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<document id="35A746EC9D20A0E06E5E99D8500A4C75" ID-DOI="10.1080/00222933.2017.1324054" ID-ISSN="1464-5262" ID-Zenodo-Dep="5181848" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628698110182" checkinUser="felipe" docAuthor="Pati, S. K., Rajesh, L., Raj, Smrithy, Sheeja, V. U., Kumar, A. Biju &amp; Sureshan, P. M." docDate="2017" docId="251287B10E67FFBCFDD2FC1E8F793554" docLanguage="en" docName="JNatHist.51.23-24.1295-1330.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 51 (23 - 24)" docSource="http://dx.doi.org/10.1080/00222933.2017.1324054" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Pilarta aroma Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan 2017, sp. nov." docType="treatment" docUuid="2A1E2FBE-4276-49AB-B3BC-D9A7F9451A07" docUuidSource="ZooBank" docVersion="4" lastPageNumber="1314" masterDocId="D92BFFC90E77FFA8FFDBFF968C6C342A" masterDocTitle="Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India" masterLastPageNumber="1330" masterPageNumber="1295" pageNumber="1310" updateTime="1699049904912" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D8A6A5B40E0226FA29FA411D90C4BD21">Karkata, a new genus of gecarcinucid freshwater crab with two new species, and four new species of Pilarta Bahir and Yeo, 2007 and Cylindrotelphusa Alcock, 1909 (Decapoda: Brachyura) from Kerala, India</mods:title>
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<mods:namePart id="86EE124EFC2639BA90BDBFDDE2AEEB87">Pati, S. K.</mods:namePart>
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<mods:namePart id="0A3B69C5AAC99614573C35716BA546AE">Rajesh, L.</mods:namePart>
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<mods:namePart id="428C0E8E9547E1F0F92A38296EF17FCD">Raj, Smrithy</mods:namePart>
</mods:name>
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<mods:namePart id="E2418E68C518E239A5ADAA52CF57C6B9">Sheeja, V. U.</mods:namePart>
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<mods:namePart id="8B6696DCB921D82FA74934D5D2146A59">Kumar, A. Biju</mods:namePart>
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<mods:namePart id="59F852C00216EFB808C9BADEE6D49A13">Sureshan, P. M.</mods:namePart>
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<mods:title id="5B36B0BD13CE3EB1F72B4675F4B1C844">Journal of Natural History</mods:title>
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<mods:date id="2E52005C030E0C613065C0F45DAB0970">2017</mods:date>
<mods:detail id="DABEB70D80D398584ED99C83B8FDA1DA" type="series">
<mods:title id="71E512C9CBA2D5416087EE2CAC262DAE">J. Nat. Hist.</mods:title>
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<mods:number id="581EE0EB4B904D897EB7220A52102BB0">2017-05-26</mods:number>
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<mods:number id="279EE6AF2BC35F2D4FFD33B65E10EDEB">51</mods:number>
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<mods:number id="9204FF875DD3AFD7837B26242287358B">23 - 24</mods:number>
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<mods:identifier id="3AFF7FE2E1DE457896695D7353651592" type="DOI">10.1080/00222933.2017.1324054</mods:identifier>
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<paragraph id="AD0436A70E67FFB8FDD2FC1E8F773788" blockId="16.[502,814,904,965]" box="[521,795,904,930]" pageId="16" pageNumber="1310">
<heading id="F64C81CB0E67FFB8FDD2FC1E8F773788" box="[521,795,904,930]" centered="true" fontSize="11" level="2" pageId="16" pageNumber="1310" reason="2">
<taxonomicName id="6ABB4D240E67FFB8FDD2FC1E8ED83788" ID-CoL="4HSGW" authority="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan, 2017" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[521,692,904,930]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="1310" phylum="Arthropoda" rank="species" species="aroma" status="sp. nov.">
<emphasis id="9FCFEAB50E67FFB8FDD2FC1E8ED83788" bold="true" box="[521,692,904,930]" italics="true" pageId="16" pageNumber="1310">Pilarta aroma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84FC57CE0E67FFB8FD65FC1F8F773788" box="[702,795,905,930]" pageId="16" pageNumber="1310" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="E5A1652C0E67FFB8FE2DFC3A8F4237EF" box="[502,814,939,965]" pageId="16" pageNumber="1310" type="description">
<paragraph id="AD0436A70E67FFB8FE2DFC3A8F4237EF" blockId="16.[502,814,904,965]" box="[502,814,939,965]" pageId="16" pageNumber="1310">
(
<figureCitation id="35802A220E67FFB8FE26FC3A8E0637EF" box="[509,618,940,965]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="16" pageNumber="1310">Figures 7</figureCitation>
(ai) and 8(ad))
</paragraph>
</subSubSection>
<subSubSection id="E5A1652C0E67FFB8FF7BFC678F5C309D" pageId="16" pageNumber="1310" type="materials_examined">
<paragraph id="AD0436A70E67FFB8FF7BFC678D373020" blockId="16.[160,1155,1009,1207]" box="[160,347,1009,1034]" pageId="16" pageNumber="1310">
<emphasis id="9FCFEAB50E67FFB8FF7BFC678D373020" bold="true" box="[160,347,1009,1034]" italics="true" pageId="16" pageNumber="1310">Type specimens</emphasis>
</paragraph>
<paragraph id="AD0436A70E67FFB8FF7BFB838F0D3058" blockId="16.[160,1155,1009,1207]" pageId="16" pageNumber="1310">
