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<document id="9F23A0F293952D96FFB7D987BCBD24F5" ID-DOI="10.11646/zootaxa.4033.2.10" ID-GBIF-Dataset="2a820c38-c23b-4c48-8388-8a54c8aea799" ID-ISSN="1175-5326" ID-Zenodo-Dep="237325" ID-ZooBank="72E4B44E-9A61-4D77-80A4-18CF7DF80B6F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461297157489" checkinUser="plazi" docAuthor="Ferrari, Rafael R." docDate="2015" docId="280F87ABFFD912147EE2FEEA9FB6FEA2" docLanguage="en" docName="zt04033p300.pdf" docOrigin="Zootaxa 4033 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geodiscelis (Nazcoediscelis) antiminera Ferrari, new species" docType="treatment" docVersion="8" lastPageNumber="298" masterDocId="D436FFD3FFDB12117E75FC489C6DFFFD" masterDocTitle="A new species of Geodiscelis Michener &amp; Rozen (Hymenoptera: Colletidae: Xeromelissinae) from Peru" masterLastPageNumber="300" masterPageNumber="293" pageNumber="295" updateTime="1698616705064" updateUser="plazi">
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<mods:title id="CDEE84D8DB01E883ADD3AA5CE78615CD">A new species of Geodiscelis Michener &amp; Rozen (Hymenoptera: Colletidae: Xeromelissinae) from Peru</mods:title>
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<mods:namePart id="BC07841E8403D0DECA15CD5FA2367737">Ferrari, Rafael R.</mods:namePart>
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<mods:date id="A5134118881927394D2AB7C70C04787C">2015</mods:date>
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<treatment id="280F87ABFFD912147EE2FEEA9FB6FEA2" ID-DOI="http://doi.org/10.5281/zenodo.6119437" ID-GBIF-Taxon="119642788" ID-Zenodo-Dep="6119437" LSID="urn:lsid:plazi:treatment:280F87ABFFD912147EE2FEEA9FB6FEA2" httpUri="http://treatment.plazi.org/id/280F87ABFFD912147EE2FEEA9FB6FEA2" lastPageId="5" lastPageNumber="298" pageId="2" pageNumber="295">
<subSubSection id="E8BC6536FFD912137EE2FEEA9C8AFD21" pageId="2" pageNumber="295" type="nomenclature">
<paragraph id="A01936BDFFD912137EE2FEEA9F14FD41" blockId="2.[151,889,674,732]" box="[151,889,674,700]" pageId="2" pageNumber="295">
<heading id="FB5181D1FFD912137EE2FEEA9F14FD41" bold="true" box="[151,889,674,700]" fontSize="11" level="1" pageId="2" pageNumber="295" reason="1">
<emphasis id="92D2EAAFFFD912137EE2FEEA9F14FD41" bold="true" box="[151,889,674,700]" pageId="2" pageNumber="295">
<taxonomicName id="67A64D3EFFD912137EE2FEEA9EB1FD41" authority="Ferrari" authorityName="Ferrari" box="[151,732,674,700]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov." subGenus="Nazcoediscelis">
<emphasis id="92D2EAAFFFD912137EE2FEEA9D4DFD41" bold="true" box="[151,288,674,700]" italics="true" pageId="2" pageNumber="295">Geodiscelis</emphasis>
(
<emphasis id="92D2EAAFFFD912137F44FEEA9D8DFD41" bold="true" box="[305,480,674,700]" italics="true" pageId="2" pageNumber="295">Nazcoediscelis</emphasis>
)
<emphasis id="92D2EAAFFFD912137F84FEEA9E1AFD41" bold="true" box="[497,631,674,700]" italics="true" pageId="2" pageNumber="295">antiminera</emphasis>
Ferrari
</taxonomicName>
,
<taxonomicNameLabel id="89E157D4FFD912137C9CFEEA9F14FD41" box="[745,889,674,700]" pageId="2" pageNumber="295" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="A01936BDFFD912137EE2FE8D9C8AFD21" blockId="2.[151,889,674,732]" box="[151,231,709,732]" pageId="2" pageNumber="295">
(
<figureCitation id="389D2A38FFD912137EEAFE8D9CB2FD21" box="[159,223,709,732]" captionStart="FIGURE 2" captionStartId="3.[151,244,1880,1902]" captionTargetBox="[159,1436,191,1859]" captionTargetId="figure@3.[151,1436,191,1859]" captionTargetPageId="3" captionText="FIGURE 2. Geodiscelis antiminera sp. nov. Holotype ♂: (A) habitus, lateral view; (C) habitus, dorsal view; (E) head, frontal view. Paratype ♀: (B) habitus, lateral view; (D) habitus, dorsal view; (F) head, frontal view." httpUri="https://zenodo.org/record/237327/files/figure.png" pageId="2" pageNumber="295">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="E8BC6536FFD912137EE2FF429951FC74" pageId="2" pageNumber="295" type="diagnosis">
<paragraph id="A01936BDFFD912137EE2FF429951FC74" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EE2FF429D61FCDC" bold="true" box="[151,268,778,801]" pageId="2" pageNumber="295">Diagnosis.</emphasis>
The following combination of characters is sufficient to differentiate
<taxonomicName id="67A64D3EFFD912137A7DFF4298F1FCDC" box="[1032,1180,778,801]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="295" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFD912137A7DFF4298F1FCDC" box="[1032,1180,778,801]" italics="true" pageId="2" pageNumber="295">G. antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFD912137AD0FF429891FCDC" box="[1189,1276,778,801]" pageId="2" pageNumber="295" rank="species">
<emphasis id="92D2EAAFFFD912137AD0FF429891FCDC" bold="true" box="[1189,1276,778,801]" pageId="2" pageNumber="295">sp. nov.</emphasis>
</taxonomicNameLabel>
from all other species in the genus (unless otherwise stated, all characters refer to both sexes): malar area ~1.5x longer than wide, but shorter than compound eye; disc of supraclypeal area flat in lateral view; wing veins dark-brown; metepisternum nearly glabrous; male S6 with apicomedian spine; and medioventral process on gonocoxa conical and well-developed.
