174 lines
17 KiB
XML
174 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="24E33431B1D63C22C7F9D48D286FCE4C" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Ornithocheirus brachyrhinus Seeley 1870, (nomen dubium)" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="42" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="42" updateTime="1634706563434" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Rodrigues, Taissa</mods:namePart>
|
|
<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
|
|
<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
|
|
<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2013</mods:date>
|
|
<mods:detail type="pubDate">
|
|
<mods:number>2013-06-12</mods:number>
|
|
</mods:detail>
|
|
<mods:detail type="volume">
|
|
<mods:number>308</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>1</mods:start>
|
|
<mods:end>112</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
|
|
<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
|
|
<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
|
|
<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-GBIF-Taxon="152045002" LSID="urn:lsid:plazi:treatment:24E33431B1D63C22C7F9D48D286FCE4C" httpUri="http://treatment.plazi.org/id/24E33431B1D63C22C7F9D48D286FCE4C" lastPageNumber="42" pageId="41" pageNumber="42">
|
|
<subSubSection pageId="41" pageNumber="42" type="nomenclature">
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<taxonomicName LSID="24E33431-B1D6-3C22-C7F9-D48D286FCE4C" authority="Seeley, 1870" authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ornithocheirus brachyrhinus" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus" status="(nomen dubium)">Ornithocheirus brachyrhinus Seeley, 1870</taxonomicName>
|
|
<taxonomicNameLabel pageId="41" pageNumber="42">(nomen dubium)</taxonomicNameLabel>
|
|
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A-D Ornithocheirus brachyrhinus, holotype CAMSM B 54443 (Albian, Cambridge Greensand), anterior part of the rostrum. A right lateral view B respective line drawing C ventral view D respective line drawing. E-H Ornithocheirus carteri, holotype CAMSM B 54437 (Albian, Cambridge Greensand), anterior part of the rostrum E anterior view F respective line drawing G left lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22084" pageId="41" pageNumber="42">Fig. 16A-D</figureCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="41" pageNumber="42" type="reference_group">
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Ornithocheiridae" genus="Ptenodactylus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus" status="(nomen dubium)">
|
|
<normalizedToken originalValue="“Ptenodactylus”">"Ptenodactylus"</normalizedToken>
|
|
brachyrhinus
|
|
</taxonomicName>
|
|
Seeley:
|
|
<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B77" refString="Seeley, HG, 1869. Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." title="Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." year="1869">Seeley 1869</bibRefCitation>
|
|
: p. xvi [disclaimed]
|
|
</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus" status="(nomen dubium)">Ornithocheirus brachyrhinus</taxonomicName>
|
|
Seeley:
|
|
<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
|
|
: p. 123
|
|
</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus" status="(nomen dubium)">Ornithocheirus brachyrhinus</taxonomicName>
|
|
Seeley:
|
|
<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
|
|
: p. 535
|
|
</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Anhanguera" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="cuvieri" status="(nomen dubium)">Anhanguera cuvieri</taxonomicName>
|
|
(Bowerbank):
|
|
<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
|
|
: table 1
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="41" pageNumber="42" type="holotype">
|
|
<paragraph pageId="41" pageNumber="42">Holotype.</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">
|
|
CAMSM B54443, anterior portion of the rostrum (
|
|
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A-D Ornithocheirus brachyrhinus, holotype CAMSM B 54443 (Albian, Cambridge Greensand), anterior part of the rostrum. A right lateral view B respective line drawing C ventral view D respective line drawing. E-H Ornithocheirus carteri, holotype CAMSM B 54437 (Albian, Cambridge Greensand), anterior part of the rostrum E anterior view F respective line drawing G left lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22084" pageId="41" pageNumber="42">Fig. 16A-D</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="41" pageNumber="42" type="type locality">
