432 lines
56 KiB
XML
432 lines
56 KiB
XML
<document id="446B1EF7C964592DFD8758CB481AB58C" ID="10.11646/zootaxa.4237.1.10SLASH10302" ID-CLB-Dataset="36435" ID-DOI="10.11646/zootaxa.4237.1.10" ID-GBIF-Dataset="8acf5793-4bfb-41f2-be40-5b4a8a80e44a" ID-ISSN="1175-5326" ID-Zenodo-Dep="322765" ID-ZooBank="21ABECB9-279F-42C5-AB95-767C3FF2D8F9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488192719361" checkinUser="plazi" docAuthor="Katongo, Cyprian, Seehausen, Ole & Snoeks, Jos" docDate="2017" docId="241387F7FFBFFF9AFF507239FB22FAE8" docLanguage="en" docName="zootaxa.4237.1.10SLASH10302.pdf" docOrigin="Zootaxa 4237 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudocrenilabrus pyrrhocaudalis Katongo, Seehausen & Snoeks, 2017, sp. nov." docType="treatment" docUuid="0D5769ED-5FA4-4413-9990-02FCCD6EC67B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="186" masterDocId="D82AFF8FFFBEFF9FFFC7756DFFFFFFAE" masterDocTitle="A new species of Pseudocrenilabrus (Perciformes: Cichlidae) from Lake Mweru in the Upper Congo River System" masterLastPageNumber="190" masterPageNumber="181" pageNumber="182" updateTime="1698437372663" updateUser="ExternalLinkService">
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<mods:mods id="61D2374503C8C6A9F602BCE232D845D1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E748FC6F429736519B11ACA71D2623CB">
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<mods:title id="CB9324D8E364E4DC18445E6424EE5BFA">A new species of Pseudocrenilabrus (Perciformes: Cichlidae) from Lake Mweru in the Upper Congo River System</mods:title>
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<mods:name id="507411C3B75F8141749425303D6666E4" type="personal">
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<mods:role id="64277E6D9CDDEF8BDB552BCF9A9AF04B">
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<mods:roleTerm id="F7DF81B081AE67933207AC31B47414F9">Author</mods:roleTerm>
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<mods:namePart id="06F1B908BE49EC757FBADCD38F7DC7BF">Katongo, Cyprian</mods:namePart>
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<mods:roleTerm id="5FC160836B4327DBCB6D12E73B452838">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D63227318DE6F377752D880A0E77CAAB">Seehausen, Ole</mods:namePart>
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</mods:name>
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<mods:name id="31F7C0E1B61A95FC8A7A6CE904264E9F" type="personal">
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<mods:roleTerm id="73890894517A6B05EBD97FF423D4A6E1">Author</mods:roleTerm>
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<mods:namePart id="0C585AE6CDD6A9EC8948D70730FC3F48">Snoeks, Jos</mods:namePart>
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<mods:typeOfResource id="B9186DBF1CA371130536086F331A4350">text</mods:typeOfResource>
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<mods:title id="D099D8A80362302E9A59E7FD41673DCF">Zootaxa</mods:title>
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<mods:part id="2883B29B2A0CA612D3480D9AB8907B16">
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<mods:date id="348A0CFD88FC263D9EA9873273ADAF75">2017</mods:date>
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<mods:number id="AD4E86F052C6F000A0659002ACAB2CDF">4237</mods:number>
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<mods:number id="78AC0A5C9333A3592D38CD18D6970892">1</mods:number>
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<mods:identifier id="49E749F1693CDC0B4B7334ED4FEB266E" type="DOI">10.11646/zootaxa.4237.1.10</mods:identifier>
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<treatment id="241387F7FFBFFF9AFF507239FB22FAE8" ID-DOI="http://doi.org/10.5281/zenodo.6044517" ID-GBIF-Taxon="127710641" ID-Zenodo-Dep="6044517" LSID="urn:lsid:zoobank.org:act:0D5769ED-5FA4-4413-9990-02FCCD6EC67B" httpUri="http://treatment.plazi.org/id/241387F7FFBFFF9AFF507239FB22FAE8" lastPageId="5" lastPageNumber="186" pageId="1" pageNumber="182">
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<subSubSection id="E4A0656AFFBFFF9EFF507239FE1FF821" pageId="1" pageNumber="182" type="nomenclature">
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<paragraph id="AC0536E1FFBFFF9EFF507239FD63F8C0" blockId="1.[151,1436,1876,2007]" box="[151,668,1876,1902]" pageId="1" pageNumber="182">
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<heading id="F74D818DFFBFFF9EFF507239FD63F8C0" bold="true" box="[151,668,1876,1902]" fontSize="11" level="1" pageId="1" pageNumber="182" reason="1">
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<emphasis id="9ECEEAF3FFBFFF9EFF507239FD63F8C0" bold="true" box="[151,668,1876,1902]" pageId="1" pageNumber="182">
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<taxonomicName id="6BBA4D62FFBFFF9EFF507239FDC9F8C0" ID-CoL="4NV7J" box="[151,566,1876,1902]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="182" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
