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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.238.115494" ID-Pensoft-Pub="1314-2003-238-85" ID-Pensoft-UUID="C9B5F40CB5E352F5918D1DA395640A4C" ModsDocID="1314-2003-238-85" checkinTime="1707310973662" checkinUser="pensoft" docAuthor="Su, Fan, Qin, Xiao-Wei, Fan, Rui, Yan, Lin, Ji, Xun-Zhi &amp; Hao, Chao-Yun" docDate="2024" docId="24A792C8C6895716AE74229B484DF470" docLanguage="en" docName="PhytoKeys 238: 85-94" docOrigin="PhytoKeys 238" docPubDate="2024-02-07" docSource="http://dx.doi.org/10.3897/phytokeys.238.115494" docTitle="Piper motuoense X. W. Qin, F. Su &amp; C. Y. Hao 2024, sp. nov." docType="treatment" docVersion="2" id="C9B5F40CB5E352F5918D1DA395640A4C" lastPageNumber="85" masterDocId="C9B5F40CB5E352F5918D1DA395640A4C" masterDocTitle="Piper motuoense, a new species of Piperaceae from Xizang, China" masterLastPageNumber="94" masterPageNumber="85" pageNumber="85" updateTime="1707311274392" updateUser="ExternalLinkService">
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<mods:title id="7D9B7101863CF40B3DCE8858D1702A76">Piper motuoense, a new species of Piperaceae from Xizang, China</mods:title>
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<mods:namePart id="CD0E37B519101B6ED25CF477F886A5E5">Su, Fan</mods:namePart>
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<mods:affiliation id="8AB44DAC90B44C16EA666A88EC04F49E">Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China &amp; Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China &amp; Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China</mods:affiliation>
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<mods:namePart id="7FA7BB7FF872EDF70E71FFBF211F6167">Qin, Xiao-Wei</mods:namePart>
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<mods:affiliation id="E9A3FC790B6CAF2E83BCF551E827CA7B">Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China &amp; Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China &amp; Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China</mods:affiliation>
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<mods:namePart id="5D46C737147675E986141DFED2A40756">Fan, Rui</mods:namePart>
<mods:affiliation id="FFF710C6BD70D69EBDBE5328E272D828">Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China &amp; Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China &amp; Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China</mods:affiliation>
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<mods:namePart id="374695F0CB487C8E80EECF17AC3CDB83">Yan, Lin</mods:namePart>
<mods:affiliation id="27C3E73F78E1C4BB3B28C92BEE1E08CE">Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China &amp; Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China &amp; Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China</mods:affiliation>
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<mods:namePart id="661E2076B78406AC66ED24EF9FA34817">Ji, Xun-Zhi</mods:namePart>
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<mods:affiliation id="430B163D376F5980A98C146D6E78F1EF">Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China &amp; Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China &amp; Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China</mods:affiliation>
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<mods:namePart id="9FC4161F22383C8538906949BB513577">Hao, Chao-Yun</mods:namePart>
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<mods:affiliation id="221303FC53CB3B001825548AF186770B">Spice and Beverage Research Institute, CATAS, Wanning 571533, Hainan, China &amp; Ministry of Agriculture Key Laboratory of Genetic Resources Utilization of Spice and Beverage Crops, Wanning 571533, Hainan, China &amp; Hainan Provincial Key Laboratory of Genetic Improvement and Quality Regulation for Tropical Spice and Beverage Crops, Wanning 571533, Hainan, China</mods:affiliation>
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<mods:date id="FD697CF8AAA9DEC02EF3EA2CC8654D84">2024</mods:date>
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<mods:start id="0B96C38D231033456C05380AFB2655EE">85</mods:start>
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<mods:identifier id="95D9350F2396D622D633703E60B63314" type="DOI">http://dx.doi.org/10.3897/phytokeys.238.115494</mods:identifier>
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<treatment id="24A792C8C6895716AE74229B484DF470" LSID="urn:lsid:plazi:treatment:24A792C8C6895716AE74229B484DF470" httpUri="http://treatment.plazi.org/id/24A792C8C6895716AE74229B484DF470" lastPageNumber="85" pageId="0" pageNumber="85">
<subSubSection id="22D74CFCA0703EEA2399D7CE0D235178" pageId="0" pageNumber="85" type="nomenclature">
<paragraph id="DB28BF838DE7086A903E9A9B21F429C4" pageId="0" pageNumber="85">
<taxonomicName id="7A5D860A2925231DD6B58872125E8903" LSID="24A792C8-C689-5716-AE74-229B484DF470" authority="X. W. Qin, F. Su &amp; C. Y. Hao" authorityName="X. W. Qin, F. Su &amp; C. Y. Hao" authorityYear="2024" class="Magnoliopsida" family="Piperaceae" genus="Piper" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piper motuoense" order="Piperales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="species" species="motuoense" status="sp. nov.">Piper motuoense X.W.Qin, F.Su &amp; C.Y.Hao</taxonomicName>