|
|
<paragraph pageId="41" pageNumber="42">Type locality.</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">Cambridge, Cambridgeshire, England.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="41" pageNumber="42" type="type horizon">
|
|
<paragraph pageId="41" pageNumber="42">Type horizon.</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="41" pageNumber="42" type="description">
|
|
<paragraph pageId="41" pageNumber="42">Description.</paragraph>
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Ornithocheirus brachyrhinus</emphasis>
|
|
</taxonomicName>
|
|
is known from the tip of a snout, with a dorsally curved palate and lacking an anteriorly located crest. It shares with
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="cuvieri">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Cimoliopterus cuvieri</emphasis>
|
|
</taxonomicName>
|
|
features such as the curved palate, the anterior end being higher than wide, lack of an anterior expansion, and absence of an anterior crest. The structure of
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Ornithocheirus brachyrhinus</emphasis>
|
|
</taxonomicName>
|
|
corresponds perfectly to the tip of the snout of
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="cuvieri">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Cimoliopterus cuvieri</emphasis>
|
|
</taxonomicName>
|
|
and it is possibly referable to that species, as
|
|
<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin (2001)</bibRefCitation>
|
|
has suggested. However, CAMSM B54443 is fragmentary and therefore it is not possible to establish if it had a posteriorly located crest and the alveoli size variation diagnostic of
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" baseAuthorityName="Rodrigues & Kellner" baseAuthorityYear="2013" class="Reptilia" genus="Cimoliopterus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="cuvieri">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Cimoliopterus cuvieri</emphasis>
|
|
</taxonomicName>
|
|
. Thus,
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Ornithocheirus brachyrhinus</emphasis>
|
|
</taxonomicName>
|
|
is here considered a
|
|
<emphasis italics="true" pageId="41" pageNumber="42">nomen dubium</emphasis>
|
|
.
|
|
</paragraph>
|
|
<caption httpUri="https://binary.pensoft.net/fig/22084" pageId="41" pageNumber="42" start="Figure 16" startId="F16">
|
|
<paragraph pageId="41" pageNumber="42">
|
|
<emphasis bold="true" pageId="41" pageNumber="42">Figure 16.</emphasis>
|
|
<emphasis bold="true" pageId="41" pageNumber="42">A-D</emphasis>
|
|
<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="brachyrhinus">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Ornithocheirus brachyrhinus</emphasis>
|
|
</taxonomicName>
|
|
, holotype CAMSM B54443 (Albian, Cambridge Greensand), anterior part of the rostrum.
|
|
<emphasis bold="true" pageId="41" pageNumber="42">A</emphasis>
|
|
right lateral view
|
|
<emphasis bold="true" pageId="41" pageNumber="42">B</emphasis>
|
|
respective line drawing
|
|
<emphasis bold="true" pageId="41" pageNumber="42">C</emphasis>
|
|
ventral view
|
|
<emphasis bold="true" pageId="41" pageNumber="42">D</emphasis>
|
|
respective line drawing.
|
|
<emphasis bold="true" pageId="41" pageNumber="42">E-H</emphasis>
|
|
<taxonomicName authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="carteri">
|
|
<emphasis italics="true" pageId="41" pageNumber="42">Ornithocheirus carteri</emphasis>
|
|
</taxonomicName>
|
|
, holotype CAMSM B 54437 (Albian, Cambridge Greensand), anterior part of the rostrum
|
|
<emphasis bold="true" pageId="41" pageNumber="42">E</emphasis>
|
|
anterior view
|
|
<emphasis bold="true" pageId="41" pageNumber="42">F</emphasis>
|
|
respective line drawing
|
|
<emphasis bold="true" pageId="41" pageNumber="42">G</emphasis>
|
|
left lateral view
|
|
<emphasis bold="true" pageId="41" pageNumber="42">H</emphasis>
|
|
respective line drawing. Abbreviations:
|
|
<emphasis bold="true" pageId="41" pageNumber="42">m</emphasis>
|
|
- maxillae,
|
|
<emphasis bold="true" pageId="41" pageNumber="42">pm</emphasis>
|
|
- premaxillae,
|
|
<emphasis bold="true" pageId="41" pageNumber="42">prid</emphasis>
|
|
- palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm.
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |