]" pageId="4" pageNumber="455">
<emphasis id="93C1BE255313DC79FF3DFA2A822FFA7B" bold="true" box="[199,322,1488,1514]" pageId="4" pageNumber="455">Diagnosis.</emphasis>
Head black with metallic violet tinge. Clypeus and neck smooth. Frons median area smooth, sides and vertex setiferous punctate. Labrum, mandibles, labial and maxillary palpi reddish brown. Antenna black except articles IIII dark brown to reddish brown. Pronotum reddish brick, cordiform, apical border straight and wider than basal margin, densely foveate with pubescence towards lateral depressions. Elytra black; base with a brick red sutural mark; each elytron with three yellowish marks; first mark towards first quarter transverse shaped from half of interval IVVIII, second mark towards third quarter oblique shaped from interval III (touching II)VIII, third towards tip of the apex triangular shaped. Elytra lateral border black. Legs long and slender, femora apex, all tibia base and apex, all tarsi apex black rest of the leg yellowish. Venter black.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF3DF90A8128F80B" pageId="4" pageNumber="455" type="discussion">
<paragraph id="A10A62375313DC79FF3DF90A8128F80B" blockId="4.[151,1437,1272,1983]" pageId="4" pageNumber="455">
<emphasis id="93C1BE255313DC79FF3DF90A8222F89B" bold="true" box="[199,335,1776,1802]" pageId="4" pageNumber="455">Comments.</emphasis>
This species is similar to
<taxonomicName id="66B519B45313DC79FD84F90B809CF89A" authority="(Motschulsky, 1859)" authorityName="Ullah &amp; Naeem &amp; Mahmood &amp; Garner" authorityYear="2022" baseAuthorityName="Motschulsky" baseAuthorityYear="1859" box="[638,1009,1776,1803]" class="Insecta" family="Carabidae" genus="Pristomachaerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="littoralis">
<emphasis id="93C1BE255313DC79FD84F90B8196F89B" box="[638,763,1776,1802]" italics="true" pageId="4" pageNumber="455">P. littoralis</emphasis>
(
<bibRefCitation id="C5241FC65313DC79FCF6F90A8085F89A" author="Motschulsky, V. von" box="[780,1000,1776,1803]" pageId="4" pageNumber="455" pagination="25 - 118" refId="ref27745" refString="Motschulsky, V. von (1859) Entomologie speciale. Insectes des Indes orientales, et de contrees analogues. 2: de serie. Etudes Entomologiques, 8, 25 - 118." type="journal article" year="1859">Motschulsky, 1859</bibRefCitation>
)
</taxonomicName>
. However,
<taxonomicName id="66B519B45313DC79FB89F90B8668F89B" authorityName="Chaudoir" authorityYear="1972" box="[1139,1285,1777,1802]" class="Insecta" family="Carabidae" genus="Callistomimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="455" phylum="Arthropoda" rank="species" species="coarctatus">
<emphasis id="93C1BE255313DC79FB89F90B8668F89B" box="[1139,1285,1777,1802]" italics="true" pageId="4" pageNumber="455">P. coarctatus</emphasis>
</taxonomicName>
differ on the basis of a large body size, pronotum reddish brown; elytra base with reddish brown longitudinal mark along suture and touching both preapical oblique marks; apical transverse yellowish white mark from interval VIX; preapical oblique mark yellowish white from interval VVIII; apex with small rounded yellowish white mark, elytralateralborderyellowishwhite.
</paragraph>
</subSubSection>
<subSubSection id="E9AF31BC5313DC79FF3DF85E8673F82F" box="[199,1310,1956,1983]" pageId="4" pageNumber="455" type="distribution">
<paragraph id="A10A62375313DC79FF3DF85E8673F82F" blockId="4.[151,1437,1272,1983]" box="[199,1310,1956,1983]" pageId="4" pageNumber="455">
<emphasis id="93C1BE255313DC79FF3DF85E820CF82F" bold="true" box="[199,353,1956,1982]" pageId="4" pageNumber="455">Distribution.</emphasis>
<collectingCountry id="D9A222A75313DC79FE9DF85E82CCF82F" box="[359,417,1956,1982]" name="India" pageId="4" pageNumber="455">India</collectingCountry>
(
<collectingRegion id="6371ACD55313DC79FE4AF85E8168F82F" box="[432,517,1956,1982]" country="India" name="Sikkim" pageId="4" pageNumber="455">Sikkim</collectingRegion>
, Darjeeling;
<collectingRegion id="6371ACD55313DC79FD6FF85E805EF82E" box="[661,819,1956,1983]" country="India" name="Uttar Pradesh" pageId="4" pageNumber="455">Uttar Pradesh</collectingRegion>
),
<collectingCountry id="D9A222A75313DC79FCBCF85E80E4F82F" box="[838,905,1956,1982]" name="Nepal" pageId="4" pageNumber="455">Nepal</collectingCountry>
and
<collectingCountry id="D9A222A75313DC79FC45F85E8773F82F" box="[959,1054,1956,1982]" name="Pakistan" pageId="4" pageNumber="455">Pakistan</collectingCountry>
(
<bibRefCitation id="C5241FC65313DC79FBD6F85E867FF82F" author="Kirschenhofer, E." box="[1068,1298,1956,1982]" pageId="4" pageNumber="455" pagination="1834" refId="ref26867" refString="Kirschenhofer, E. (2017) Tribe Chlaeniini Brulle, 1834. In: Lobel, I. &amp; Lobel, D. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata-Myxophaga-Adephaga. Brill, Leiden, pp. 482 - 497. https: // doi. org / 10.1163 / 9789004330290" type="book chapter" year="2017">Kirschenhofer 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>