<taxonomicNameLabel id="3C15073FBC5F9E6470FB7E0F5C33E2A6" pageId="0" pageNumber="85">sp. nov.</taxonomicNameLabel>
</paragraph>
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<paragraph id="0980CAB1819208BA121E6DF2A930959C" pageId="0" pageNumber="85">
<figureCitation id="C4E15074DEC726A9DDBC98DBD5312CF8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Piper motuoense X. W. Qin, F. Su &amp; C. Y. Hao, sp. nov. A habit B branch with infructescence C branch with staminate spike D branch with trophophyll leaf E adaxial and abaxial surface of gonophyll leaf F adaxial and abaxial surface of gonophyll leaf base G detail of the indument along the mid-vein of the gonophyll leaf adaxial and abaxial surface H pistillate spike I staminate spike J infructescence K close-up of portion of the staminate spike L close-up of portion of the infructescence M cross-section of infructescence N seed (side view). Photographs by Fan Su." figureDoi="10.3897/phytokeys.238.115494.figure1" httpUri="https://binary.pensoft.net/fig/980526" pageId="0" pageNumber="85">Figs 1</figureCitation>
<figureCitation id="233400FA3C412D45150FD318287BF495" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Line drawing of Piper motuoense X. W. Qin, F. Su &amp; C. Y. Hao A branch with infructescence B branch with staminate spike C magnified view of pistillate spike D magnified view of staminate spike E magnified view of infructescence F detail of the indument along the secondary nerves of the abaxial leaf surface G adaxial surface of gonophyll leaf H adaxial surface of gonophyll leaf base I abaxial surface of gonophyll leaf J abaxial surface of gonophyll leaf base K cross-section of infructescence L stamen M carpel N seed. Illustration by Fan Su, based on the holotype." figureDoi="10.3897/phytokeys.238.115494.figure2" httpUri="https://binary.pensoft.net/fig/980527" pageId="0" pageNumber="85">, 2</figureCitation>
</paragraph>
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<subSubSection id="28C79841A6DA60F5E8922CD55BCBF440" pageId="0" pageNumber="85" type="diagnosis">
<paragraph id="D4B3E99FF89D0E8B2D02F8F3F855E685" pageId="0" pageNumber="85">Diagnosis.</paragraph>
<paragraph id="74FC012075FBDDCEF0A94FD151C765D5" pageId="0" pageNumber="85">
The new species is morphologically similar to
<taxonomicName id="1927A7E5D600DD939878E1A48AA8896B" lsidName="P. yinkiangense" pageId="0" pageNumber="85" rank="species" species="yinkiangense">
<emphasis id="5A8E5EA8A0B3CAA3A4F4ED10AE7DD325" italics="true" pageId="0" pageNumber="85">P. yinkiangense</emphasis>
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, but can be easily distinguished from the latter in several aspects. The leaf-blades 12.5-18
<normalizedToken id="F9DF745AFEEBEF57DF2393BF1D5EA8E1" originalValue="×">x</normalizedToken>
3.5-6.5 cm, elliptic or ovate to lanceolate (vs. 11-14
<normalizedToken id="BDC8F5D2F6A756E2C275D3A1B510E612" originalValue="×">x</normalizedToken>
6.5-8.5 cm, oblique-ovate), chartaceous (vs. membranous), abaxially sparsely villous along the mid-vein (vs. abaxially sparsely hispidulous), gonophyll leaves 7-9 secondary vein pairs, the outer pair arising 2-4 cm above base (vs. 8-9, outer pair arising 1-2 cm above base), base bilaterally clinging to the petiole and overlap together (vs. basal sinus 1-2 mm wide on side of longer and wider lobe, 4-5 mm wide on other side, bilaterally free for 2-3 mm), floral bracts sessile (vs. petiolate) and stigmas 3, 4 or 5 (vs. 4).
<taxonomicName id="1844F77588D5FFAA7A58630B4E75E38B" lsidName="P. motuoense" pageId="0" pageNumber="85" rank="species" species="motuoense">
<emphasis id="1E1CF81E5A140C0419EC879D92C2E0F2" italics="true" pageId="0" pageNumber="85">P. motuoense</emphasis>
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also resembles
<taxonomicName id="81E633DDED2427B19E69834355610FBA" lsidName="P. anisotis" pageId="0" pageNumber="85" rank="species" species="anisotis">
<emphasis id="BABE76FAFE3F0238805B5FB06CEEC976" italics="true" pageId="0" pageNumber="85">P. anisotis</emphasis>
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in the shape of leaves and fruit, but differs from the latter in the leaves vein 7-9 (vs. 5-7), leave base bilateral clinging to petiole and overlap together (vs. bilateral free for 1-2 mm) (Table
<tableCitation id="6557E3C6FDF970154D97EA62B623E8CD" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of key characteristics in P. motuoense, P. yinkiangense and P. anisotis." httpUri="http://table.plazi.org/id/C442A2EE40412627CD580B30ABB6BEB5" pageId="0" pageNumber="85" tableUuid="C442A2EE40412627CD580B30ABB6BEB5">1</tableCitation>
).
</paragraph>
<caption id="C442A2EE40412627CD580B30ABB6BEB5" ID-Table-UUID="C442A2EE40412627CD580B30ABB6BEB5" httpUri="http://table.plazi.org/id/C442A2EE40412627CD580B30ABB6BEB5" pageId="0" pageNumber="85" start="Table 1" startId="T1">
<paragraph id="98126A0785CC6818570F4024CF8BC84F" pageId="0" pageNumber="85">
<emphasis id="3ABE4492A6FF732D296D2BE5A80EE91A" bold="true" pageId="0" pageNumber="85">Table 1.</emphasis>
Morphological comparison of key characteristics in