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<emphasis id="9ECEEAF3FFBFFF9EFF507239FDC9F8C0" bold="true" box="[151,566,1876,1902]" italics="true" pageId="1" pageNumber="182">Pseudocrenilabrus pyrrhocaudalis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="85FD5788FFBFFF9EFDF97239FD63F8C0" box="[574,668,1876,1902]" pageId="1" pageNumber="182" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="AC0536E1FFBFFF9EFF50721BFE1FF821" blockId="1.[151,1436,1876,2007]" box="[151,480,1910,1935]" pageId="1" pageNumber="182">
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Fire-tailed
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<taxonomicName id="6BBA4D62FFBFFF9EFED4721BFE1FF821" box="[275,480,1910,1935]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="182" phylum="Chordata" rank="genus">Pseudocrenilabrus</taxonomicName>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E4A0656AFFBFFF9EFF5072F7FE71F81D" box="[151,398,1946,1971]" pageId="1" pageNumber="182" type="description">
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<paragraph id="AC0536E1FFBFFF9EFF5072F7FE71F81D" blockId="1.[151,1436,1876,2007]" box="[151,398,1946,1971]" pageId="1" pageNumber="182">
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||
<figureCitation id="34812A64FFBFFF9EFF5072F7FF1CF81D" box="[151,227,1946,1971]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1747]" captionTargetBox="[250,1335,464,1669]" captionTargetId="figure@2.[226,1360,447,1700]" captionTargetPageId="2" captionText="FIGURE 1. Pseudocrenilabrus pyrrhocaudalis sp. nov., male holotype (SAIAB 191528, 62.8 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322766/files/figure.png" pageId="1" pageNumber="182">Figs. 1</figureCitation>
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,
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<figureCitation id="34812A64FFBFFF9EFF3672F6FEFDF81D" box="[241,258,1947,1971]" captionStart="FIGURE 2" captionStartId="3.[151,250,1425,1447]" captionTargetBox="[241,1344,399,1380]" captionTargetId="figure@3.[226,1360,373,1404]" captionTargetPageId="3" captionText="FIGURE 2. Pseudocrenilabrus pyrrhocaudalis sp. nov, female, paratype (SAIAB 191530, 66.7 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322767/files/figure.png" pageId="1" pageNumber="182">2</figureCitation>
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.
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<tableCitation id="E138035AFFBFFF9EFECA72F7FE92F81D" box="[269,365,1946,1971]" captionStart="TABLE 2" captionStartId="6.[119,195,560,580]" captionText="TABLE 2. Morphometric ratios for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 2 1) from Lake Mweru. Ranges of P pyrrhocaudalis incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (73 mm SL), inđicate the relative contribution of the highlighteđ characters to the đifferences between the two species. These characters are eye điameter, interorbital wiđth," pageId="1" pageNumber="182">Tables 2</tableCitation>
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,
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<tableCitation id="E138035AFFBFFF9EFEBE72F6FE76F81D" box="[377,393,1947,1971]" captionStart="TABLE 3" captionStartId="7.[125,201,546,566]" captionText="TABLE 3. Meristic counts for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 21) from Lake Mweru. Summary values incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (" pageId="1" pageNumber="182">3</tableCitation>
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||
.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="E4A0656AFFBFFF9DFF5072D2FE20FE26" lastPageId="2" lastPageNumber="183" pageId="1" pageNumber="182" type="materials_examined">
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<paragraph id="AC0536E1FFBFFF9DFF5072D2FD9BFF7A" blockId="1.[151,1436,1876,2007]" lastBlockId="2.[151,1436,151,392]" lastPageId="2" lastPageNumber="183" pageId="1" pageNumber="182">
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<materialsCitation id="1CD23CBCFFBFFF9DFF5072D2FD9BFF7A" ID-GBIF-Occurrence="1437749144" collectingDate="2005-09-15" collectorName="C. Katongo & O. Seehausen" country="Zambia" county="Luapula-Congo River" lastPageId="2" lastPageNumber="183" latitude="-8.95" location="Kalobwa Beach" longLatPrecision="21" longitude="29.1" municipality="Lake Mweru" pageId="1" pageNumber="182" specimenCount="1" typeStatus="holotype">
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<typeStatus id="73018843FFBFFF9EFF5072D2FEFBF879" box="[151,260,1983,2007]" pageId="1" pageNumber="182" type="holotype">
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<emphasis id="9ECEEAF3FFBFFF9EFF5072D2FEFBF879" bold="true" box="[151,260,1983,2007]" pageId="1" pageNumber="182">Holotype</emphasis>
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</typeStatus>