</paragraph>
</subSubSection>
<subSubSection id="E8BC6536FFD912157EB2FFDD9ED9FE3F" lastPageId="4" lastPageNumber="297" pageId="2" pageNumber="295" type="description">
<paragraph id="A01936BDFFD912137EB2FFDD9D02FC32" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2FFDD9D3FFC51" bold="true" box="[199,338,917,940]" pageId="2" pageNumber="295">Description.</emphasis>
<emphasis id="92D2EAAFFFD912137F2DFFDD9DE2FC51" box="[344,399,917,940]" italics="true" pageId="2" pageNumber="295">Male</emphasis>
(
<typeStatus id="7F1D881FFFD912137FE9FFDD9E6EFC51" box="[412,515,917,940]" pageId="2" pageNumber="295" type="holotype">Holotype</typeStatus>
,
<figureCitation id="389D2A38FFD912137C7BFFDD9E3AFC51" box="[526,599,917,940]" captionStart="FIGURE 2" captionStartId="3.[151,244,1880,1902]" captionTargetBox="[159,1436,191,1859]" captionTargetId="figure@3.[151,1436,191,1859]" captionTargetPageId="3" captionText="FIGURE 2. Geodiscelis antiminera sp. nov. Holotype ♂: (A) habitus, lateral view; (C) habitus, dorsal view; (E) head, frontal view. Paratype ♀: (B) habitus, lateral view; (D) habitus, dorsal view; (F) head, frontal view." httpUri="https://zenodo.org/record/237327/files/figure.png" pageId="2" pageNumber="295">Figs. 2</figureCitation>
A, 2C, 2E):
<emphasis id="92D2EAAFFFD912137CADFFDD9FF9FC51" box="[728,916,917,940]" italics="true" pageId="2" pageNumber="295">Dimensions (mm)</emphasis>
: Approximate body length 3.5; head width 0.91; forewing length 2.3.
</paragraph>
<paragraph id="A01936BDFFD912137EB2FF929848FAFA" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2FF929D24FC0C" box="[199,329,986,1009]" italics="true" pageId="2" pageNumber="295">Colouration</emphasis>
: Black except yellow as follows: malar area anteriorly; clypeus (except extreme lateral margins); ventral surface of antenna; spot on pronotal lobe; foreleg (except for dark-brown smudge on femur dorsobasally, and tibia and basitarsus anteriorly whitish-yellow); apical ring on mesocoxa, mesotrochanter; mesofemur ventrally; mesotibia anteriorly and posterodistally; apical half of metacoxa; metatrochanter ventrally; metafemur ventral and posterodistally; metatibia anteromedially; T1T6 each with median transverse band; S1 distally, S2 and S3S6 (except for brown marking on discs). Whitish-yellow as follows: mandible (except apex dark amber); labrum; malar area posteriorly; hypostomal area; anterior 1/3 of tegula; mesotibia anterodistally; mesobasitarsus, external margin, proximal and distal 1/3 of anterior surface of metatibia; metabasitarsus; posterior 1/4 of T1T6; T7. Dark brown on: pedicel and flagellum dorsally; wing veins; proximal half of T1T6. Light brown on: mesotarsus.
</paragraph>
<paragraph id="A01936BDFFD912137EB2F95A9D15FA49" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2F95A9D28FAD4" box="[199,325,1298,1321]" italics="true" pageId="2" pageNumber="295">Pubescence</emphasis>
: White, thickened, appressed, dense on: paraocular, interalveolar, vertexal and genal areas, pronotum, scutellum, mesepisternum, metanotum, dorsolateral area of propodeum, anterior 1/3 of T1T6; sparser on frontal area disc of mesoscutum and metanotum. White, thin, suberect, sparse on: mandible, labrum, clypeus, hypostomal area, trochanters, femora, foretibia, mesotibia; slightly longer and denser on metatibia; much longer on posterior margins of mesotarsomeres 14.
</paragraph>
<paragraph id="A01936BDFFD912137EB2F988988AF9C2" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2F9889D41FA2A" box="[199,300,1472,1495]" italics="true" pageId="2" pageNumber="295">Sculpture</emphasis>
: Microreticulate on clypeus, supraclypeal, paraocularand genal areas, pronotum, scutellum, metanotum, metasomal terga. Granulose on frontal and vertexal areas, mesepisternum, metepisternum; strongly granulose on metapostnotum. Punctures irregularly spaced on clypeus (i=46d above, i=2d below), and paraocular area (i=d anteriorly, i=2d posteriorly); indistinct on frontal and vertexal areas; i&lt;d on mesoscutum and scutellum.