<materialsCitation id="1DD33CFA0E67FFB8FF7BFB838F0D3058" collectingDate="2015-02-01" collectorName="Smrithy Raj" country="India" elevation="1784" latitude="8.616" location="Pongalappara Agasthyamala" longLatPrecision="77" longitude="77.246" municipality="Thiruvananthapuram district" pageId="16" pageNumber="1310" specimenCount="1" stateProvince="Kerala" typeStatus="holotype">
<typeStatus id="720088050E67FFB8FF7BFB838D653007" box="[160,265,1045,1069]" pageId="16" pageNumber="1310" type="holotype">Holotype</typeStatus>
: male (cw
<quantity id="6A439B420E67FFB8FE52FB838D933007" box="[393,511,1045,1069]" metricMagnitude="-2" metricUnit="m" metricValue="1.907" pageId="16" pageNumber="1310" unit="mm" value="19.07">19.07 mm</quantity>
, cl
<quantity id="6A439B420E67FFB8FDF3FB838EF23007" box="[552,670,1045,1069]" metricMagnitude="-2" metricUnit="m" metricValue="1.353" pageId="16" pageNumber="1310" unit="mm" value="13.53">13.53 mm</quantity>
, ch
<quantity id="6A439B420E67FFB8FD0BFB838F543007" box="[720,824,1045,1069]" metricMagnitude="-3" metricUnit="m" metricValue="8.36" pageId="16" pageNumber="1310" unit="mm" value="8.36">8.36 mm</quantity>
, fw
<quantity id="6A439B420E67FFB8FCB0FB838FBF3007" box="[875,979,1045,1069]" metricMagnitude="-3" metricUnit="m" metricValue="5.39" pageId="16" pageNumber="1310" unit="mm" value="5.39">5.39 mm</quantity>
),
<location id="A864607C0E67FFB8FC33FB838D2D3065" LSID="urn:lsid:plazi:treatment:251287B10E67FFBCFDD2FC1E8F793554:A864607C0E67FFB8FC33FB838D2D3065" country="India" latitude="8.616" longLatPrecision="77" longitude="77.246" municipality="Thiruvananthapuram district" name="Pongalappara Agasthyamala" pageId="16" pageNumber="1310" stateProvince="Kerala">Pongalappara Agasthyamala</location>
,
<collectingMunicipality id="4D60ACDD0E67FFB8FE8AFBA18EF63065" box="[337,666,1079,1103]" pageId="16" pageNumber="1310">Thiruvananthapuram district</collectingMunicipality>
,
<collectingRegion id="6F7FF8450E67FFB8FD70FBA18E9E3065" box="[683,754,1079,1103]" country="India" name="Kerala" pageId="16" pageNumber="1310">Kerala</collectingRegion>
,
<collectingCountry id="D5AC76370E67FFB8FCD8FBA18F563065" box="[771,826,1079,1103]" name="India" pageId="16" pageNumber="1310">India</collectingCountry>
(
<geoCoordinate id="C88F50600E67FFB8FC94FBA18FC53065" box="[847,937,1079,1103]" degrees="8.616" direction="north" orientation="latitude" pageId="16" pageNumber="1310" precision="55" value="8.616">8.616°N</geoCoordinate>
,
<geoCoordinate id="C88F50600E67FFB8FC61FBA188703065" box="[954,1052,1079,1103]" degrees="77.246" direction="east" orientation="longitude" pageId="16" pageNumber="1310" precision="55" value="77.246">77.246°E</geoCoordinate>
; altitude
<quantity id="6A439B420E67FFB8FF7BFBCC8C943058" box="[160,248,1114,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="16" pageNumber="1310" unit="m" value="1784.0">
<elevation id="2696D1940E67FFB8FF7BFBCC8C943058" box="[160,248,1114,1138]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="16" pageNumber="1310" unit="m" value="1784.0">1784 m</elevation>
</quantity>
),
<date id="D90510670E67FFB8FED7FBCC8DAB3058" box="[268,455,1114,1138]" pageId="16" pageNumber="1310" value="2015-02-01">
<collectingDate id="C941E98F0E67FFB8FED7FBCC8DAB3058" box="[268,455,1114,1138]" pageId="16" pageNumber="1310" value="2015-02-01">1 February 2015</collectingDate>
</date>
, coll.
<collectorName id="004E53710E67FFB8FDD2FBCC8EE63058" box="[521,650,1114,1138]" pageId="16" pageNumber="1310">Smrithy Raj</collectorName>
(ZSI, WRC-C.1529).
</materialsCitation>
</paragraph>
<paragraph id="AD0436A70E67FFB8FF1BFBEB8F5C309D" blockId="16.[160,1155,1009,1207]" pageId="16" pageNumber="1310">
<typeStatus id="720088050E67FFB8FF1BFBEB8D5C30BF" box="[192,304,1149,1173]" pageId="16" pageNumber="1310" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="BBBDFD2E0E67FFB8FEE5FBEB8DF430BF" box="[318,408,1149,1173]" pageId="16" pageNumber="1310" type="male">2 males</specimenCount>
(cw
<quantity id="6A439B420E67FFB8FE0EFBEB8EF930BF" box="[469,661,1148,1173]" metricMagnitude="-2" metricUnit="m" metricValue="1.8975" metricValueMax="1.972" metricValueMin="1.823" pageId="16" pageNumber="1310" unit="mm" value="18.975" valueMax="19.72" valueMin="18.23">18.2319.72 mm</quantity>
, cl
<quantity id="6A439B420E67FFB8FD1AFBEB8FED30BF" box="[705,897,1148,1173]" metricMagnitude="-2" metricUnit="m" metricValue="1.4075" metricValueMax="1.434" metricValueMin="1.381" pageId="16" pageNumber="1310" unit="mm" value="14.075" valueMax="14.34" valueMin="13.81">13.8114.34 mm</quantity>
, ch
<quantity id="6A439B420E67FFB8FC6EFBEB883630BF" box="[949,1114,1148,1173]" metricMagnitude="-3" metricUnit="m" metricValue="8.66" metricValueMax="9.27" metricValueMin="8.05" pageId="16" pageNumber="1310" unit="mm" value="8.66" valueMax="9.27" valueMin="8.05">8.059.27 mm</quantity>
, fw
<quantity id="6A439B420E67FFB8FF7BFB098D2E309D" box="[160,322,1183,1207]" metricMagnitude="-3" metricUnit="m" metricValue="5.515" metricValueMax="5.53" metricValueMin="5.5" pageId="16" pageNumber="1310" unit="mm" value="5.515" valueMax="5.53" valueMin="5.5">5.505.53 mm</quantity>
), same data as holotype (ZSI, WRC-C.1530).