<taxonomicName id="018EB02F22147F31DD8568224976E36B" lsidName="P. motuoense" pageId="0" pageNumber="85" rank="species" species="motuoense">
<emphasis id="B31326148C025920DA467CB4CF34DB04" italics="true" pageId="0" pageNumber="85">P. motuoense</emphasis>
</taxonomicName>
,
<taxonomicName id="B1618442E5FD0E1AEF764445A77ECE83" lsidName="P. yinkiangense" pageId="0" pageNumber="85" rank="species" species="yinkiangense">
<emphasis id="5AA6999675272D39B5C3D84307D5785F" italics="true" pageId="0" pageNumber="85">P. yinkiangense</emphasis>
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and
<taxonomicName id="B7555F38E3958CA5E5A2C723C24496E6" lsidName="P. anisotis" pageId="0" pageNumber="85" rank="species" species="anisotis">
<emphasis id="BE90383B69531140527D9D0526CDBEA5" italics="true" pageId="0" pageNumber="85">P. anisotis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="C05CD94635A7C4C48B582002F32A2768" pageId="0" pageNumber="85">
<table id="7979FD6AC5487A44C384C0DE2426FF4F" pageId="0" pageNumber="85">
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<th id="5CE505686F82A4AC8215270F3E70EB53" colspan="2" pageId="0" pageNumber="85" rowspan="1">Characters</th>
<th id="D6A4DBF109314BF5BFED981667AC2264" colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName id="B1647ABD68236EC69D7DEDD37FE319EE" lsidName="P. motuoense" pageId="0" pageNumber="85" rank="species" species="motuoense">
<emphasis id="02C873FEA35B369550B1A4F176EEB483" italics="true" pageId="0" pageNumber="85">P. motuoense</emphasis>
</taxonomicName>
</th>
<th id="354BEB66E4F223601813628782FF5372" colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName id="0AAFA48584174AEE37B8B60390533173" lsidName="P. yinkiangense" pageId="0" pageNumber="85" rank="species" species="yinkiangense">
<emphasis id="F22A1EC310C17307AFFD67195AA262F3" italics="true" pageId="0" pageNumber="85">P. yinkiangense</emphasis>
</taxonomicName>
</th>
<th id="01E4F173F62D504708B9C07730306A81" colspan="1" pageId="0" pageNumber="85" rowspan="1">
<taxonomicName id="A9103BEE304A3E90041EDB3ED362DEB9" lsidName="P. anisotis" pageId="0" pageNumber="85" rank="species" species="anisotis">
<emphasis id="1A843AF8A4888BC24BCD1D1BD8133405" italics="true" pageId="0" pageNumber="85">P. anisotis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="79BD5EAA2B13BE42F3E38119BC67940E" pageId="0" pageNumber="85">
<td id="4617C8D448134023C8094F320BDDE6CC" colspan="2" pageId="0" pageNumber="85" rowspan="1">Stem</td>
<td id="70494446CB7B00F59FEC2B13811B9DFE" colspan="1" pageId="0" pageNumber="85" rowspan="1">Densely villous when young, glabrous when mature</td>
<td id="5EA7AB96BACFAB565A5815789E9A8600" colspan="1" pageId="0" pageNumber="85" rowspan="1">Densely villous</td>
<td id="509FDBFE0F7A70FEE6B52AB47F199DFD" colspan="1" pageId="0" pageNumber="85" rowspan="1">Densely short tomentose</td>
</tr>
<tr id="59CCF3F0622106C483001BAA0CD81CD3" pageId="0" pageNumber="85">
<td id="EB584B9C82BE3CEDC9338E6AD2D33CF4" colspan="1" pageId="0" pageNumber="85" rowspan="4">
<taxonomicName id="C68398034B0E66E311060688BCBCCAFB" genus="Gonophyll" lsidName="Gonophyll leaves" pageId="0" pageNumber="85" rank="species" species="leaves">Gonophyll leaves</taxonomicName>
</td>
<td id="7019B8C5EEA32C77B88884E625219D59" colspan="1" pageId="0" pageNumber="85" rowspan="1">petiole</td>
<td id="8400D3C2DCCDCEC666C8107C491BB1A7" colspan="1" pageId="0" pageNumber="85" rowspan="1">2.5-3 cm long</td>
<td id="6AF46B6DE161A6B2C7949126872772B5" colspan="1" pageId="0" pageNumber="85" rowspan="1">Ca. 2 mm long</td>
<td id="CC4DA6AA13D24FC66AB3B8577547A6FC" colspan="1" pageId="0" pageNumber="85" rowspan="1">Ca. 3 mm long</td>
</tr>
<tr id="2860EEA00055B3A445EB1BD2CCF71B04" pageId="0" pageNumber="85">
<td id="D52C61BF6395B536472D6A9E97EDA45A" colspan="1" pageId="0" pageNumber="85" rowspan="1">blade</td>
<td id="7E3BB6EE8619E213A458E01F65118FA2" colspan="1" pageId="0" pageNumber="85" rowspan="1">
12.5-18
<normalizedToken id="7726B9F888441AAEE044A46EF747960F" originalValue="×">x</normalizedToken>
3.5-6.5 cm, elliptic or ovate to lanceolate, chartaceous, abaxially densely villous, especially along the veins, adaxially sparsely villous along the mid-vein
</td>
<td id="48F606146F7C35B19BD661FCECE5560D" colspan="1" pageId="0" pageNumber="85" rowspan="1">
11-14
<normalizedToken id="3B1237F389A147450C6C70ADAC710BFE" originalValue="×">x</normalizedToken>
6.5-8.5 cm, oblique-ovate, membranous, abaxially densely pubescent, usually along veins, adaxial sparsely hispidulous
</td>
<td id="E4DEB43A59BA055DA3D6D2D080C6F5BB" colspan="1" pageId="0" pageNumber="85" rowspan="1">
7.5-13
<normalizedToken id="EE3AD8B43D88F25747EAE2FE85AFD16C" originalValue="×">x</normalizedToken>
2.5-5 cm, oblique-oblong, membranaceous, abaxially densely villous, especially along the veins, adaxially densely hispidulous
</td>
</tr>
<tr id="7D8C72FEC5395F1A57E0A8E5DCBE6FD0" pageId="0" pageNumber="85">
<td id="BA5ADC134E4280D4CF0AC56E77DBBE51" colspan="1" pageId="0" pageNumber="85" rowspan="1">base</td>
<td id="B60AB2FEED1D90946715DDF83B3B0B07" colspan="1" pageId="0" pageNumber="85" rowspan="1">Auriculate-cordate, bilaterally clinging to leaves petiole and overlap together</td>