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. SAIAB-191528, male
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<quantity id="6B429B04FFBFFF9EFDE672D3FD7AF878" box="[545,645,1982,2007]" metricMagnitude="-2" metricUnit="m" metricValue="6.279999999999999" pageId="1" pageNumber="182" unit="mm" value="62.8">62.8 mm</quantity>
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SL;
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<collectingCountry id="D4AD7671FFBFFF9EFD0072D3FCDFF879" box="[711,800,1982,2007]" name="Zambia" pageId="1" pageNumber="182">Zambia</collectingCountry>
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;
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<location id="A965603AFFBFFF9EFCF572D3FC15F879" LSID="urn:lsid:plazi:treatment:241387F7FFBFFF9AFF507239FB22FAE8:A965603AFFBFFF9EFCF572D3FC15F879" box="[818,1002,1982,2007]" country="Zambia" county="Luapula-Congo River" latitude="-8.95" longLatPrecision="21" longitude="29.1" municipality="Lake Mweru" name="Kalobwa Beach" pageId="1" pageNumber="182">Kalobwa Beach</location>
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(1,134 meters above sea level with coordinates
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<geoCoordinate id="C98E5026FFBCFF9DFEE675FAFE72FF1E" box="[289,397,151,176]" direction="south" orientation="latitude" pageId="2" pageNumber="183" precision="15" value="-8.95">8°57'0" S</geoCoordinate>
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and
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<geoCoordinate id="C98E5026FFBCFF9DFE0F75FAFDC7FF1E" box="[456,568,151,176]" direction="east" orientation="longitude" pageId="2" pageNumber="183" precision="15" value="29.1">29°6'0" E</geoCoordinate>
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),
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||
<collectingMunicipality id="4C61AC9BFFBCFF9DFD8875FAFD1BFF01" box="[591,740,151,176]" pageId="2" pageNumber="183">Lake Mweru</collectingMunicipality>
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,
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||
<collectingCounty id="45644E6DFFBCFF9DFD3675FAFC18FF1E" box="[753,999,151,176]" pageId="2" pageNumber="183">Luapula-Congo River</collectingCounty>
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system,
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<collectingMethod id="75FB4EF6FFBCFF9DFB8875FAFB49FF01" box="[1103,1206,151,176]" pageId="2" pageNumber="183">seine net</collectingMethod>
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,
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||
<collectorName id="014F5337FFBCFF9DFB0475F5FAB9FF1E" box="[1219,1350,152,176]" pageId="2" pageNumber="183">C. Katongo</collectorName>
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||
and
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||
<collectorName id="014F5337FFBCFF9DFA4675F5FEEFFF7B" pageId="2" pageNumber="183">O. Seehausen</collectorName>
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||
,
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||
<collectingDate id="C840E9C9FFBCFF9DFEDC75D1FE0AFF7B" box="[283,501,188,213]" pageId="2" pageNumber="183" value="2005-09-15">
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||
<date id="D8041021FFBCFF9DFEDC75D1FE0AFF7B" box="[283,501,188,213]" pageId="2" pageNumber="183" value="2005-09-15">15 September 2005</date>
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||
</collectingDate>
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(
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||
<figureCitation id="34812A64FFBCFF9DFDC375D1FDBCFF7A" box="[516,579,188,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1747]" captionTargetBox="[250,1335,464,1669]" captionTargetId="figure@2.[226,1360,447,1700]" captionTargetPageId="2" captionText="FIGURE 1. Pseudocrenilabrus pyrrhocaudalis sp. nov., male holotype (SAIAB 191528, 62.8 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322766/files/figure.png" pageId="2" pageNumber="183">Fig. 1</figureCitation>
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||
a).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="AC0536E1FFBCFF9DFF00758DFE20FE26" blockId="2.[151,1436,151,392]" pageId="2" pageNumber="183">
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||
<materialsCitation id="1CD23CBCFFBCFF9DFF00758DFDB1FEB2" ID-GBIF-Occurrence="1437749145" collectionCode="MRAC" collectorName="P. van Zwieten" country="Zambia" county="Lake Mweru" location="Luapula-Congo River" municipality="Mukwakwa" pageId="2" pageNumber="183" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="73018843FFBCFF9DFF00758DFEC0FF56" box="[199,319,224,248]" pageId="2" pageNumber="183" type="paratype">
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||
<emphasis id="9ECEEAF3FFBCFF9DFF00758DFEC0FF56" bold="true" box="[199,319,224,248]" pageId="2" pageNumber="183">Paratypes</emphasis>
|
||
</typeStatus>
|
||
.
|
||
<collectionCode id="CAABAE24FFBCFF9DFE89758DFE5FFF56" box="[334,416,224,248]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Mus<75>e Royal de l<>Afrique Centrale" pageId="2" pageNumber="183">MRAC</collectionCode>
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||
A9-034-P-0238-247,
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<quantity id="6B429B04FFBCFF9DFD5D75B2FCC1FF59" box="[666,830,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="6.519999999999999" metricValueMax="7.31" metricValueMin="5.7299999999999995" pageId="2" pageNumber="183" unit="mm" value="65.19999999999999" valueMax="73.1" valueMin="57.3">57.3–73.1 mm</quantity>
|
||
SL,
|
||
<location id="A965603AFFBCFF9DFCB175B2FB93FF56" LSID="urn:lsid:plazi:treatment:241387F7FFBFFF9AFF507239FB22FAE8:A965603AFFBCFF9DFCB175B2FB93FF56" box="[886,1132,223,248]" country="Zambia" county="Lake Mweru" municipality="Mukwakwa" name="Luapula-Congo River" pageId="2" pageNumber="183">Luapula-Congo River</location>
|
||
system,
|
||
<collectingMunicipality id="4C61AC9BFFBCFF9DFB1675B2FAA7FF56" box="[1233,1368,223,248]" pageId="2" pageNumber="183">Mukwakwa</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="45644E6DFFBCFF9DFAA375B2FF14FEB2" pageId="2" pageNumber="183">Lake Mweru</collectingCounty>
|
||
,
|
||
<collectingCountry id="D4AD7671FFBCFF9DFF307469FEB0FEB3" box="[247,335,260,285]" name="Zambia" pageId="2" pageNumber="183">Zambia</collectingCountry>
|
||
,
|
||
<collectorName id="014F5337FFBCFF9DFE9C7468FDF9FEB3" box="[347,518,260,285]" pageId="2" pageNumber="183">P. van Zwieten</collectorName>
|
||
, 1994
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="1CD23CBCFFBCFF9DFD9D7469FACBFEEE" ID-GBIF-Occurrence="1437749143" collectionCode="MRAC" collectorName="C. Katongo" country="Zambia" county="Lake Mweru" location="Luapula-Congo River" municipality="Mwatishi River" pageId="2" pageNumber="183" specimenCount="1" typeStatus="paratype">
|
||
<collectionCode id="CAABAE24FFBCFF9DFD9D7469FD53FEB2" box="[602,684,260,284]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Mus<75>e Royal de l<>Afrique Centrale" pageId="2" pageNumber="183">MRAC</collectionCode>
|
||
A4-025-P0103-07 and
|
||
<collectionCode id="CAABAE24FFBCFF9DFC7F7469FBF4FEB2" box="[952,1035,260,284]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Mus<75>e Royal de l<>Afrique Centrale" pageId="2" pageNumber="183">MRAC</collectionCode>
|
||
A4-025-0137-38,
|
||
<quantity id="6B429B04FFBCFF9DFB197469FA92FEB2" box="[1246,1389,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="5.025" metricValueMax="5.4" metricValueMin="4.65" pageId="2" pageNumber="183" unit="mm" value="50.25" valueMax="54.0" valueMin="46.5">46.5–54 mm</quantity>
|
||
SL,
|
||
<location id="A965603AFFBCFF9DFF50744AFE6CFEEE" LSID="urn:lsid:plazi:treatment:241387F7FFBFFF9AFF507239FB22FAE8:A965603AFFBCFF9DFF50744AFE6CFEEE" box="[151,403,295,320]" country="Zambia" county="Lake Mweru" municipality="Mwatishi River" name="Luapula-Congo River" pageId="2" pageNumber="183">Luapula-Congo River</location>
|
||
system,
|
||
<collectingMunicipality id="4C61AC9BFFBCFF9DFDC1744AFD42FEEE" box="[518,701,295,320]" pageId="2" pageNumber="183">Mwatishi River</collectingMunicipality>
|
||
/
|
||
<collectingCounty id="45644E6DFFBCFF9DFD02744AFCA2FE91" box="[709,861,295,320]" pageId="2" pageNumber="183">Lake Mweru</collectingCounty>
|
||
confluence, gillnet,
|
||
<collectorName id="014F5337FFBCFF9DFB9D7445FB19FEEE" box="[1114,1254,296,320]" pageId="2" pageNumber="183">C. Katongo</collectorName>
|
||
, 2002
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="1CD23CBCFFBCFF9DFA8F7445FE20FE26" ID-GBIF-Occurrence="1437749146" collectionCode="SAIAB" collectorName="Kalobwa & Lake Mweru, C. & O. Seehausen & September" country="Zambia" location="Luapula-Congo River" municipality="Lake Mweru" pageId="2" pageNumber="183" specimenCode="SAIAB 191530" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="FC1C9E9AFFBCFF9DFA8F7445FF09FECB" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="2" pageNumber="183">
|
||