</paragraph>
</subSubSection>
<subSubSection id="E5A1652C0E67FFB9FF7BFB728DF03250" lastPageId="17" lastPageNumber="1311" pageId="16" pageNumber="1310" type="diagnosis">
<paragraph id="AD0436A70E67FFB8FF7BFB728D7830D7" blockId="16.[160,1156,1252,1727]" box="[160,276,1252,1277]" pageId="16" pageNumber="1310">
<emphasis id="9FCFEAB50E67FFB8FF7BFB728D7830D7" bold="true" box="[160,276,1252,1277]" italics="true" pageId="16" pageNumber="1310">Diagnosis</emphasis>
</paragraph>
<paragraph id="AD0436A70E67FFB9FF7BFA918DF03250" blockId="16.[160,1156,1252,1727]" lastBlockId="17.[160,1155,1530,1658]" lastPageId="17" lastPageNumber="1311" pageId="16" pageNumber="1310">
Carapace broader than long (cw/cl = ca. 1.31.4), sparsely punctate; dorsal surface convex in frontal view, almost smooth; anterolateral and posterolateral margins glabrous; front slightly broad (fw/cw = ca. 0.3); epigastric cristae visible as 2 low, broad, almost smooth protuberances; postorbital cristae not well developed, smooth; external orbital angle broadly triangular, with long outer margin, ca. 3 times length of inner margin; epibranchial tooth very small, blunt; branchial regions inflated, lacking striae; cervical grooves distinct, broad; H-shaped groove deep; epistome posterior margin with rounded median lobe, distinctly sinuous lateral lobes (
<figureCitation id="35802A220E67FFB8FCC5FA6C8F133238" box="[798,895,1530,1554]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="16" pageNumber="1310">Figure 7</figureCitation>
(a,b)). Ambulatory legs with dense setae on external surfaces of dactylus, propodus and carpus (
<figureCitation id="35802A220E67FFB8FC35F98B8822321F" box="[1006,1102,1565,1589]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="16" pageNumber="1310">Figure 7</figureCitation>
(a,f)). Sutures between male thoracic sternites s2/s3 and s3/s4 undiscernible as grooves (
<figureCitation id="35802A220E67FFB8FF73F9F48D603250" box="[168,268,1634,1658]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="16" pageNumber="1310">Figure 7</figureCitation>
(c,g)). Male abdomen broadly T-shaped; sixth abdominal somite distinctly longer than telson; telson broader than long (
<figureCitation id="35802A220E67FFB8FD63F9138F7B32B7" box="[696,791,1669,1693]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="16" pageNumber="1310">Figure 7</figureCitation>
(c,h)). G1 terminal segment less stout, slightly long, ca. 0.4 times length of subterminal segment, with almost straight outer margin, tip distinctly narrow, straight or slightly curved outwards; subterminal segment with broad median portion, not as broad as widest part of base, outer margin almost straight (
<figureCitation id="35802A220E66FFB9FE8CF9A98DD9327D" box="[343,437,1599,1623]" captionStart="Figure 8" captionStartId="16.[160,225,748,771]" captionTargetBox="[178,1137,142,716]" captionTargetId="figure-248@16.[178,1138,142,717]" captionTargetPageId="16" captionText="Figure 8. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view of left G1; (b) dorsal view of left G1 terminal segment; (c) ventral view of left G1; (d) left G2. Scale bars: a, c, d = 1 mm; b = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5181864" httpUri="https://zenodo.org/record/5181864/files/figure.png" pageId="17" pageNumber="1311">Figure 8</figureCitation>
(ac)). G2 with long distal segment, ca. 0.5 times length of basal segment (
<figureCitation id="35802A220E66FFB9FEC9F9F48D1C3250" box="[274,368,1634,1658]" captionStart="Figure 8" captionStartId="16.[160,225,748,771]" captionTargetBox="[178,1137,142,716]" captionTargetId="figure-248@16.[178,1138,142,717]" captionTargetPageId="16" captionText="Figure 8. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view of left G1; (b) dorsal view of left G1 terminal segment; (c) ventral view of left G1; (d) left G2. Scale bars: a, c, d = 1 mm; b = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5181864" httpUri="https://zenodo.org/record/5181864/files/figure.png" pageId="17" pageNumber="1311">Figure 8</figureCitation>
(d)).
</paragraph>
</subSubSection>
<caption id="F9C4662F0E66FFB9FF7BFAD28F2C319E" ID-DOI="http://doi.org/10.5281/zenodo.5181866" ID-Zenodo-Dep="5181866" httpUri="https://zenodo.org/record/5181866/files/figure.png" pageId="17" pageNumber="1311" startId="17.[160,225,1348,1371]" targetBox="[178,1138,142,1318]" targetPageId="17">
<paragraph id="AD0436A70E66FFB9FF7BFAD28F2C319E" blockId="17.[160,1155,1348,1460]" pageId="17" pageNumber="1311">
<emphasis id="9FCFEAB50E66FFB9FF7BFAD28C973171" bold="true" box="[160,251,1348,1371]" pageId="17" pageNumber="1311">Figure 9.</emphasis>
<taxonomicName id="6ABB4D240E66FFB9FED8FAD28DA23171" authority="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan, 2017" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[259,462,1348,1371]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="1311" phylum="Arthropoda" rank="species" species="punctatissima" status="sp. nov.">
<emphasis id="9FCFEAB50E66FFB9FED8FAD28DA23171" box="[259,462,1348,1371]" italics="true" pageId="17" pageNumber="1311">Pilarta punctatissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84FC57CE0E66FFB9FE0EFAD38E493176" box="[469,549,1349,1372]" pageId="17" pageNumber="1311" rank="species">sp. nov.</taxonomicNameLabel>
, holotype male (ZSI, WRC-C.1531). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) right or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm.
</paragraph>
</caption>
<subSubSection id="E5A1652C0E65FFBAFF7BFF1B8F8532AB" pageId="18" pageNumber="1312" type="description">
<paragraph id="AD0436A70E65FFBAFF7BFF1B8D90348C" blockId="18.[160,1156,141,1275]" box="[160,508,141,166]" pageId="18" pageNumber="1312">
<emphasis id="9FCFEAB50E65FFBAFF7BFF1B8D90348C" bold="true" box="[160,508,141,166]" italics="true" pageId="18" pageNumber="1312">
Description of male
<typeStatus id="720088050E65FFBAFE4FFF1B8D90348C" box="[404,508,141,166]" pageId="18" pageNumber="1312" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="AD0436A70E65FFBAFF7BFF278870366C" blockId="18.[160,1156,141,1275]" pageId="18" pageNumber="1312">
Carapace broader than long (cw/cl = 1.4), sparsely punctate; dorsal surface convex in frontal view, smooth; anterolateral surface slightly inflated in frontal view; anterolateral and posterolateral margins glabrous; frontal margin bilobed, slightly broad (fw/cw = 0.3); epigastric cristae visible as 2 low, broad, smooth protuberances; postorbital cristae not well developed, smooth; external orbital angle broadly triangular, outer margin long, ca. 3 times length of inner margin, anterior half of outer margin concave; epibranchial tooth very small, blunt, separated from external orbital angle by distinct cleft; postorbital region gently concave; branchial regions inflated, lacking striae; cervical grooves distinct, broad; epigastric groove short, broad; mesogastric groove shallow, long, broad, bifurcated posteriorly; H-shaped groove distinct, deep; frontal median triangle incomplete, with cristate dorsal margin only, lateral margins not developed; epistome posterior margin with prominent, rounded median lobe, distinctly sinuous lateral lobes (
<figureCitation id="35802A220E65FFBAFC58FDB88F8C366C" box="[899,992,558,582]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="18" pageNumber="1312">Figure 7</figureCitation>
(a,b)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF1BFDC78ED03684" blockId="18.[160,1156,141,1275]" pageId="18" pageNumber="1312">
Chelipeds unequal, left chela larger; fingers of larger cheliped gaping when fingers closed; outer surface of carpus rugose; major tooth of carpus large, broad, blunt; external surfaces of chelipeds glabrous (
<figureCitation id="35802A220E65FFBAFDD3FD008E0B3684" box="[520,615,662,686]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="18" pageNumber="1312">Figure 7</figureCitation>
(a,ce)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF1BFD2F8F8336D9" blockId="18.[160,1156,141,1275]" pageId="18" pageNumber="1312">
Ambulatory legs long, moderately stout, with distinct setae; p3 longest; external surfaces of dactylus, propodus and carpus with dense setae (
<figureCitation id="35802A220E65FFBAFC80FD4D8FD636D9" box="[859,954,731,755]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="18" pageNumber="1312">Figure 7</figureCitation>
(a,f)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF1BFD6888E8373C" blockId="18.[160,1156,141,1275]" box="[192,1156,766,790]" pageId="18" pageNumber="1312">
Sutures between thoracic sternites s2/s3 and s3/s4 undiscernible as grooves (
<figureCitation id="35802A220E65FFBAFC28FD688826373C" box="[1011,1098,766,790]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="18" pageNumber="1312">Figure 7</figureCitation>
(c,g)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF1BFCB78F5C378B" blockId="18.[160,1156,141,1275]" pageId="18" pageNumber="1312">
Abdomen broadly T-shaped; fifth abdominal somite distinctly broader than long; sixth abdominal somite broader than long, distinctly longer than telson; telson broader than long, apex round; sternoabdominal cavity deep, long, extending up to imaginary line joining median part of cheliped coxae (
<figureCitation id="35802A220E65FFBAFD4DFC1F8E98378B" box="[662,756,905,929]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="18" pageNumber="1312">Figure 7</figureCitation>
(c,h)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF1BFC3D8E873064" blockId="18.[160,1156,141,1275]" pageId="18" pageNumber="1312">
Mandibular palp with 2 segments, terminal segment bilobed. First, second maxillipeds each with long flagellum on exopod. Third maxillipeds cover most of buccal cavity when closed; ischium subrectangular, longer than broad, with almost indistinct median sulcus; merus subrectangular, broader than long; exopod longer than ischium, reaching lower third of merus, lacking flagellum (
<figureCitation id="35802A220E65FFBAFDB1FBA08EA43064" box="[618,712,1078,1102]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="18" pageNumber="1312">Figure 7</figureCitation>
(i)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF1BFBCF8E8A30D1" blockId="18.[160,1156,141,1275]" pageId="18" pageNumber="1312">
G1 stout, straight; terminal segment less stout, slightly long, ca. 0.4 times length of subterminal segment, with almost straight outer margin, tip distinctly narrow, slightly curved outwards; subterminal segment with broad median portion, not as broad as widest part of base, outer margin straight (
<figureCitation id="35802A220E65FFBAFD4BFB578E8330F3" box="[656,751,1217,1241]" captionStart="Figure 8" captionStartId="16.[160,225,748,771]" captionTargetBox="[178,1137,142,716]" captionTargetId="figure-248@16.[178,1138,142,717]" captionTargetPageId="16" captionText="Figure 8. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view of left G1; (b) dorsal view of left G1 terminal segment; (c) ventral view of left G1; (d) left G2. Scale bars: a, c, d = 1 mm; b = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5181864" httpUri="https://zenodo.org/record/5181864/files/figure.png" pageId="18" pageNumber="1312">Figure 8</figureCitation>
(ac)). G2 with long distal segment, ca. 0.5 times length of basal segment (
<figureCitation id="35802A220E65FFBAFD86FB758ED730D1" box="[605,699,1251,1275]" captionStart="Figure 8" captionStartId="16.[160,225,748,771]" captionTargetBox="[178,1137,142,716]" captionTargetId="figure-248@16.[178,1138,142,717]" captionTargetPageId="16" captionText="Figure 8. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view of left G1; (b) dorsal view of left G1 terminal segment; (c) ventral view of left G1; (d) left G2. Scale bars: a, c, d = 1 mm; b = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5181864" httpUri="https://zenodo.org/record/5181864/files/figure.png" pageId="18" pageNumber="1312">Figure 8</figureCitation>
(d)).
</paragraph>
<paragraph id="AD0436A70E65FFBAFF7BFAAB8D4E317C" blockId="18.[160,794,1341,1401]" box="[160,290,1341,1366]" pageId="18" pageNumber="1312">
<emphasis id="9FCFEAB50E65FFBAFF7BFAAB8D4E317C" bold="true" box="[160,290,1341,1366]" italics="true" pageId="18" pageNumber="1312">Live colour</emphasis>
</paragraph>
<paragraph id="AD0436A70E65FFBAFF7BFAF78F763153" blockId="18.[160,794,1341,1401]" box="[160,794,1377,1401]" pageId="18" pageNumber="1312">Carapace, chelipeds and ambulatory legs are all brown.</paragraph>
<paragraph id="AD0436A70E65FFBAFF7BFA2D8D7931FE" blockId="18.[160,1156,1467,1665]" box="[160,277,1467,1492]" pageId="18" pageNumber="1312">
<emphasis id="9FCFEAB50E65FFBAFF7BFA2D8D7931FE" bold="true" box="[160,277,1467,1492]" italics="true" pageId="18" pageNumber="1312">
<typeStatus id="720088050E65FFBAFF7BFA2D8D7931FE" box="[160,277,1467,1492]" pageId="18" pageNumber="1312" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="AD0436A70E65FFBAFF7BFA488F8532AB" blockId="18.[160,1156,1467,1665]" pageId="18" pageNumber="1312">
The male
<typeStatus id="720088050E65FFBAFECFFA488DEE31DC" box="[276,386,1502,1526]" pageId="18" pageNumber="1312" type="paratype">paratypes</typeStatus>
(ZSI, WRC-C.1530) resemble the
<typeStatus id="720088050E65FFBAFD24FA488F0F31DC" box="[767,867,1502,1526]" pageId="18" pageNumber="1312" type="holotype">holotype</typeStatus>
in their overall carapace and gonopod characters. Nevertheless, the
<typeStatus id="720088050E65FFBAFD46F9978F673233" box="[669,779,1537,1561]" pageId="18" pageNumber="1312" type="paratype">paratypes</typeStatus>
show the following differences: epigastric cristae that are slightly rugose in the smaller male (vs smooth); a median sulcus on the third maxilliped ischium that is distinct (vs indistinct); and a G1 tip that is straight in the smaller male (vs slightly curved outwards in the
<typeStatus id="720088050E65FFBAFCACF9FF8FB332AB" box="[887,991,1641,1665]" pageId="18" pageNumber="1312" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="E5A1652C0E64FFBBFF7BFF188F7F34C6" pageId="19" pageNumber="1313" type="etymology">
<paragraph id="AD0436A70E64FFBBFF7BFF188D71348D" blockId="19.[160,1155,142,236]" box="[160,285,142,167]" pageId="19" pageNumber="1313">
<emphasis id="9FCFEAB50E64FFBBFF7BFF188D71348D" bold="true" box="[160,285,142,167]" italics="true" pageId="19" pageNumber="1313">Etymology</emphasis>
</paragraph>
<paragraph id="AD0436A70E64FFBBFF7BFF248F7F34C6" blockId="19.[160,1155,142,236]" pageId="19" pageNumber="1313">Aroma means hairless in Sanskrit. Used as a noun in apposition, alluding to the smooth or hairless carapace and chelipeds of the crab.</paragraph>
</subSubSection>
<subSubSection id="E5A1652C0E64FFBBFF7BFEBC8DBC35A2" pageId="19" pageNumber="1313" type="materials_examined">
<paragraph id="AD0436A70E64FFBBFF7BFEBC8D5B3569" blockId="19.[160,1156,298,392]" box="[160,311,298,323]" pageId="19" pageNumber="1313">
<emphasis id="9FCFEAB50E64FFBBFF7BFEBC8D5B3569" bold="true" box="[160,311,298,323]" italics="true" pageId="19" pageNumber="1313">Type locality</emphasis>
</paragraph>
<paragraph id="AD0436A70E64FFBBFF7BFED88DBC35A2" blockId="19.[160,1156,298,392]" pageId="19" pageNumber="1313">
<materialsCitation id="1DD33CFA0E64FFBBFF7BFED88DA035A2" country="India" county="Thiruvananthapuram district" elevation="1784" latitude="8.616" location="Pongalappara" longLatPrecision="77" longitude="77.246" municipality="Agasthyamala" pageId="19" pageNumber="1313" specimenCount="1" stateProvince="Kerala">
<location id="A864607C0E64FFBBFF7BFED88D52354C" LSID="urn:lsid:plazi:treatment:251287B10E67FFBCFDD2FC1E8F793554:A864607C0E64FFBBFF7BFED88D52354C" box="[160,318,334,358]" country="India" county="Thiruvananthapuram district" latitude="8.616" longLatPrecision="77" longitude="77.246" municipality="Agasthyamala" name="Pongalappara" pageId="19" pageNumber="1313" stateProvince="Kerala">Pongalappara</location>
,
<collectingMunicipality id="4D60ACDD0E64FFBBFE8DFED88D9B354C" box="[342,503,334,358]" pageId="19" pageNumber="1313">Agasthyamala</collectingMunicipality>
,
<collectingCounty id="44654E2B0E64FFBBFDD5FED88F31354C" box="[526,861,334,358]" pageId="19" pageNumber="1313">Thiruvananthapuram district</collectingCounty>
,
<collectingRegion id="6F7FF8450E64FFBBFCAEFED88FD0354C" box="[885,956,334,358]" country="India" name="Kerala" pageId="19" pageNumber="1313">Kerala</collectingRegion>
,
<collectingCountry id="D5AC76370E64FFBBFC0FFED88867354C" box="[980,1035,334,358]" name="India" pageId="19" pageNumber="1313">India</collectingCountry>
(
<geoCoordinate id="C88F50600E64FFBBFBFDFED888EC354C" box="[1062,1152,334,358]" degrees="8.616" direction="north" orientation="latitude" pageId="19" pageNumber="1313" precision="55" value="8.616">8.616°N</geoCoordinate>
,
<geoCoordinate id="C88F50600E64FFBBFF7BFEE68D6E35A2" box="[160,258,368,392]" degrees="77.246" direction="east" orientation="longitude" pageId="19" pageNumber="1313" precision="55" value="77.246">77.246°E</geoCoordinate>
; altitude
<quantity id="6A439B420E64FFBBFEB6FEE68DA835A2" box="[365,452,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="19" pageNumber="1313" unit="m" value="1784.0">
<elevation id="2696D1940E64FFBBFEB6FEE68DA835A2" box="[365,452,368,392]" metricMagnitude="3" metricUnit="m" metricValue="1.784" pageId="19" pageNumber="1313" unit="m" value="1784.0">1784 m</elevation>
</quantity>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E5A1652C0E64FFBCFF7BFE508F793554" lastPageId="20" lastPageNumber="1314" pageId="19" pageNumber="1313" type="discussion">
<paragraph id="AD0436A70E64FFBBFF7BFE508D6A35F5" blockId="19.[160,1156,454,1727]" box="[160,262,454,479]" pageId="19" pageNumber="1313">
<emphasis id="9FCFEAB50E64FFBBFF7BFE508D6A35F5" bold="true" box="[160,262,454,479]" italics="true" pageId="19" pageNumber="1313">Remarks</emphasis>
</paragraph>
<paragraph id="AD0436A70E64FFBBFF7BFE7C881C3006" blockId="19.[160,1156,454,1727]" pageId="19" pageNumber="1313">
The new species,
<taxonomicName id="6ABB4D240E64FFBBFEB1FE7C8E6D3628" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[362,513,490,514]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="aroma">
<emphasis id="9FCFEAB50E64FFBBFEB1FE7C8E6D3628" box="[362,513,490,514]" italics="true" pageId="19" pageNumber="1313">Pilarta aroma</emphasis>
</taxonomicName>
, is most similar to
<taxonomicName id="6ABB4D240E64FFBBFD07FE7C8F503628" authorityName="Bahir and Yeo" authorityYear="2007" box="[732,828,490,514]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="anuka">
<emphasis id="9FCFEAB50E64FFBBFD07FE7C8F503628" box="[732,828,490,514]" italics="true" pageId="19" pageNumber="1313">P. anuka</emphasis>
</taxonomicName>
due to its sparsely punctate carapace, glabrous anterolateral and posterolateral margins of the carapace, epistome posterior margin with distinctly sinuous lateral lobes, sixth male abdominal somite longer than telson, male telson broader than long, less stout G1 terminal segment, and comparatively short G2 distal segment, ca. 0.40.5 times the length of the basal segment (
<figureCitation id="35802A220E64FFBBFECAFD018D153685" box="[273,377,663,687]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="19" pageNumber="1313">Figures 7</figureCitation>
(ac,h) and 8(ad) for
<taxonomicName id="6ABB4D240E64FFBBFDABFD018EBF3685" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[624,723,663,687]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="aroma">
<emphasis id="9FCFEAB50E64FFBBFDABFD018EBF3685" box="[624,723,663,687]" italics="true" pageId="19" pageNumber="1313">P. aroma</emphasis>
</taxonomicName>
; see
<bibRefCitation id="C92A4B560E64FFBBFCD7FD018F843685" author="Bahir MM &amp; Yeo DCJ" box="[780,1000,663,687]" pageId="19" pageNumber="1313" pagination="309 - 354" refId="ref17281" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
, figs 25(a,d,f h) and 26(ac) for
<taxonomicName id="6ABB4D240E64FFBBFEA3FD2C8DB536F8" authorityName="Bahir and Yeo" authorityYear="2007" box="[376,473,698,722]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="anuka">
<emphasis id="9FCFEAB50E64FFBBFEA3FD2C8DB536F8" box="[376,473,698,722]" italics="true" pageId="19" pageNumber="1313">P. anuka</emphasis>
</taxonomicName>
). However,
<taxonomicName id="6ABB4D240E64FFBBFDBFFD2C8EA536F8" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[612,713,698,722]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="aroma">
<emphasis id="9FCFEAB50E64FFBBFDBFFD2C8EA536F8" box="[612,713,698,722]" italics="true" pageId="19" pageNumber="1313">P. aroma</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="6ABB4D240E64FFBBFC15FD2C884336F8" authorityName="Bahir and Yeo" authorityYear="2007" box="[974,1071,698,722]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="anuka">
<emphasis id="9FCFEAB50E64FFBBFC15FD2C884336F8" box="[974,1071,698,722]" italics="true" pageId="19" pageNumber="1313">P. anuka</emphasis>
</taxonomicName>
by the smooth postorbital cristae (
<figureCitation id="35802A220E64FFBBFE3AFD4A8E2E36DE" box="[481,578,732,756]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="19" pageNumber="1313">Figure 7</figureCitation>
(a,b)) (vs rugose postorbital cristae; see
<bibRefCitation id="C92A4B560E64FFBBFBCFFD4A8D7C373D" author="Bahir MM &amp; Yeo DCJ" pageId="19" pageNumber="1313" pagination="309 - 354" refId="ref17281" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
, figs 25(a) and 26(a,b)), branchial regions lacking striae (
<figureCitation id="35802A220E64FFBBFC15FD69885E373D" box="[974,1074,767,791]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="19" pageNumber="1313">Figure 7</figureCitation>
(a)) (vs branchial regions with distinct striae adjacent to lateral margins; see
<bibRefCitation id="C92A4B560E64FFBBFC07FCB48CBB3776" author="Bahir MM &amp; Yeo DCJ" pageId="19" pageNumber="1313" pagination="309 - 354" refId="ref17281" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
, fig. 26(a)), deep H-shaped groove (
<figureCitation id="35802A220E64FFBBFDB2FCD28EAA3776" box="[617,710,836,860]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="19" pageNumber="1313">Figure 7</figureCitation>
(a)) (vs comparatively shallow H-shaped groove; see
<bibRefCitation id="C92A4B560E64FFBBFEF7FCF18E603755" author="Bahir MM &amp; Yeo DCJ" box="[300,524,871,895]" pageId="19" pageNumber="1313" pagination="309 - 354" refId="ref17281" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
, fig. 26(a)), G1 terminal segment relatively long, ca. 0.4 times the length of the subterminal segment, with almost straight outer margin (
<figureCitation id="35802A220E64FFBBFF73FC3A8D6B37EE" box="[168,263,940,964]" captionStart="Figure 8" captionStartId="16.[160,225,748,771]" captionTargetBox="[178,1137,142,716]" captionTargetId="figure-248@16.[178,1138,142,717]" captionTargetPageId="16" captionText="Figure 8. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view of left G1; (b) dorsal view of left G1 terminal segment; (c) ventral view of left G1; (d) left G2. Scale bars: a, c, d = 1 mm; b = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5181864" httpUri="https://zenodo.org/record/5181864/files/figure.png" pageId="19" pageNumber="1313">Figure 8</figureCitation>
(ac)) (vs G1 terminal segment relatively short, ca. 0.3 times the length of the subterminal segment, with slightly convex outer margin; see
<bibRefCitation id="C92A4B560E64FFBBFC8FFC59885D37CD" author="Bahir MM &amp; Yeo DCJ" box="[852,1073,975,999]" pageId="19" pageNumber="1313" pagination="309 - 354" refId="ref17281" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
, fig. 25 (f,g)), G1 subterminal segment with almost straight outer margin (
<figureCitation id="35802A220E64FFBBFC47FC648F973020" box="[924,1019,1010,1034]" captionStart="Figure 8" captionStartId="16.[160,225,748,771]" captionTargetBox="[178,1137,142,716]" captionTargetId="figure-248@16.[178,1138,142,717]" captionTargetPageId="16" captionText="Figure 8. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view of left G1; (b) dorsal view of left G1 terminal segment; (c) ventral view of left G1; (d) left G2. Scale bars: a, c, d = 1 mm; b = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5181864" httpUri="https://zenodo.org/record/5181864/files/figure.png" pageId="19" pageNumber="1313">Figure 8</figureCitation>
(a,b)) (vs G1 subterminal segment with convex outer margin; see
<bibRefCitation id="C92A4B560E64FFBBFD25FB828FB13006" author="Bahir MM &amp; Yeo DCJ" box="[766,989,1044,1068]" pageId="19" pageNumber="1313" pagination="309 - 354" refId="ref17281" refString="Bahir MM, Yeo DCJ. 2007. The gecarcinucid freshwater crabs of southern India (Crustacea: Decapoda: Brachyura). Raffles Bull Zool. Suppl. No. 16: 309 - 354." type="book chapter" year="2007">Bahir and Yeo 2007</bibRefCitation>
, figs 25(f,g)).
</paragraph>
<paragraph id="AD0436A70E64FFBBFF1BFBA18FC23295" blockId="19.[160,1156,454,1727]" pageId="19" pageNumber="1313">
<taxonomicName id="6ABB4D240E64FFBBFF1BFBA18D3A3065" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[192,342,1079,1103]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="aroma">
<emphasis id="9FCFEAB50E64FFBBFF1BFBA18D3A3065" box="[192,342,1079,1103]" italics="true" pageId="19" pageNumber="1313">Pilarta aroma</emphasis>
</taxonomicName>
is very distinct from
<taxonomicName id="6ABB4D240E64FFBBFD99FBA18E983065" authority="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan, 2017" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[578,756,1079,1103]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="punctatissima" status="sp. nov.">
<emphasis id="9FCFEAB50E64FFBBFD99FBA18E983065" box="[578,756,1079,1103]" italics="true" pageId="19" pageNumber="1313">P. punctatissima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="84FC57CE0E64FFBBFD20FBA18F3E3065" box="[763,850,1079,1103]" pageId="19" pageNumber="1313" rank="species">sp. nov.</taxonomicNameLabel>
in features of the carapace and gonopods: carapace sparsely punctate (vs densely punctate), dorsal surface of the carapace almost smooth (vs smooth except epigastric and postorbital cristae, postorbital region, frontal region and lateral margins), anterolateral and posterolateral margins of the carapace glabrous (vs with dense setae), postorbital cristae smooth (vs rugose), external orbital angle with comparatively short outer margin, ca. 3 times the length of the inner margin (vs with comparatively long outer margin, ca. 4 times the length of the inner margin), branchial regions lacking striae (vs with distinct striae adjacent to lateral margins), H-shaped groove deep (vs relatively shallow), epistome posterior margin with rounded median lobe, distinctly sinuous lateral lobes (vs with broadly triangular median lobe, medially notched lateral lobes), sixth male abdominal somite longer than telson (vs slightly shorter than telson), male telson broader than long (vs as long as broad), G1 terminal segment less stout, relatively long, ca. 0.4 times the length of the subterminal segment, with almost straight outer margin (vs more stout, relatively short, ca. 0.3 times the length of the subterminal segment, with strongly convex outer margin), G1 subterminal segment with almost straight outer margin (vs with convex outer margin), G2 distal segment relatively short, ca. 0.5 times the length of the basal segment (vs relatively long, ca. 0.6 times the length of the basal segment) (