</paragraph>
<paragraph id="A01936BDFFD912137EB2FA029822F8CC" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2FA029D45F99C" box="[199,296,1610,1633]" italics="true" pageId="2" pageNumber="295">Structure</emphasis>
:
<emphasis id="92D2EAAFFFD912137F42FA029D1CF99C" box="[311,369,1610,1633]" italics="true" pageId="2" pageNumber="295">Head</emphasis>
: Shorter than mesosoma (59:65), longer than wide (59:49). Labrum broader than long (12:8), apex rounded. Mandible lacking subapical tooth. Malar space much longer than basal depth of mandible (31:14), shorter than compound eye (31:55), malar line absent. Clypeus 1.5x longer than greatest breadth ignoring portion laterad to epistomal lobe (17:11); epistomal lobe strongly protruding into clypeus. Compound eye 28:19 (length to breadth), in profile margin coincident with that of genal profile ventrally; eyes slightly converging below. Genal area expanded posteriorly, 2/3 as wide as breadth of compound eye (19:21), ocelloccipital distance greater than diameter of lateral ocellus (10:8). Scape ~
<date id="D418107DFFD912137ED0FB529CADF8CC" box="[165,192,1818,1841]" pageId="2" pageNumber="295">2x</date>
as long as greatest width (21:10); pedicel cylindrical, slightly wider than long (10:8).
</paragraph>
<paragraph id="A01936BDFFD912137EB2FB759FECF864" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2FB759D54F8A9" box="[199,313,1853,1876]" italics="true" pageId="2" pageNumber="295">Mesosoma</emphasis>
: Anterior surface of pronotum nearly horizontal; ratio of lengths of scutellum:metanotum:horizontal surface of metaposnotum 19:10:20. Legs unmodified; hind tibial spurs long and narrow, inner spur half as long as metabasitarsus (23:37). Stigma with margin basal to vein R divergent.
</paragraph>
<paragraph id="A01936BDFFD912137EB2FBED9CD3F822" blockId="2.[151,1436,778,2015]" pageId="2" pageNumber="295">
<emphasis id="92D2EAAFFFD912137EB2FBED9D5BF841" box="[199,310,1957,1980]" italics="true" pageId="2" pageNumber="295">Metasoma</emphasis>
: Flattened, broadest at marginal zone of T3. S6 with apex angulate, bearing a spine. Genitalia as in
<figureCitation id="389D2A38FFD912137B04FBED9CC9F822" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="2" pageNumber="295">Fig. 3</figureCitation>
A.
</paragraph>
<caption id="F4D96635FFD812127EE2FB109FA3F876" httpUri="https://zenodo.org/record/237327/files/figure.png" pageId="3" pageNumber="296" targetBox="[159,1436,191,1859]" targetPageId="3">
<paragraph id="A01936BDFFD812127EE2FB109FA3F876" blockId="3.[151,1437,1880,1931]" pageId="3" pageNumber="296">
<emphasis id="92D2EAAFFFD812127EE2FB109D60F893" bold="true" box="[151,269,1880,1902]" pageId="3" pageNumber="296">FIGURE 2.</emphasis>
<taxonomicName id="67A64D3EFFD812127F61FB119D99F893" box="[276,500,1881,1902]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="296" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFD812127F61FB119D99F893" box="[276,500,1881,1902]" italics="true" pageId="3" pageNumber="296">Geodiscelis antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFD812127F89FB109E24F893" box="[508,585,1880,1902]" pageId="3" pageNumber="296" rank="species">
<emphasis id="92D2EAAFFFD812127F89FB109E24F893" bold="true" box="[508,585,1880,1902]" pageId="3" pageNumber="296">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype ♂:
<emphasis id="92D2EAAFFFD812127CA5FB109EB5F893" bold="true" box="[720,728,1880,1902]" pageId="3" pageNumber="296">(</emphasis>
A) habitus, lateral view; (C) habitus, dorsal view; (E) head, frontal view. Paratype ♀: (B) habitus, lateral view; (D) habitus, dorsal view; (F) head, frontal view.
</paragraph>
</caption>
<paragraph id="A01936BDFFDF12157EB2FCD09F5CFF52" blockId="4.[151,1436,152,971]" box="[199,817,152,175]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FCD09D7AFF52" box="[199,279,152,175]" italics="true" pageId="4" pageNumber="297">Female</emphasis>
(
<figureCitation id="389D2A38FFDF12157F50FCD09D1DFF52" box="[293,368,152,175]" captionStart="FIGURE 2" captionStartId="3.[151,244,1880,1902]" captionTargetBox="[159,1436,191,1859]" captionTargetId="figure@3.[151,1436,191,1859]" captionTargetPageId="3" captionText="FIGURE 2. Geodiscelis antiminera sp. nov. Holotype ♂: (A) habitus, lateral view; (C) habitus, dorsal view; (E) head, frontal view. Paratype ♀: (B) habitus, lateral view; (D) habitus, dorsal view; (F) head, frontal view." httpUri="https://zenodo.org/record/237327/files/figure.png" pageId="4" pageNumber="297">Figs. 2</figureCitation>
B, 2D, 2F): As in male, except as follows:
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FCF29859FF2C" blockId="4.[151,1436,152,971]" box="[199,1076,186,209]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FCF29D20FF2C" box="[199,333,186,209]" italics="true" pageId="4" pageNumber="297">Dimensions:</emphasis>
Approximate body length 3.7; head width 0.90; fore wing length 2.4.
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FC94993FFEC5" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FC949D3FFF0E" box="[199,338,220,243]" italics="true" pageId="4" pageNumber="297">Colouration:</emphasis>
Yellow marking on malar area slightly reduced; supraclypeal area entirely black; ventral surface of antenna brown, F1F3 darker; foretibia and forebasitarsus yellow anteriorly; mesotrochanter mostly dark-brown dorsally; outer surface of metatibia suffused with light-brown; white transverse band on metasomal terga broader.
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FD0C9FAFFE80" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FD0C9D20FEA6" box="[199,333,324,347]" italics="true" pageId="4" pageNumber="297">Pubescence:</emphasis>
Denser on frontal area; mesotarsal rake of longer, more robust setae (2
<date id="D418107DFFDF12157AF2FD0C98F9FEA6" box="[1159,1172,324,347]" pageId="4" pageNumber="297">4</date>
x MOD); metatibia and metabasitarsus with scopa (≤2.5x MOD); S2 with sparse scopa (≤
<date id="D418107DFFDF12157D26FD2E9F00FE80" box="[851,877,358,381]" pageId="4" pageNumber="297">2x</date>
MOD).
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FDC09F68FE62" blockId="4.[151,1436,152,971]" box="[199,773,392,415]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FDC09D58FE62" box="[199,309,392,415]" italics="true" pageId="4" pageNumber="297">Sculpture:</emphasis>
Less strongly granulose on metaposnotum.
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FDE39ED9FE3F" blockId="4.[151,1436,152,971]" box="[199,692,427,450]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FDE39D45FE3F" box="[199,296,427,450]" italics="true" pageId="4" pageNumber="297">Structure</emphasis>
: Pedicel globose; mesotibia swollen.
</paragraph>
</subSubSection>
<subSubSection id="E8BC6536FFDF12157EB2FD879D38FD45" pageId="4" pageNumber="297" type="materials_examined">
<paragraph id="A01936BDFFDF12157EB2FD879D2EFDD6" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FD879DE2FE1B" bold="true" box="[199,399,463,486]" pageId="4" pageNumber="297">Material studied.</emphasis>
<typeStatus id="7F1D881FFFDF12157FECFD879E6DFE1B" box="[409,512,463,486]" pageId="4" pageNumber="297" type="holotype">Holotype</typeStatus>
: “
<collectingCountry id="D8B1762DFFDF12157C6CFD879E0EFE1B" box="[537,611,463,486]" name="Peru" pageId="4" pageNumber="297">PERU</collectingCountry>
: Arequipa,
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W; La Reparticion
<date id="D418107DFFDF12157A73FD879810FE1B" box="[1030,1149,463,486]" pageId="4" pageNumber="297" value="2015-06-08">08/vi/2015</date>
;
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;
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yellow pan trap; Ferrari and Cañote” “
<taxonomicName id="67A64D3EFFDF12157CA7FDBA9F23FDF4" box="[722,846,498,521]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="92D2EAAFFFDF12157CA7FDBA9F23FDF4" box="[722,846,498,521]" italics="true" pageId="4" pageNumber="297">Geodiscelis</emphasis>
</taxonomicName>
♂;
<taxonomicName id="67A64D3EFFDF12157DF8FDBA986FFDF4" box="[909,1026,498,521]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFDF12157DF8FDBA986FFDF4" box="[909,1026,498,521]" italics="true" pageId="4" pageNumber="297">antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFDF12157A60FDBA981BFDF4" box="[1045,1142,498,521]" pageId="4" pageNumber="297" rank="species">
<emphasis id="92D2EAAFFFDF12157A60FDBA981BFDF4" bold="true" box="[1045,1142,498,521]" pageId="4" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
; R.R.Ferrari, det. 2015” “
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”.
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FE7198CCFD8E" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
<typeStatus id="7F1D881FFFDF12157EB2FE719D44FDAD" box="[199,297,569,592]" pageId="4" pageNumber="297" type="paratype">Paratype</typeStatus>
: “
<collectingCountry id="D8B1762DFFDF12157F49FE719DEBFDAD" box="[316,390,569,592]" name="Peru" pageId="4" pageNumber="297">PERU</collectingCountry>
: Arequipa,
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E; La Reparticion
<date id="D418107DFFDF12157D7CFE719FEDFDAD" box="[777,896,569,592]" pageId="4" pageNumber="297" value="2015-06-08">08/vi/2015</date>
;
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<geoCoordinate id="C592507AFFDF12157A79FE7198FEFDAD" box="[1036,1171,566,592]" direction="west" orientation="longitude" pageId="4" pageNumber="297" precision="9" value="-71.653336">71°39.20W</geoCoordinate>
;
<quantity id="675E9B58FFDF12157AE8FE71988AFDAD" box="[1181,1255,569,592]" metricMagnitude="3" metricUnit="m" metricValue="2.375" pageId="4" pageNumber="297" unit="m" value="2375.0">2375m</quantity>
yellow pan trap; Ferrari and Cañote” “
<taxonomicName id="67A64D3EFFDF12157F08FE149D94FD8E" box="[381,505,604,627]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="92D2EAAFFFDF12157F08FE149D94FD8E" box="[381,505,604,627]" italics="true" pageId="4" pageNumber="297">Geodiscelis</emphasis>
</taxonomicName>
♀;
<taxonomicName id="67A64D3EFFDF12157C57FE149EFAFD8E" box="[546,663,604,627]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFDF12157C57FE149EFAFD8E" box="[546,663,604,627]" italics="true" pageId="4" pageNumber="297">antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFDF12157CEBFE149E9EFD8E" box="[670,755,604,627]" pageId="4" pageNumber="297" rank="species">
<emphasis id="92D2EAAFFFDF12157CEBFE149E9EFD8E" bold="true" box="[670,755,604,627]" pageId="4" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
; R.R.Ferrari, det. 2015” “
<typeStatus id="7F1D881FFFDF12157A72FE1498F9FD8E" box="[1031,1172,604,627]" pageId="4" pageNumber="297" type="paratype">PARATYPE</typeStatus>
”.