<collectionCode id="CAABAE24FFBCFF9DFA8F7445FA63FEEE" box="[1352,1436,296,320]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="2" pageNumber="183">SAIAB</collectionCode>
|
||
191530
|
||
</specimenCode>
|
||
(1-13),
|
||
<quantity id="6B429B04FFBCFF9DFEF87421FE1DFECA" box="[319,482,332,357]" metricMagnitude="-2" metricUnit="m" metricValue="5.68" metricValueMax="6.83" metricValueMin="4.529999999999999" pageId="2" pageNumber="183" unit="mm" value="56.8" valueMax="68.3" valueMin="45.3">45.3–68.3 mm</quantity>
|
||
SL,
|
||
<location id="A965603AFFBCFF9DFDD07421FCF3FECB" LSID="urn:lsid:plazi:treatment:241387F7FFBFFF9AFF507239FB22FAE8:A965603AFFBCFF9DFDD07421FCF3FECB" box="[535,780,332,357]" country="Zambia" municipality="Lake Mweru" name="Luapula-Congo River" pageId="2" pageNumber="183">Luapula-Congo River</location>
|
||
system,
|
||
<collectorName id="014F5337FFBCFF9DFCB77421FC2AFECB" box="[880,981,332,357]" pageId="2" pageNumber="183">Kalobwa</collectorName>
|
||
beach,
|
||
<collectorName id="014F5337FFBCFF9DFBEC7421FB1CFECA" box="[1067,1251,332,357]" pageId="2" pageNumber="183">
|
||
<collectingMunicipality id="4C61AC9BFFBCFF9DFBEC7421FB41FECA" box="[1067,1214,332,357]" pageId="2" pageNumber="183">Lake Mweru</collectingMunicipality>
|
||
, C.
|
||
</collectorName>
|
||
Katongo and
|
||
<collectorName id="014F5337FFBCFF9DFA467421FEEFFE26" pageId="2" pageNumber="183">O. Seehausen</collectorName>
|
||
,
|
||
<collectorName id="014F5337FFBCFF9DFEDC7402FE6AFE26" box="[283,405,367,392]" pageId="2" pageNumber="183">September</collectorName>
|
||
, 2005.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F8C56669FFBCFF9DFF5073D0FD12F95C" httpUri="https://zenodo.org/record/322766/files/figure.png" pageId="2" pageNumber="183" targetBox="[250,1335,464,1669]" targetPageId="2">
|
||
<paragraph id="AC0536E1FFBCFF9DFF5073D0FD12F95C" blockId="2.[151,1436,1725,1778]" pageId="2" pageNumber="183">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFF5073D0FEEBF97D" bold="true" box="[151,276,1725,1747]" pageId="2" pageNumber="183">FIGURE 1.</emphasis>
|
||
<taxonomicName id="6BBA4D62FFBCFF9DFEDD73D0FD85F97D" box="[282,634,1725,1747]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="183" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFEDD73D0FD85F97D" box="[282,634,1725,1747]" italics="true" pageId="2" pageNumber="183">Pseudocrenilabrus pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBCFF9DFD4673D0FD2DF97D" box="[641,722,1725,1747]" pageId="2" pageNumber="183" rank="species">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFD4673D0FD2DF97D" bold="true" box="[641,722,1725,1747]" pageId="2" pageNumber="183">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, male holotype (SAIAB 191528, 62.8 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E4A0656AFFBCFF9CFF007275FDA4FEEE" lastPageId="3" lastPageNumber="184" pageId="2" pageNumber="183" type="diagnosis">
|
||
<paragraph id="AC0536E1FFBCFF9CFF007275FDA4FEEE" blockId="2.[151,1436,1816,2022]" lastBlockId="3.[151,1436,151,320]" lastPageId="3" lastPageNumber="184" pageId="2" pageNumber="183">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFF007275FEBEF89F" bold="true" box="[199,321,1816,1841]" pageId="2" pageNumber="183">Diagnosis.</emphasis>
|
||
<taxonomicName id="6BBA4D62FFBCFF9DFE887274FD25F89F" box="[335,730,1817,1841]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="183" phylum="Chordata" rank="species" species="pyrrhocaudalis">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFE887274FD25F89F" box="[335,730,1817,1841]" italics="true" pageId="2" pageNumber="183">Pseudocrenilabrus pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
can be differentiated from the other species of the genus
|
||
<taxonomicName id="6BBA4D62FFBCFF9DFF507250FE95F8FB" box="[151,362,1853,1877]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="183" phylum="Chordata" rank="genus">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFF507250FE95F8FB" box="[151,362,1853,1877]" italics="true" pageId="2" pageNumber="183">Pseudocrenilabrus</emphasis>
|
||
</taxonomicName>
|
||
on the basis of its colour pattern and its subtruncate caudal fin. Male
|
||
<taxonomicName id="6BBA4D62FFBCFF9DFB437253FAB4F8FB" box="[1156,1355,1853,1877]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="183" phylum="Chordata" rank="species" species="pyrrhocaudalis">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFB437253FAB4F8FB" box="[1156,1355,1853,1877]" italics="true" pageId="2" pageNumber="183">P. pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
have a unique colour pattern characterized by some orange colour on their anal and caudal fins that can become bright orange-red in breeding males, extending over the proximal parts of the anal and caudal fins and the distal part of the caudal and the upper and lower parts of the caudal peduncle (
|
||
<figureCitation id="34812A64FFBCFF9DFC4E72C4FC34F86F" box="[905,971,1961,1986]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1747]" captionTargetBox="[250,1335,464,1669]" captionTargetId="figure@2.[226,1360,447,1700]" captionTargetPageId="2" captionText="FIGURE 1. Pseudocrenilabrus pyrrhocaudalis sp. nov., male holotype (SAIAB 191528, 62.8 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322766/files/figure.png" pageId="2" pageNumber="183">Fig. 1</figureCitation>
|
||
a). In addition,
|
||
<taxonomicName id="6BBA4D62FFBCFF9DFB4772C7FAB6F86F" box="[1152,1353,1961,1985]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="183" phylum="Chordata" rank="species" species="pyrrhocaudalis">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFB4772C7FAB6F86F" box="[1152,1353,1961,1985]" italics="true" pageId="2" pageNumber="183">P. pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from its sympatric congener
|
||
<taxonomicName id="6BBA4D62FFBCFF9DFD4D72A3FCE8F84B" box="[650,791,1997,2021]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="183" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBCFF9DFD4D72A3FCE8F84B" box="[650,791,1997,2021]" italics="true" pageId="2" pageNumber="183">P. philander</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="34812A64FFBCFF9DFCED72A0FC8EF848" box="[810,881,1997,2022]" captionStart="FIGURE 3" captionStartId="3.[151,250,1955,1977]" captionTargetBox="[340,1247,1528,1934]" captionTargetId="figure@3.[340,1247,1528,1934]" captionTargetPageId="3" captionText="FIGURE 3. Pseudocrenilabrus philander philander, male (SAIAB 191531, 70.4 mm SL) from Isokwe Island, Lake Mweru." httpUri="https://zenodo.org/record/322768/files/figure.png" pageId="2" pageNumber="183">Fig. 3</figureCitation>
|
||
), by a combination of the following characters (
|
||
<figureCitation id="34812A64FFBDFF9CFF6775FAFF18FF1E" box="[160,231,151,176]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1747]" captionTargetBox="[250,1335,464,1669]" captionTargetId="figure@2.[226,1360,447,1700]" captionTargetPageId="2" captionText="FIGURE 1. Pseudocrenilabrus pyrrhocaudalis sp. nov., male holotype (SAIAB 191528, 62.8 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322766/files/figure.png" pageId="3" pageNumber="184">Figs.1</figureCitation>
|
||
,
|
||
<figureCitation id="34812A64FFBDFF9CFF3175F5FEF9FF1E" box="[246,262,152,176]" captionStart="FIGURE 2" captionStartId="3.[151,250,1425,1447]" captionTargetBox="[241,1344,399,1380]" captionTargetId="figure@3.[226,1360,373,1404]" captionTargetPageId="3" captionText="FIGURE 2. Pseudocrenilabrus pyrrhocaudalis sp. nov, female, paratype (SAIAB 191530, 66.7 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322767/files/figure.png" pageId="3" pageNumber="184">2</figureCitation>
|
||
,
|
||
<figureCitation id="34812A64FFBDFF9CFED375F5FEDAFF1E" box="[276,293,152,176]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2002]" captionTargetBox="[229,1353,1192,1944]" captionTargetId="figure@4.[225,1359,1187,1948]" captionTargetPageId="4" captionText="FIGURE 4. Scatter plot of the specimens of Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) and P. philander (n = 21) on the first and second axes of a Principal Component analysis on log-transformed measurements." httpUri="https://zenodo.org/record/322769/files/figure.png" pageId="3" pageNumber="184">4</figureCitation>
|
||
;
|
||
<tableCitation id="E138035AFFBDFF9CFEF475FAFE6AFF1E" box="[307,405,151,176]" captionStart="TABLE 2" captionStartId="6.[119,195,560,580]" captionText="TABLE 2. Morphometric ratios for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 2 1) from Lake Mweru. Ranges of P pyrrhocaudalis incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (73 mm SL), inđicate the relative contribution of the highlighteđ characters to the đifferences between the two species. These characters are eye điameter, interorbital wiđth," pageId="3" pageNumber="184">Tables 2</tableCitation>
|
||
,
|
||
<tableCitation id="E138035AFFBDFF9CFE6475F5FE4CFF1E" box="[419,435,152,176]" captionStart="TABLE 3" captionStartId="7.[125,201,546,566]" captionText="TABLE 3. Meristic counts for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 21) from Lake Mweru. Summary values incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (" pageId="3" pageNumber="184">3</tableCitation>
|
||
): pelvic fin white (vs. black), posterior part of dorsal fin orange (vs. olive green in the other species), comparatively thinner lips (vs. comparatively thicker lips), larger eye diameter 27.0–39.3% HL (vs 22.9–33.1% HL), narrower head width 37.0–47.3% HL (vs. 38.5–53.7% HL), narrower interorbital distance 14.6– 22.3% HL (vs. 20.5–29.4% HL) and a more slender caudal peduncle 8.8–11.8% SL (vs. 11.3–13.9% SL), and 50.0–64.7% CPL (vs 64.3–93.5% CPL).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F8C56669FFBDFF9CFF5070FCFD12FA68" httpUri="https://zenodo.org/record/322767/files/figure.png" pageId="3" pageNumber="184" targetBox="[241,1344,399,1380]" targetPageId="3">
|
||
<paragraph id="AC0536E1FFBDFF9CFF5070FCFD12FA68" blockId="3.[151,1436,1425,1478]" pageId="3" pageNumber="184">
|
||
<emphasis id="9ECEEAF3FFBDFF9CFF5070FCFEF3FA09" bold="true" box="[151,268,1425,1447]" pageId="3" pageNumber="184">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="6BBA4D62FFBDFF9CFEDE70FCFD88FA09" box="[281,631,1425,1447]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="184" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBDFF9CFEDE70FCFD88FA09" box="[281,631,1425,1447]" italics="true" pageId="3" pageNumber="184">Pseudocrenilabrus pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBDFF9CFDB970FCFD3DFA09" box="[638,706,1425,1447]" pageId="3" pageNumber="184" rank="species">
|
||
<emphasis id="9ECEEAF3FFBDFF9CFDB970FCFD3DFA09" bold="true" box="[638,706,1425,1447]" pageId="3" pageNumber="184">sp.nov</emphasis>
|
||
</taxonomicNameLabel>
|
||
, female, paratype (SAIAB 191530, 66.7 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F8C56669FFBDFF9CFF5072CEFA79F817" box="[151,1414,1955,1977]" httpUri="https://zenodo.org/record/322768/files/figure.png" pageId="3" pageNumber="184" targetBox="[340,1247,1528,1934]" targetPageId="3">
|
||
<paragraph id="AC0536E1FFBDFF9CFF5072CEFA79F817" blockId="3.[151,1414,1955,1977]" box="[151,1414,1955,1977]" pageId="3" pageNumber="184">
|
||
<emphasis id="9ECEEAF3FFBDFF9CFF5072CEFEF2F817" bold="true" box="[151,269,1955,1977]" pageId="3" pageNumber="184">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="6BBA4D62FFBDFF9CFEDE72CEFD57F817" box="[281,680,1955,1977]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="184" phylum="Chordata" rank="subSpecies" species="philander" subSpecies="philander">
|
||
<emphasis id="9ECEEAF3FFBDFF9CFEDE72CEFD57F817" box="[281,680,1955,1977]" italics="true" pageId="3" pageNumber="184">Pseudocrenilabrus philander philander</emphasis>
|
||
</taxonomicName>
|
||
, male (SAIAB 191531, 70.4 mm SL) from Isokwe Island, Lake Mweru.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E4A0656AFFBAFF9BFF0075FAFAB0FBD3" pageId="4" pageNumber="185" type="description">
|
||
<paragraph id="AC0536E1FFBAFF9BFF0075FAFF3DFD92" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="185">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFF0075FAFEA6FF1E" bold="true" box="[199,345,151,176]" pageId="4" pageNumber="185">Description.</emphasis>
|
||