<td id="1EE740649A0769947B1D87169C330FF6" colspan="1" pageId="0" pageNumber="85" rowspan="1">Obliquely auriculate-cordate, basal sinus 1-2 mm wide on side of longer and wider lobe, 4-5 mm wide on other side, bilateral difference to 2-3 mm</td>
<td id="7E228B7866A52571EEC5147472625136" colspan="1" pageId="0" pageNumber="85" rowspan="1">Unequal-sided and more cordate, basal sinus 1-2 mm wide on side of longer and wider lobe, 3-4 mm wide on other side, bilateral difference to 1-2 mm</td>
</tr>
<tr id="FB6D6F79FEBE03C9F0F40FD886774F34" pageId="0" pageNumber="85">
<td id="D019AC3F2A507612DB91FDC1C308E7A3" colspan="1" pageId="0" pageNumber="85" rowspan="1">secondaryveins</td>
<td id="03331A515B51606B9FD683A924C8DAB0" colspan="1" pageId="0" pageNumber="85" rowspan="1">7-9, when 9 veins, the outermost pair is very weak, apical pair arising 2-4 cm above base</td>
<td id="F66A1BF789B513BDC122C91103E5D133" colspan="1" pageId="0" pageNumber="85" rowspan="1">8-9, apical pair arising 1-2 cm above base</td>
<td id="C1B73E4DA07A206C7A6D19ED14734FFE" colspan="1" pageId="0" pageNumber="85" rowspan="1">5-7, apical pair arising 1.3-2.5 cm above base</td>
</tr>
<tr id="5A0EBE4886E3E4664CBB660D8823BB78" pageId="0" pageNumber="85">
<td id="05A14F2C6120A25DDF3336930FA5EE2A" colspan="1" pageId="0" pageNumber="85" rowspan="4">Pistillate spikes</td>
<td id="AE6210EBFC01036CCBBB17A0167EEBF2" colspan="1" pageId="0" pageNumber="85" rowspan="1">spikes</td>
<td id="39E21E61777D1F84AD3D41C2782DEAF3" colspan="1" pageId="0" pageNumber="85" rowspan="1">
4-5
<normalizedToken id="A1173866737FF9076F75E1DF2E6D27B5" originalValue="×">x</normalizedToken>
0.5-0.7 cm
</td>
<td id="2BB67C802C2F8C4D86F683CC19E14286" colspan="1" pageId="0" pageNumber="85" rowspan="1">
3
<normalizedToken id="708F137835B91D34BEC795520D8144B2" originalValue="×">x</normalizedToken>
0.4 cm
</td>
<td id="7DF57CB778017D69666C0B214833CFBB" colspan="1" pageId="0" pageNumber="85" rowspan="1">
1.5-3
<normalizedToken id="B60A1B235CFF923961A955DF78CAB7CB" originalValue="×">x</normalizedToken>
0.4-0.6 cm
</td>
</tr>
<tr id="C984D64796B4B445F3F9AD9D7484062A" pageId="0" pageNumber="85">
<td id="8E40D34BEAFED8D48001C5AAF205332E" colspan="1" pageId="0" pageNumber="85" rowspan="1">peduncles</td>
<td id="A93EEF9D2565D7F0FD3686249E9370C1" colspan="1" pageId="0" pageNumber="85" rowspan="1">2-3 cm long, slightly shorter than spike</td>
<td id="CDA030DF86170F3F429AE91AF7D66497" colspan="1" pageId="0" pageNumber="85" rowspan="1">2.5 cm long, equal to or longer than spike</td>
<td id="22CF505B0F5B61CEE2090195F95A2F2D" colspan="1" pageId="0" pageNumber="85" rowspan="1">2 cm long, equal to or longer than spike</td>
</tr>
<tr id="01108879DE71CBC0DA8A85EF4490DDED" pageId="0" pageNumber="85">
<td id="23B578FF08E3AECC6FA9DD697FEF4A98" colspan="1" pageId="0" pageNumber="85" rowspan="1">floral bracts</td>
<td id="8E7D1E5C4CADDC64799DD8FC8E9F9197" colspan="1" pageId="0" pageNumber="85" rowspan="1">suborbicular, sessile</td>
<td id="D1C649AE99BA55B793FD84A23E662FED" colspan="1" pageId="0" pageNumber="85" rowspan="1">suborbicular, short-pedicellate</td>
<td id="4FBA6050FB9DD4FBAB3F2B904DC768DB" colspan="1" pageId="0" pageNumber="85" rowspan="1">suborbicular, short-pedicellate</td>
</tr>
<tr id="A2926B09B28576A18B52BF193885CA83" pageId="0" pageNumber="85">
<td id="4A95A6455AC944603BFF4D340CFC8809" colspan="1" pageId="0" pageNumber="85" rowspan="1">stigmas</td>
<td id="5872D5D7DA98F74529F218E4A900E707" colspan="1" pageId="0" pageNumber="85" rowspan="1">3-4-5, 0.8-1 mm long</td>
<td id="DF1898426DAC012D558333F6213C2D58" colspan="1" pageId="0" pageNumber="85" rowspan="1">4, ca. 1 mm or longer</td>
<td id="9632D67E8EF6D15502AC2ED2C233538A" colspan="1" pageId="0" pageNumber="85" rowspan="1">4, ca. 1 mm or longer</td>
</tr>
<tr id="3CE236AFB0BD52E9AFD290550E6E3F75" pageId="0" pageNumber="85">
<td id="FA652958AB9AD7BE6B2E4CF8596771D1" colspan="2" pageId="0" pageNumber="85" rowspan="1">Fruit</td>
<td id="30B4C1EC8D3CFCA2F91161B22349E0C7" colspan="1" pageId="0" pageNumber="85" rowspan="1">
3-3.5
<normalizedToken id="F02FCE6DCAAAECBB8E2C19A2CA733BA1" originalValue="×">x</normalizedToken>
2.5-3 mm
</td>
<td id="F35B0CD5F80497E49F520C53259530D0" colspan="1" pageId="0" pageNumber="85" rowspan="1">Ca. 3 mm in diam.</td>
<td id="9B0FFE4594BC6E09D495D85B15389C68" colspan="1" pageId="0" pageNumber="85" rowspan="1">Ca. 3 mm in diam.</td>
</tr>
</table>
</paragraph>
<caption id="210A912EFD769FEF12641232A54601D2" doi="10.3897/phytokeys.238.115494.figure1" httpUri="https://binary.pensoft.net/fig/980526" pageId="0" pageNumber="85" start="Figure 1" startId="F1">
<paragraph id="BCBA6676300C3BCE8445DC797AB1A25E" pageId="0" pageNumber="85">
<emphasis id="37BF579921F19247CC99A66DAA76A21C" bold="true" pageId="0" pageNumber="85">Figure 1.</emphasis>
<taxonomicName id="8F1BDF18BAB8A1F57E37F57373BB428E" authorityName="X. W. Qin, F. Su &amp; C. Y. Hao" authorityYear="2024" class="Magnoliopsida" family="Piperaceae" genus="Piper" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piper motuoense" order="Piperales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="species" species="motuoense">
<emphasis id="E83B74E11865614A00147EDB9711C43E" italics="true" pageId="0" pageNumber="85">Piper motuoense</emphasis>
</taxonomicName>
X.W.Qin, F.Su &amp; C.Y.Hao, sp. nov.
<emphasis id="D0BB392ACF5E45E4744EDE434AD81DEE" bold="true" pageId="0" pageNumber="85">A</emphasis>
habit
<emphasis id="1DBD4B9A04B9C5EFEE98C2E1791FA411" bold="true" pageId="0" pageNumber="85">B</emphasis>
branch with infructescence
<emphasis id="0B2782E3969434DC993BBD053F83809E" bold="true" pageId="0" pageNumber="85">C</emphasis>
branch with staminate spike
<emphasis id="93A5E805549E67DE3DE38C0FFD812D5F" bold="true" pageId="0" pageNumber="85">D</emphasis>
branch with trophophyll leaf
<emphasis id="0EDCC32636DFC7DE8E50541137447FD2" bold="true" pageId="0" pageNumber="85">E</emphasis>
adaxial and abaxial surface of gonophyll leaf
<emphasis id="CC6876AC3548A11F60ACBC240026FD78" bold="true" pageId="0" pageNumber="85">F</emphasis>
adaxial and abaxial surface of gonophyll leaf base
<emphasis id="B2F71D4DAE0E3D1950D60223271D97C4" bold="true" pageId="0" pageNumber="85">G</emphasis>
detail of the indument along the mid-vein of the gonophyll leaf adaxial and abaxial surface
<emphasis id="39B6F14450074B32FEF911791937B4F7" bold="true" pageId="0" pageNumber="85">H</emphasis>
pistillate spike
<emphasis id="D8A56BDF4E4748410E1FE81102960A71" bold="true" pageId="0" pageNumber="85">I</emphasis>
staminate spike
<emphasis id="5A76DE2A3A94E7F9F78E398292B5143C" bold="true" pageId="0" pageNumber="85">J</emphasis>
infructescence
<emphasis id="511AFC744480FEBDC2274E13ECC1E4E7" bold="true" pageId="0" pageNumber="85">K</emphasis>
close-up of portion of the staminate spike
<emphasis id="4C2477D0C3470D70AF1A9039B9659FF0" bold="true" pageId="0" pageNumber="85">L</emphasis>
close-up of portion of the infructescence
<emphasis id="569849F2E0AE5C3CF7941892034F3049" bold="true" pageId="0" pageNumber="85">M</emphasis>
cross-section of infructescence
<emphasis id="AFD4C8F9F38F846BC94DCD9A8443B3C5" bold="true" pageId="0" pageNumber="85">N</emphasis>
seed (side view). Photographs by Fan Su.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="A9A9F20D98C664CBCA18598F4C9FA37D" pageId="0" pageNumber="85" type="materials_examined">
<paragraph id="13D89AD3E1B0BF3B8CCE4E393B77F860" pageId="0" pageNumber="85">
<typeStatus id="019D50150C34263FDA5712793FBA1C0F">Type</typeStatus>
.
</paragraph>
<paragraph id="A3766330FF7629528C404BA291DA6A9A" pageId="0" pageNumber="85">
<materialsCitation id="F8768D6DB8870BE16488896F921C436A" collectingDate="2021-10-03" country="China" county="Linzhi" elevation="490" latitude="29.18" location="Beibeng" longLatPrecision="20" longitude="95.00166" municipality="Motuo City" specimenCount="1" stateProvince="Xizang" typeStatus="Holotype">
<collectingCountry id="4781EE2CFD074D56D407AA20765DC43F" name="China">China</collectingCountry>
. (
<collectingRegion id="249695F43736965497827FD958FCDE7F" country="China" name="Xizang">Xizang</collectingRegion>
):
<collectingCounty id="7F8A23F65CEFC6071987692D5B5D5B0A">Linzhi</collectingCounty>
,
<collectingMunicipality id="EF8563180ECD3D147B687ACC92BA2952">Motuo City</collectingMunicipality>
,
<location id="285572749081F1DA53A187767AE2D85A" LSID="urn:lsid:plazi:treatment:24A792C8C6895716AE74229B484DF470:285572749081F1DA53A187767AE2D85A" country="China" county="Linzhi" latitude="29.18" longLatPrecision="20" longitude="95.00166" municipality="Motuo City" name="Beibeng" stateProvince="Xizang">Beibeng</location>
, climbing on the taller trees in tropical rainforest,
<geoCoordinate id="616EBB6CA0CB4AF63412CA2C99A17133" degrees="29" direction="north" minutes="10" orientation="latitude" precision="15" seconds="48" value="29.18">29°1048″N</geoCoordinate>
,
<geoCoordinate id="DC0BEEF8D688C574F17B5F57F8D3DBC9" degrees="95" direction="east" minutes="00" orientation="longitude" precision="15" seconds="06" value="95.00166">95°0006″E</geoCoordinate>
, elevation ca.
<quantity id="570BEB811EF9CE2C62F360DCB78B7302" metricMagnitude="2" metricUnit="m" metricValue="4.9" unit="m" value="490.0">
<elevation id="CE818C0626644EE2C648BAD7095F7A83" metricMagnitude="2" metricUnit="m" metricValue="4.9" unit="m" value="490.0">490 m</elevation>
</quantity>
,
<collectingDate id="1ADA0CBFFF97AEDC7FB320C6FDC9D98C" value="2021-10-03">3 Oct. 2021</collectingDate>
,
<emphasis id="CE3EFFAB27AC338FB19C438A2E0DD901" italics="true" pageId="0" pageNumber="85">Xiao-Wei Qin et al. 20211003, 20231016</emphasis>
(
<typeStatus id="687A8010D8900D95FB4176B1BD9A377F" type="holotype">Holotype</typeStatus>
: IBSC0918558;
<typeStatus id="D3B317CA20E64695C6C6410178A07E3F" type="isotype">Isotype</typeStatus>
: IBSC0918559, IBSC0918560, IBSC0918561)
</materialsCitation>
.
</paragraph>
<caption id="D4FFB3382DDD1E80DE51654B1B461B30" doi="10.3897/phytokeys.238.115494.figure2" httpUri="https://binary.pensoft.net/fig/980527" pageId="0" pageNumber="85" start="Figure 2" startId="F2">
<paragraph id="9AB116A22DF630AFA5275B5993B4993F" pageId="0" pageNumber="85">
<emphasis id="082BE75162203AB56644242126C23DC9" bold="true" pageId="0" pageNumber="85">Figure 2.</emphasis>
Line drawing of
<taxonomicName id="0F0A6F1A5E80D44FAD63C764E4B315EF" authorityName="X. W. Qin, F. Su &amp; C. Y. Hao" authorityYear="2024" class="Magnoliopsida" family="Piperaceae" genus="Piper" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piper motuoense" order="Piperales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="species" species="motuoense">
<emphasis id="DB3137DBE00860443D0F29EEC6BD3D42" italics="true" pageId="0" pageNumber="85">Piper motuoense</emphasis>
</taxonomicName>
X.W. Qin, F. Su &amp; C.Y. Hao
<emphasis id="7DDE343DC7C485C3149DA7583EED0FA6" bold="true" pageId="0" pageNumber="85">A</emphasis>
branch with infructescence
<emphasis id="10384EFBD2B75EE9558D5D0BB7C82EC8" bold="true" pageId="0" pageNumber="85">B</emphasis>
branch with staminate spike
<emphasis id="97A0ADB0DC3E8E8AB699DAC1F14C76CE" bold="true" pageId="0" pageNumber="85">C</emphasis>
magnified view of pistillate spike
<emphasis id="C5C1C6ABEC21AFC2F165A7017DA48C37" bold="true" pageId="0" pageNumber="85">D</emphasis>
magnified view of staminate spike
<emphasis id="C3E6A09F91F6F627059FB5CD5697597A" bold="true" pageId="0" pageNumber="85">E</emphasis>
magnified view of infructescence
<emphasis id="640C9C3E195BA483942061E3567C52A0" bold="true" pageId="0" pageNumber="85">F</emphasis>
detail of the indument along the secondary nerves of the abaxial leaf surface
<emphasis id="080AFDC1D226EF48FC1EC288B567D640" bold="true" pageId="0" pageNumber="85">G</emphasis>
adaxial surface of gonophyll leaf
<emphasis id="5F7874DC7017FBEB6573AC08F53F05A6" bold="true" pageId="0" pageNumber="85">H</emphasis>
adaxial surface of gonophyll leaf base
<emphasis id="B3705B15A5ECE97C9D0176768BA61D37" bold="true" pageId="0" pageNumber="85">I</emphasis>
abaxial surface of gonophyll leaf
<emphasis id="955AC17507D432456AA9DB59C4FBAD58" bold="true" pageId="0" pageNumber="85">J</emphasis>
abaxial surface of gonophyll leaf base
<emphasis id="6C48B010F1AD413BD263D84ECFCC1360" bold="true" pageId="0" pageNumber="85">K</emphasis>
cross-section of infructescence
<emphasis id="8AAF2AEBA586C984846C89C29C823961" bold="true" pageId="0" pageNumber="85">L</emphasis>
stamen
<emphasis id="2C6FA914174549689AA15E5B4127F2D1" bold="true" pageId="0" pageNumber="85">M</emphasis>
carpel
<emphasis id="FB74AC1EE2842563A368D8B0DD884361" bold="true" pageId="0" pageNumber="85">N</emphasis>
seed. Illustration by Fan Su, based on the
<typeStatus id="590DA519F528D83E818FC6B290533B24" type="holotype">holotype</typeStatus>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="7F780F3EB7EFF343E4D55260901B7C88" pageId="0" pageNumber="85" type="description">
<paragraph id="2B9B76C027BAE06EB4D8D8AAE8566586" pageId="0" pageNumber="85">Description.</paragraph>
<paragraph id="92DC018D0EFE99D1BDB2481D677990FC" pageId="0" pageNumber="85">
<emphasis id="AAB000031619BF0707F441F90CEF9A4C" bold="true" italics="true" pageId="0" pageNumber="85">Lianas</emphasis>
over 5 m long, with climbing adventitious roots, dioecious, perennial, densely villous when young, becoming glabrous when mature.
<emphasis id="7B15D21B3555A5562D44EB6762C65A79" bold="true" italics="true" pageId="0" pageNumber="85">Roots</emphasis>
dimorphic; basal roots terrestrial, feeding; adventitious roots produced along the aerial nodes, clasping the phorophyte.
<emphasis id="CDECFF9101DF353990D9AC0F2FF36E4D" bold="true" italics="true" pageId="0" pageNumber="85">Stems</emphasis>
climbing upwards, internodes terete, striated, with red stripes when they are young, swollen nodes.
<emphasis id="C49D556CB322F91D50659545C8EDA6E5" bold="true" italics="true" pageId="0" pageNumber="85">Leaves</emphasis>
dimorphic, distichously-alternate, petiolate, blades chartaceous, glandular.
<emphasis id="E909A931531140CE0E3FB18F11C7F4CE" bold="true" italics="true" pageId="0" pageNumber="85">Trophophyll leaves</emphasis>
with petiole 2.5-3 cm long, cylinder-shaped in cross-section, red, pubescence; leaf-blades 7-9
<normalizedToken id="F3DB3E2538F87478EDA71167BB5AB7EC" originalValue="×">x</normalizedToken>
5-6 cm, cordate to elliptic, adaxially dark green, glabrous, abaxially surface pale green, sparsely villous, especially along the veins, base usually cordate, symmetric, apex short-acuminate to long-acuminate; mid-vein red, 5-7 pairs, red, all basal, when 7, the outermost pair inconspicuous.
<emphasis id="4D5ADFB0FFE41541C23EB5B109F090D0" bold="true" italics="true" pageId="0" pageNumber="85">Gonophyll leaves</emphasis>
with petiole 0.3-0.4 cm long, cylinder-shaped in cross-section, pale green, densely pubescence; leaf-blades 12.5-18
<normalizedToken id="A309534F8A490A416C6CAD0FA2465E4C" originalValue="×">x</normalizedToken>
3.5-6.5 cm, elliptic or ovate-lanceolate; 7-9 pairs of secondary veins; when 9 veins, the outermost pair is very weak; apical pair 2-4 cm above base, reaching leaf apex, alternate, others basal, reticulate veins prominent; adaxial surface dark green, glabrous or sparsely villous on the mid-vein, abaxial surface pale green, densely villous, especially on the veins; base auriculate-cordate, asymmetrical, bilateral clinging to leaves petiole and healing together, apex long acuminate.
<emphasis id="847689A8712EE84BC98850D20B7A2BEF" bold="true" italics="true" pageId="0" pageNumber="85">Inflorescence</emphasis>
a pedunculate spike, leaf-opposed, pendulous; peduncle flexible, cylindrical; spadix cylindrical, the fertile rachis hairy, with densely compacted flowers; floral bracts sessile, imbricate, ca. 1 mm in diam., orbicular or suborbicular, piligerous, margin irregular, undulate.
<emphasis id="E41E33441927E54CB63A08B0A6E65B9B" bold="true" italics="true" pageId="0" pageNumber="85">Staminate inflorescences</emphasis>
5.5-8 cm long, peduncles 3-4 cm long, villous; spike 2.5-4
<normalizedToken id="9DB3C8FE40292F8AE2452BB1BBDB9C1E" originalValue="×">x</normalizedToken>
0.5-0.7 cm, villous, pale green when young, pale yellow to white when mature;
<emphasis id="C1A9C3667B4707D49FF6CEEA446DB718" bold="true" pageId="0" pageNumber="85">stamens</emphasis>
1, filaments 0.6-1 mm long, stout, hyaline, anthers 0.4-0.7
<normalizedToken id="1EE98B924B2540316825313D6B8A1297" originalValue="×">x</normalizedToken>
0.2-0.5 mm, 2-thecous, reniform, white before dehiscence, black after dehiscence, dehiscence lateral.
<emphasis id="115946275EA13382EFDFF1B5362A78ED" bold="true" italics="true" pageId="0" pageNumber="85">Pistillate inflorescences</emphasis>
6-8 cm long, peduncles 2-3 cm long, piliferous; spike 4-5
<normalizedToken id="7EA498CDB4C69A7499AD988879E0E5B9" originalValue="×">x</normalizedToken>
0.5-0.7 cm, pale green when young, pale yellow to white when mature;
<emphasis id="F15CD65CA0D7C8112E2E33F5E004C8BF" bold="true" pageId="0" pageNumber="85">ovary</emphasis>
1.6-2
<normalizedToken id="9F13CD57BD4644377369DE11B0EE79E6" originalValue="×">x</normalizedToken>
1.4-1.8 mm, sessile, free from the neighbouring ones, obovoid, green,
<emphasis id="58AACAED5655161616489F9913C2C880" bold="true" pageId="0" pageNumber="85">style</emphasis>
1-1.2 mm long,
<emphasis id="ED4DD085EC468C4B3967EA64C58DFE95" bold="true" pageId="0" pageNumber="85">stigmas</emphasis>
3-5, 0.8-1 mm long, filiform, reflexed, cream-coloured at anthesis, becoming tan to light grey post-anthesis.
<emphasis id="266610F1D54F4CDC23404B61B9975381" bold="true" italics="true" pageId="0" pageNumber="85">Infructescence</emphasis>
leaf-opposed, 5-7
<normalizedToken id="E26E4579E6CA9801F8DB0CE7DBBE2716" originalValue="×">x</normalizedToken>
1-1.2 cm; cylindrical; with densely compacted fruits; pendulous, cylindrical, piliferous.
<emphasis id="7E8C58214C88A62FE560D24A02CE97B6" bold="true" italics="true" pageId="0" pageNumber="85">Drupes</emphasis>
3-3.5
<normalizedToken id="247E1840103DA7654FCF0C761A048008" originalValue="×">x</normalizedToken>
2.5-3 mm, sessile, free from the neighbouring ones, subglobose, remaining attached to rachis at maturity, piligerous, persistent style 0.3-0.5 mm long, cylindrical; epicarp green, mesocarp pale green, translucent, endocarp dark yellow.
<emphasis id="6DF4FA6A46A73057AD761C69B8DAA69B" bold="true" italics="true" pageId="0" pageNumber="85">Seeds</emphasis>
1-1.2
<normalizedToken id="99743D4E320F4BBB887A8B2CAD798AEC" originalValue="×">x</normalizedToken>
0.6-0.8 mm, obovoid, ochre to dark yellow, testa smooth.
</paragraph>
</subSubSection>
<subSubSection id="34F60CA5BD9A26977FE48410F090E52A" pageId="0" pageNumber="85" type="phenology">
<paragraph id="136A4EE247EE3ED4020DE45729AE2022" pageId="0" pageNumber="85">Phenology.</paragraph>
<paragraph id="BAA258A8A20E7D79076C66A88E09CDAC" pageId="0" pageNumber="85">Flowering from June to October; fruiting from September to November.</paragraph>
</subSubSection>
<subSubSection id="4E0BCC3AF9CFD043C11F263061426365" pageId="0" pageNumber="85" type="etymology">
<paragraph id="82A14F3860205D0F10DED69AD61B0ABE" pageId="0" pageNumber="85">Etymology.</paragraph>
<paragraph id="6599B78B4177B5D2B9CDAE5E8CC74FF9" pageId="0" pageNumber="85">The specific epithet refers to its distribution, Motuo County, Xizang, in China.</paragraph>
</subSubSection>
<subSubSection id="16B58EC73A57EE2A90ADDB4B7640A08E" pageId="0" pageNumber="85" type="vernacular name">
<paragraph id="F03CAAD766CB5DF2838E13BA168670F7" pageId="0" pageNumber="85">Vernacular name.</paragraph>
<paragraph id="14DEDC0C93E01F847362EC31AC101946" pageId="0" pageNumber="85">
Chinese:
<normalizedToken id="61BA5F09877E5148DC64873AE29CB051" originalValue="墨脱胡椒">墨脱胡椒</normalizedToken>
(
<normalizedToken id="18CF33E5EF27C45DC82AA3E07ABC3468" originalValue="mò">mo</normalizedToken>
<normalizedToken id="A371BB0215FE2AEE06840F050992C5E2" originalValue="tuō">tuō</normalizedToken>
<normalizedToken id="078561618FFB60B51D2DB8140CC90CF0" originalValue="hú">hu</normalizedToken>
<normalizedToken id="2698A6565A10DEE6DE94FE58202D2C33" originalValue="jiāo">jiao</normalizedToken>
).
<normalizedToken id="C9E54F08613073FCFD6FCA4F957DD4A5" originalValue="Mò">'Mo</normalizedToken>
<normalizedToken id="8D359A27928CD33191A03AADE2FBE4C6" originalValue="Tuō">Tuō'</normalizedToken>
is a place name, which is the literal translation of the specific epithet
<emphasis id="61FDD9EDCA7AC3970412166B96C356F1" italics="true" pageId="0" pageNumber="85">motuoense</emphasis>
and
<normalizedToken id="DC082E22A947EEA037FAAB13D0498A92" originalValue="hú">'hu</normalizedToken>
<normalizedToken id="62D8AD826639C22748E1DFA1052E8711" originalValue="jiāo">jiao'</normalizedToken>
is the Chinese name of
<taxonomicName id="79BFB01B720818B9F5F233F7D355F08E" class="Magnoliopsida" family="Piperaceae" genus="Piper" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Piper" order="Piperales" pageId="0" pageNumber="85" phylum="Tracheophyta" rank="genus">
<emphasis id="5327A965830C47DAB9FC93F6753CE9B4" italics="true" pageId="0" pageNumber="85">Piper</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="25638037D98D3D35B4BB9EE91D4B82DB" pageId="0" pageNumber="85" type="habitat">
<paragraph id="36AB2E796C401FE5E277C63B5C54C73C" pageId="0" pageNumber="85">Habitat and distribution.</paragraph>
<paragraph id="EE0B83F5044307376F6BA706958B964F" pageId="0" pageNumber="85">
The new species is currently known only from its type locality in Beibeng Town, Motuo County, Xizang Autonomous Region (Fig.
<figureCitation id="A53DC597E0EBF18E2642F7855112D39C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. The geographical distribution of P. motuoense sp. nov., P. yinkiangense and P. anisotis." figureDoi="10.3897/phytokeys.238.115494.figure3" httpUri="https://binary.pensoft.net/fig/980528" pageId="0" pageNumber="85">3</figureCitation>
). The new species grows very well in the type locality, as it has been recorded in three different sites (Beibeng, Damu and
<normalizedToken id="0C74F63B1E9210857BCB8DF5A23C818B" originalValue="Tigers">Tiger's</normalizedToken>
Mouth). It occurs in wet tropical rainforest at elevations of 490-1700 m and often climbs on taller trees or rocks.
</paragraph>
<caption id="55AFE22F0F601D016DB8BC17326258EF" doi="10.3897/phytokeys.238.115494.figure3" httpUri="https://binary.pensoft.net/fig/980528" pageId="0" pageNumber="85" start="Figure 3" startId="F3">
<paragraph id="3D7EA79D04D5A2042242DF1B217D6D80" pageId="0" pageNumber="85">
<emphasis id="0DBADF06766F8847F9058D5918B4DA97" bold="true" pageId="0" pageNumber="85">Figure 3.</emphasis>
The geographical distribution of
<taxonomicName id="D3A100F39F98DECBFA9E334EF49C8868" lsidName="P. motuoense" pageId="0" pageNumber="85" rank="species" species="motuoense">
<emphasis id="AA7B3F1EDA87D428EF9C5698EE14F3C2" italics="true" pageId="0" pageNumber="85">P. motuoense</emphasis>
</taxonomicName>
sp. nov.,
<taxonomicName id="6087C64908CFEDEC312A12189DC055A3" lsidName="P. yinkiangense" pageId="0" pageNumber="85" rank="species" species="yinkiangense">
<emphasis id="2485503741A105A13ABF665EC94B7BA1" italics="true" pageId="0" pageNumber="85">P. yinkiangense</emphasis>
</taxonomicName>
and
<taxonomicName id="69116582AAB7D48F857E8690008BCD96" lsidName="P. anisotis" pageId="0" pageNumber="85" rank="species" species="anisotis">
<emphasis id="D331949BE9B0D98BE060BEC0CDB890FA" italics="true" pageId="0" pageNumber="85">P. anisotis</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="82FB3893A2460A26A40C491FFB92D2BE" pageId="0" pageNumber="85" type="comments">
<paragraph id="C40D7FEE2B637CC6235B4F8844D871C3" pageId="0" pageNumber="85">Comments.</paragraph>
<paragraph id="922DCBB2E5133CA3670DC8564BD9B68D" pageId="0" pageNumber="85">
<taxonomicName id="8D7E7C5200621C46FAC25F4934B8D19B" lsidName="P. motuoense" pageId="0" pageNumber="85" rank="species" species="motuoense">
<emphasis id="F89D8234F916489552A65FB302EB449C" italics="true" pageId="0" pageNumber="85">P. motuoense</emphasis>
</taxonomicName>
was initially misidentified as
<taxonomicName id="C2FEEC4E38C10040CA9AEF772A249587" lsidName="P. anisotis" pageId="0" pageNumber="85" rank="species" species="anisotis">
<emphasis id="9A191C6479BE51E1C0E66EC1FA005B53" italics="true" pageId="0" pageNumber="85">P. anisotis</emphasis>
</taxonomicName>
from India, and Motuo is close to Assam in India. After examination of material of
<taxonomicName id="3CE2CE9A8B2D6FD2A3652542EA5BB048" lsidName="P. anisotis" pageId="0" pageNumber="85" rank="species" species="anisotis">
<emphasis id="BAE39BF35E8AAEEA317A70FA8F45C9C5" italics="true" pageId="0" pageNumber="85">P. anisotis</emphasis>
</taxonomicName>
we found that the new species differs in a number of characters. We also compared it with
<taxonomicName id="99404542BED5890EF05017A290E25D01" lsidName="P. yinkiangense" pageId="0" pageNumber="85" rank="species" species="yinkiangense">
<emphasis id="AAC1BD41F3E92EC0E9A42D5BB76F3E03" italics="true" pageId="0" pageNumber="85">P. yinkiangense</emphasis>
</taxonomicName>
. The differences amongst these three species are summarised in Table
<tableCitation id="6CAD722F8313320027A10CFDEA6DA8A6" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological comparison of key characteristics in P. motuoense, P. yinkiangense and P. anisotis." httpUri="http://table.plazi.org/id/C442A2EE40412627CD580B30ABB6BEB5" pageId="0" pageNumber="85" tableUuid="C442A2EE40412627CD580B30ABB6BEB5">1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>