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FE369D38FD45" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
Both specimens are currently deposited at
<collectionCode id="C6B7AE78FFDF12157CD4FE369E86FD68" box="[673,747,638,661]" httpUri="http://grbio.org/cool/66ra-d50z" name="The Packer Collection at York University" pageId="4" pageNumber="297">PCYU</collectionCode>
, however, they will be sent to
<collectionCode id="C6B7AE78FFDF12157A32FE3698F4FD68" box="[1095,1177,638,661]" pageId="4" pageNumber="297">MUSM</collectionCode>
pending completion of ongoing research.
</paragraph>
</subSubSection>
<subSubSection id="E8BC6536FFDF12157EB2FE8B9DA6FC99" pageId="4" pageNumber="297" type="etymology">
<paragraph id="A01936BDFFDF12157EB2FE8B9E80FCBF" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FE8B9D2AFD27" bold="true" box="[199,327,707,730]" pageId="4" pageNumber="297">Etymology.</emphasis>
<taxonomicName id="67A64D3EFFDF12157F24FE8B9E21FD27" box="[337,588,707,730]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFDF12157F24FE8B9E21FD27" box="[337,588,707,730]" italics="true" pageId="4" pageNumber="297">Geodiscelis antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFDF12157C23FE8B9EC3FD27" box="[598,686,707,730]" pageId="4" pageNumber="297" rank="species">
<emphasis id="92D2EAAFFFDF12157C23FE8B9EC3FD27" bold="true" box="[598,686,707,730]" pageId="4" pageNumber="297">sp. nov.</emphasis>
</taxonomicNameLabel>
is known only from the Department of Arequipa, a region of great importance for the mining industry in
<collectingCountry id="D8B1762DFFDF12157C5AFEAE9E0EFD00" box="[559,611,742,765]" name="Peru" pageId="4" pageNumber="297">Peru</collectingCountry>
. In Spanish, “antiminero means “anti-mining”, a feeling commonly shared by those inhabitants of Arequipa who rely on the soil and rivers to survive, and who are therefore opposed to the pollution of their natural resources caused by the mines.
</paragraph>
<paragraph id="A01936BDFFDF12157EB2FF059DA6FC99" blockId="4.[151,1436,152,971]" box="[199,459,845,868]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FF059D39FC99" bold="true" box="[199,340,845,868]" pageId="4" pageNumber="297">Floral hosts.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="E8BC6536FFDF12157EB2FF27987EFC36" pageId="4" pageNumber="297" type="discussion">
<paragraph id="A01936BDFFDF12157EB2FF27987EFC36" blockId="4.[151,1436,152,971]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EB2FF279D24FC7B" bold="true" box="[199,329,879,902]" pageId="4" pageNumber="297">Comments.</emphasis>
All photographs of the male
<typeStatus id="7F1D881FFFDF12157CF2FF279E89FC7B" box="[647,740,879,902]" pageId="4" pageNumber="297" type="holotype">holotype</typeStatus>
were taken prior to dissection. Currently, some metasomal terga are broken, and the metasoma is glued to a card point. The locality where the female
<typeStatus id="7F1D881FFFDF12157A42FFDA98FEFC54" box="[1079,1171,914,937]" pageId="4" pageNumber="297" type="paratype">paratype</typeStatus>
was collected is located ~
<quantity id="675E9B58FFDF12157ED1FFFC9C8FFC36" box="[164,226,948,971]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="297" unit="km" value="50.0">50km</quantity>
East of the type-locality. Both specimens were captured in yellow pan traps.