<figureCitation id="35802A220E64FFBBFC59F9128F8132B6" box="[898,1005,1668,1692]" captionStart="Figure 7" captionStartId="15.[160,225,1279,1302]" captionTargetBox="[178,1138,142,1248]" captionTargetId="figure-54@15.[178,1138,142,1248]" captionTargetPageId="15" captionText="Figure 7. Pilarta aroma sp. nov., holotype male (ZSI, WRC-C.1529). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) left or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181862" httpUri="https://zenodo.org/record/5181862/files/figure.png" pageId="19" pageNumber="1313">Figures 7</figureCitation>
(ac,h) and 8 (ad) for
<taxonomicName id="6ABB4D240E64FFBBFED2F9318D013295" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[265,365,1703,1727]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="aroma">
<emphasis id="9FCFEAB50E64FFBBFED2F9318D013295" box="[265,365,1703,1727]" italics="true" pageId="19" pageNumber="1313">P. aroma</emphasis>
</taxonomicName>
;
<figureCitation id="35802A220E64FFBBFEA0F9318D883295" box="[379,484,1703,1727]" captionStart="Figure 9" captionStartId="17.[160,225,1348,1371]" captionTargetBox="[178,1138,142,1318]" captionTargetId="figure-51@17.[178,1138,142,1318]" captionTargetPageId="17" captionText="Figure 9. Pilarta punctatissima sp. nov., holotype male (ZSI, WRC-C.1531). (a) Dorsal view; (b) frontal view; (c) ventral view; (d) right or major cheliped; (e) carpus of major cheliped; (f) right ambulatory leg (p3); (g) anterior thoracic sternites (s1s4); (h) abdominal somites 46 and telson; (i) left third maxilliped. Scale bars: ac = 10 mm; d, f = 5 mm; e, gi = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5181866" httpUri="https://zenodo.org/record/5181866/files/figure.png" pageId="19" pageNumber="1313">Figures 9</figureCitation>
(ac,h) and 10(ad) for
<taxonomicName id="6ABB4D240E64FFBBFD35F9318FCE3295" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[750,930,1703,1727]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="19" pageNumber="1313" phylum="Arthropoda" rank="species" species="punctatissima">
<emphasis id="9FCFEAB50E64FFBBFD35F9318FCE3295" box="[750,930,1703,1727]" italics="true" pageId="19" pageNumber="1313">P. punctatissima</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="AD0436A70E63FFBCFF7BFF188D0D348D" blockId="20.[160,1156,142,236]" box="[160,353,142,167]" pageId="20" pageNumber="1314">
<emphasis id="9FCFEAB50E63FFBCFF7BFF188D0D348D" bold="true" box="[160,353,142,167]" italics="true" pageId="20" pageNumber="1314">Ecological notes</emphasis>
</paragraph>
<paragraph id="AD0436A70E63FFBCFF7BFF248F1834C6" blockId="20.[160,1156,142,236]" pageId="20" pageNumber="1314">
Crabs were collected from underneath cobblestones in a high-altitude mountain stream (
<quantity id="6A439B420E63FFBCFF7CFF428CB834C6" box="[167,212,212,236]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="1314" unit="m" value="2.0">2 m</quantity>
wide, up to
<quantity id="6A439B420E63FFBCFEB2FF428DF934C6" box="[361,405,212,236]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="1314" unit="m" value="1.0">1 m</quantity>
deep) with clear, cold water (10 to 20°C).
</paragraph>
<paragraph id="AD0436A70E63FFBCFF7BFEB68DD73513" blockId="20.[160,1155,288,382]" box="[160,443,288,313]" pageId="20" pageNumber="1314">
<emphasis id="9FCFEAB50E63FFBCFF7BFEB68DD73513" bold="true" box="[160,443,288,313]" italics="true" pageId="20" pageNumber="1314">Geographic distribution</emphasis>
</paragraph>
<paragraph id="AD0436A70E63FFBCFF7BFED58F793554" blockId="20.[160,1155,288,382]" pageId="20" pageNumber="1314">
<materialsCitation id="1DD33CFA0E63FFBCFF7BFED58F7C3554" location="Thiruvananthapuram district" municipality="Western Ghats" pageId="20" pageNumber="1314" specimenCount="1" stateProvince="Kerala">
The new species,
<taxonomicName id="6ABB4D240E63FFBCFEB3FED58D923571" authorityName="Pati &amp; Rajesh &amp; Raj &amp; Sheeja &amp; Kumar &amp; Sureshan" authorityYear="2017" box="[360,510,323,347]" class="Malacostraca" family="Gecarcinucidae" genus="Pilarta" kingdom="Animalia" order="Decapoda" pageId="20" pageNumber="1314" phylum="Arthropoda" rank="species" species="aroma">
<emphasis id="9FCFEAB50E63FFBCFEB3FED58D923571" box="[360,510,323,347]" italics="true" pageId="20" pageNumber="1314">Pilarta aroma</emphasis>
</taxonomicName>
, is known only from the
<collectingMunicipality id="4D60ACDD0E63FFBCFCC0FED58FD33571" box="[795,959,323,347]" pageId="20" pageNumber="1314">Western Ghats</collectingMunicipality>
at Agasthyamala (
<typeStatus id="720088050E63FFBCFF7DFEF08CB53554" box="[166,217,358,382]" pageId="20" pageNumber="1314">type</typeStatus>
locality) in
<location id="A864607C0E63FFBCFEBBFEF08ECD3554" LSID="urn:lsid:plazi:treatment:251287B10E67FFBCFDD2FC1E8F793554:A864607C0E63FFBCFEBBFEF08ECD3554" box="[352,673,358,382]" municipality="Western Ghats" name="Thiruvananthapuram district" pageId="20" pageNumber="1314" stateProvince="Kerala">Thiruvananthapuram district</location>
of
<collectingRegion id="6F7FF8450E63FFBCFD12FEF08F7C3554" box="[713,784,358,382]" country="India" name="Kerala" pageId="20" pageNumber="1314">Kerala</collectingRegion>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>