</paragraph>
</subSubSection>
<subSubSection id="E8BC6536FFDF12147EE2F8599FB6FEA2" lastPageId="5" lastPageNumber="298" pageId="4" pageNumber="297" type="key">
<paragraph id="A01936BDFFDF12157EE2F8599E3AFBB6" blockId="4.[151,971,1041,1099]" pageId="4" pageNumber="297">
<emphasis id="92D2EAAFFFDF12157EE2F8599E6DFBD4" bold="true" box="[151,512,1041,1065]" pageId="4" pageNumber="297">
Key to the species of
<taxonomicName id="67A64D3EFFDF12157FF1F8599E6DFBD4" ID-CoL="4MWS" box="[388,512,1041,1065]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="92D2EAAFFFDF12157FF1F8599E6DFBD4" bold="true" box="[388,512,1041,1065]" italics="true" pageId="4" pageNumber="297">Geodiscelis</emphasis>
</taxonomicName>
</emphasis>
[modified from
<bibRefCitation id="C4374B4CFFDF12157CC7F8599FABFBD5" author="Packer" box="[690,966,1041,1064]" pageId="4" pageNumber="297" refString="Packer, L. &amp; Dumesh, S. (2014) Two new species of Geodiscelis Michener and Rozen (Hymenoptera: Apoidea: Colletidae) with a phylogenetic analysis and subgeneric classification of the genus. Zootaxa, 3857 (2), 275 - 291. http: // dx. doi. org / 10.11646 / zootaxa. 3857.2.7" type="journal article" year="2014">Packer &amp; Dumesh (2014)</bibRefCitation>
] Note: the female
<taxonomicName id="67A64D3EFFDF12157F3AF87C9DA1FBB6" box="[335,460,1076,1099]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="species" species="phisquiri">
<emphasis id="92D2EAAFFFDF12157F3AF87C9DA1FBB6" box="[335,460,1076,1099]" italics="true" pageId="4" pageNumber="297">G. phisquiri</emphasis>
</taxonomicName>
is unknown.
</paragraph>
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<keyStep id="1B522E18FFDF12157EE2F83199F6FB32" pageId="4" pageNumber="297">
<paragraph id="A01936BDFFDF12157EE2F83199F6FB4D" blockId="4.[151,1436,1145,1797]" pageId="4" pageNumber="297">
<keyLead id="1B579588FFDF12157EE2F83199F6FB4D" pageId="4" pageNumber="297">
1 Malar space at least
<date id="D418107DFFDF12157FD0F8319DADFB6D" box="[421,448,1145,1168]" pageId="4" pageNumber="297">2x</date>
longer than wide (
<figureCitation id="389D2A38FFDF12157CFFF8319EA7FB6D" box="[650,714,1145,1168]" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
C); disc of supraclypeal area flat (
<figureCitation id="389D2A38FFDF12157A4DF831981AFB6D" box="[1080,1143,1145,1168]" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
C) (subgenus
<emphasis id="92D2EAAFFFDF12157B7BF8319C8DFB4D" italics="true" pageId="4" pageNumber="297">Nazcoediscelis</emphasis>
) ................................................................................................................................................................................ 3
</keyLead>
</paragraph>
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<keyLead id="1B579588FFDF12157EE2F8F099F6FB32" box="[151,1435,1208,1231]" pageId="4" pageNumber="297">
- Malar space at most only slightly longer than wide (
<figureCitation id="389D2A38FFDF12157C86F8F09F5FFB32" box="[755,818,1208,1231]" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
D); disc of supraclypeal area convex (
<figureCitation id="389D2A38FFDF12157AB2F8F09968FB32" box="[1223,1285,1208,1231]" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
D) .............. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="1B522E18FFDF12157EE2F89099F1FA71" pageId="4" pageNumber="297">
<paragraph id="A01936BDFFDF12157EE2F89099F6FAD0" blockId="4.[151,1436,1145,1797]" pageId="4" pageNumber="297">
<keyLead id="1B579588FFDF12157EE2F89099F6FAD0" pageId="4" pageNumber="297">
2 Malar space relatively long, about as long as wide [as in fig.
<quantity id="675E9B58FFDF12157D27F8909FE4FB12" box="[850,905,1240,1263]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="4" pageNumber="297" unit="in" value="16.0">16 in</quantity>
<bibRefCitation id="C4374B4CFFDF12157DFAF89098CBFB12" author="Packer" box="[911,1190,1240,1263]" pageId="4" pageNumber="297" refString="Packer, L. &amp; Dumesh, S. (2014) Two new species of Geodiscelis Michener and Rozen (Hymenoptera: Apoidea: Colletidae) with a phylogenetic analysis and subgeneric classification of the genus. Zootaxa, 3857 (2), 275 - 291. http: // dx. doi. org / 10.11646 / zootaxa. 3857.2.7" type="journal article" year="2014">Packer &amp; Dumesh (2014)</bibRefCitation>
]; female with two hind tibial spurs [as in fig.
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]; male with hind legs unmodified [as in fig.
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<bibRefCitation id="C4374B4CFFDF12157B43F8BF9D1DFAD0" author="Packer" pageId="4" pageNumber="297" refString="Packer, L. &amp; Dumesh, S. (2014) Two new species of Geodiscelis Michener and Rozen (Hymenoptera: Apoidea: Colletidae) with a phylogenetic analysis and subgeneric classification of the genus. Zootaxa, 3857 (2), 275 - 291. http: // dx. doi. org / 10.11646 / zootaxa. 3857.2.7" type="journal article" year="2014">Packer &amp; Dumesh (2014)</bibRefCitation>
]
<emphasis id="92D2EAAFFFDF12157FF7F95E9F3BFAD0" box="[386,854,1302,1325]" italics="true" pageId="4" pageNumber="297">
...................................................................
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</emphasis>
. (
<emphasis id="92D2EAAFFFDF12157D19F95E9F85FAD0" box="[876,1000,1302,1325]" italics="true" pageId="4" pageNumber="297">Geodiscelis</emphasis>
)
<emphasis id="92D2EAAFFFDF12157D8DF95E98EBFAD0" box="[1016,1158,1302,1325]" italics="true" pageId="4" pageNumber="297">megacephala</emphasis>
<bibRefCitation id="C4374B4CFFDF12157AF9F95E99F6FAD0" author="Michener" box="[1164,1435,1302,1325]" pageId="4" pageNumber="297" refString="Michener, C. D. &amp; Rozen, J. G. Jr. (1999) A new ground-nesting genus of xeromelissine bees from Argentina and the tribal classification of the subfamily (Hymenoptera: Colletidae). American Museum Novitates, 3281, 1 - 10." type="journal article" year="1999">Michener &amp; Rozen, 1999</bibRefCitation>
</keyLead>
</paragraph>
<paragraph id="A01936BDFFDF12157EE2F97E99F1FA71" blockId="4.[151,1436,1145,1797]" pageId="4" pageNumber="297">
<keyLead id="1B579588FFDF12157EE2F97E99F1FA71" pageId="4" pageNumber="297">
- Malar space extremely reduced, many times wider than long (
<figureCitation id="389D2A38FFDF12157D14F97E9FCDFAB0" box="[865,928,1334,1357]" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
D); female with only one hind tibial spur [as in fig.
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]; male with metafemur, metatibia and metabasitarsus considerably swollen [as in fig.
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3 Malar space longer than compound eye [as in fig.
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] .................................................... ..................................................................................................................
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- Malar space shorter than compound eye (
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D) ..................................................................................................... 4
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4 Wing veins dark-brown (
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E, 3F); male S6 with apicomedian spine [as in fig.
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] .. ..................................................................................................................................................................................... 5
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- Wing veins light-yellow [as in fig.
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]; male S6 apicomedially obtusely angulate, not spinose [as in fig.
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] ...........................................
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5 Malar area ~1.5x longer than wide (
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C); metepisternum nearly glabrous (
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E); medioventral process on gonocoxa conical and strongly developed (
<figureCitation id="389D2A38FFDF12157CE5FAD89EA3F95A" box="[656,718,1680,1703]" captionStart="FIGURE 3" captionStartId="5.[151,244,1823,1845]" captionTargetBox="[151,1436,432,1782]" captionTargetId="figure@5.[151,1436,432,1782]" captionTargetPageId="5" captionText="FIGURE 3. Figures illustrating characters used in the taxonomic key. A B: dorsal view of genital capsule; (A) medioventral process on gonocoxa conical and strongly developed in Geodiscelis antiminera sp. nov., (B) rounded and weakly developed medioventral process of gonocoxa in G. nazcalinea. C D: lateral view of head; (C) supraclypeal area flat in G. antiminera sp. nov., (D) convex and protuberant in G. thaumaskelos. E F: lateral view of mesosoma; (E) metepisternum glabrous in G. antiminera sp. nov., (F) densely pilose in G. nazcalinea." httpUri="https://zenodo.org/record/237328/files/figure.png" pageId="4" pageNumber="297">Fig. 3</figureCitation>
A) ....................................................
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Ferrari
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- Malar area
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longer than wide [as in fig.
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]; metepisternum densely pilose (
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F); medioventral process on gonocoxa rounded and weakly developed (
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B) .............................................................
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...........................................................................................................................
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<emphasis id="92D2EAAFFFDF12157EB2FB639D2EF8BF" bold="true" box="[199,323,1835,1858]" pageId="4" pageNumber="297">Phylogeny.</emphasis>
The reanalysis of the morphology-based data matrix of 68 characters by
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, with the inclusion of the species described herein, resulted in a unique, completely dichotomous phylogenetic tree with the same topology of that obtained by those authors (
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). Tree length (L), consistency (CI) and retention (RI) indexes are 158, 0.703 and 0.801, respectively. Overall, group support was reasonably high for most clades. The genus
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<emphasis id="92D2EAAFFFDF12157B6CFBDA99F8F854" box="[1305,1429,1938,1961]" italics="true" pageId="4" pageNumber="297">Geodiscelis</emphasis>
</taxonomicName>
, as well as its only non-monotypic subgenus,
<emphasis id="92D2EAAFFFDF12157C0BFBFC9F71F836" box="[638,796,1972,1995]" italics="true" pageId="4" pageNumber="297">Nazcoediscelis</emphasis>
, are both monophyletic. Within
<taxonomicName id="67A64D3EFFDF12157A09FBFC9895F836" ID-CoL="4MWS" box="[1148,1272,1972,1995]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="genus">
<emphasis id="92D2EAAFFFDF12157A09FBFC9895F836" box="[1148,1272,1972,1995]" italics="true" pageId="4" pageNumber="297">Geodiscelis</emphasis>
</taxonomicName>
, the subgenera
<emphasis id="92D2EAAFFFDF12157EE2FB9E9D58F810" box="[151,309,2006,2029]" italics="true" pageId="4" pageNumber="297">Nazcoediscelis</emphasis>
and
<emphasis id="92D2EAAFFFDF12157F1CFB9E9E49F810" box="[361,548,2006,2029]" italics="true" pageId="4" pageNumber="297">
<taxonomicName id="67A64D3EFFDF12157F1CFB9E9D88F810" ID-CoL="4MWS" box="[361,485,2006,2029]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="297" phylum="Arthropoda" rank="genus" sensu="stricto">Geodiscelis</taxonomicName>
<taxonomicNameLabel id="89E157D4FFDF12157F99FB9E9E49F810" box="[492,548,2006,2029]" pageId="4" pageNumber="297" sensu="stricto">s. str.</taxonomicNameLabel>
</emphasis>
are sister-groups, and together they comprise the sister-group of
<emphasis id="92D2EAAFFFDF12157AAFFB9E99E0F810" box="[1242,1421,2006,2029]" italics="true" pageId="4" pageNumber="297">Thaumoediscelis</emphasis>
.
</paragraph>
<paragraph id="A01936BDFFDE12147EB2FCD09FB6FEA2" blockId="5.[151,1436,152,351]" pageId="5" pageNumber="298">
The results of the phylogenetic analysis strongly suggest that
<taxonomicName id="67A64D3EFFDE12147DF1FCD09876FF52" box="[900,1051,152,175]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFDE12147DF1FCD09876FF52" box="[900,1051,152,175]" italics="true" pageId="5" pageNumber="298">G. antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFDE12147A52FCD098EFFF52" box="[1063,1154,152,175]" pageId="5" pageNumber="298" rank="species">
<emphasis id="92D2EAAFFFDE12147A52FCD098EFFF52" bold="true" box="[1063,1154,152,175]" pageId="5" pageNumber="298">sp. nov.</emphasis>
</taxonomicNameLabel>
belongs to the subgenus
<emphasis id="92D2EAAFFFDE12147EE2FCF29D5BFF2C" box="[151,310,186,209]" italics="true" pageId="5" pageNumber="298">Nazcoediscelis</emphasis>
and is sister to all other species in the subgenus (
<figureCitation id="389D2A38FFDE12147D21FCF29FF5FF2C" box="[852,920,186,209]" captionStart="FIGURE 4" captionStartId="6.[151,244,822,844]" captionTargetBox="[151,1430,182,807]" captionTargetId="figure@6.[151,1436,179,811]" captionTargetPageId="6" captionText="FIGURE 4. Most equally parsimonious tree found (L = 158, CI = 0.703 and RI = 0.801) in the phylogenetic analysis of the species of Geodiscelis. Filled in circles represent unique apomorphies, open circles indicate homoplasious apomorphies. Small numbers above and below the circles indicate the character number and state, respectively. Large numbers above internodes are GC values from symmetric resampling, below are Bremer support values. Character and character states are those of Packer &amp; Dumesh (2014)." httpUri="https://zenodo.org/record/237329/files/figure.png" pageId="5" pageNumber="298">Fig. 4</figureCitation>
). Character states for this species are similar to those for
<taxonomicName id="67A64D3EFFDE12147E89FC959EB0FF09" authority="Packer &amp; Dumesh, 2014:285" authorityName="Packer &amp; Dumesh" authorityPageNumber="285" authorityYear="2014" box="[252,733,221,244]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="nazcalinea">
<emphasis id="92D2EAAFFFDE12147E89FC959DE1FF09" box="[252,396,221,244]" italics="true" pageId="5" pageNumber="298">G. nazcalinea</emphasis>
(
<bibRefCitation id="C4374B4CFFDE12147FEFFC959EB8FF09" author="Packer" box="[410,725,221,244]" pageId="5" pageNumber="298" refString="Packer, L. &amp; Dumesh, S. (2014) Two new species of Geodiscelis Michener and Rozen (Hymenoptera: Apoidea: Colletidae) with a phylogenetic analysis and subgeneric classification of the genus. Zootaxa, 3857 (2), 275 - 291. http: // dx. doi. org / 10.11646 / zootaxa. 3857.2.7" type="journal article" year="2014">Packer &amp; Dumesh, 2014:285</bibRefCitation>
)
</taxonomicName>
, except as follows: 24(2) malar space 1.5x the depth of mandible; 42(0) M+Cu length of 2nd abscissa subequal to that of 1st; 65(2) galeal comb with no tooth. The internal mesosomal characters 3840 could not be scored for
<taxonomicName id="67A64D3EFFDE12147C38FD6D9EB2FEC1" box="[589,735,293,316]" class="Insecta" family="Colletidae" genus="Geodiscelis" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="298" phylum="Arthropoda" rank="species" species="antiminera" status="sp. nov.">
<emphasis id="92D2EAAFFFDE12147C38FD6D9EB2FEC1" box="[589,735,293,316]" italics="true" pageId="5" pageNumber="298">G. antiminera</emphasis>
</taxonomicName>
<taxonomicNameLabel id="89E157D4FFDE12147C93FD6D9F56FEC1" box="[742,827,293,316]" pageId="5" pageNumber="298" rank="species">
<emphasis id="92D2EAAFFFDE12147C93FD6D9F56FEC1" bold="true" box="[742,827,293,316]" pageId="5" pageNumber="298">sp. nov.</emphasis>
</taxonomicNameLabel>
due to lack of additional material for dissection, whereas the loss of S7 and S8 did not allow for the scoring of characters 46 and 4956.
</paragraph>
</subSubSection>
</treatment>
</document>