In both sexes: body relatively deep, head profile somewhat convex, mouth inclined upwards with relatively thin lips, interorbital distance narrow, caudal fin sub-truncate. Lower pharyngeal bone relatively slender and slightly longer than wide. Pharyngeal teeth all fine; those of the posterior row clearly larger than the others. Teeth of the two median rows somewhat enlarged. There is a small gradient in the orientation of the major cusp of the pharyngeal teeth from slightly backwards on the anterior parts of the pharyngeal jaw towards more erect posteriorly and slightly forward in the posteriormost rows. Breeding males with a bright orange coloration on caudal and anal fins; an orange spot on the anal fin, distal parts of caudal and anal fins white. Base of the caudal fin bright orange more than 2/3 of fin. Head and upper lateral part of body grey, while the lower part yellowish orange. Dorsal fin with white lateral dots arranged as stripes radiating from base to tips of dorsal-fin rays. Pelvic fins bright white (
|
||
<figureCitation id="34812A64FFBAFF9BFF2374B1FEDBFE5A" box="[228,292,476,501]" captionStart="FIGURE 1" captionStartId="2.[151,250,1725,1747]" captionTargetBox="[250,1335,464,1669]" captionTargetId="figure@2.[226,1360,447,1700]" captionTargetPageId="2" captionText="FIGURE 1. Pseudocrenilabrus pyrrhocaudalis sp. nov., male holotype (SAIAB 191528, 62.8 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322766/files/figure.png" pageId="4" pageNumber="185">Fig. 1</figureCitation>
|
||
a). Females generally grey, with no anal-fin spot; pelvic fins bright white; bright orange lower half of caudal; some orange flashes at base of caudal and anal fins; some spots and streaks on dorsal and caudal fins (
|
||
<figureCitation id="34812A64FFBAFF9BFAA87492FF58FD92" captionStart="FIGURE 2" captionStartId="3.[151,250,1425,1447]" captionTargetBox="[241,1344,399,1380]" captionTargetId="figure@3.[226,1360,373,1404]" captionTargetPageId="3" captionText="FIGURE 2. Pseudocrenilabrus pyrrhocaudalis sp. nov, female, paratype (SAIAB 191530, 66.7 mm SL) from Kalobwa Beach, Lake Mweru in live (a) and in preserved colour pattern (b)." httpUri="https://zenodo.org/record/322767/files/figure.png" pageId="4" pageNumber="185">Fig. 2</figureCitation>
|
||
a).
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBAFF9BFF007725FAB0FBD3" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="185">
|
||
A scatter plot of mainly male specimens of
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFD7B7724FC7BFDCE" box="[700,900,584,608]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFD7B7724FC7BFDCE" box="[700,900,584,608]" italics="true" pageId="4" pageNumber="185">P. pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBAFF9BFC4B7725FC18FDCE" box="[908,999,584,608]" pageId="4" pageNumber="185" rank="species">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFC4B7725FC18FDCE" bold="true" box="[908,999,584,608]" pageId="4" pageNumber="185">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and Lake Mweru
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFB7D7724FABBFDCE" box="[1210,1348,584,608]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFB7D7724FABBFDCE" box="[1210,1348,584,608]" italics="true" pageId="4" pageNumber="185">P. philander</emphasis>
|
||
</taxonomicName>
|
||
on the first and second axes of a PCA on log transformed measurements (
|
||
<figureCitation id="34812A64FFBAFF9BFC1A7701FBB9FD2A" box="[989,1094,620,645]" captionStart="FIGURE 4" captionStartId="4.[151,250,1980,2002]" captionTargetBox="[229,1353,1192,1944]" captionTargetId="figure@4.[225,1359,1187,1948]" captionTargetPageId="4" captionText="FIGURE 4. Scatter plot of the specimens of Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) and P. philander (n = 21) on the first and second axes of a Principal Component analysis on log-transformed measurements." httpUri="https://zenodo.org/record/322769/files/figure.png" pageId="4" pageNumber="185">Figure 4</figureCitation>
|
||
) shows that there are clear morphological differences between the two species. The characters with the highest loadings on the second axis (responsible for the observed morphological differences) are caudal peduncle length (CPL), interorbital width (IOW), eye diameter (ED), anal fin base (AFB), head width (HW) and caudal peduncle depth (CPD), (
|
||
<tableCitation id="E138035AFFBAFF9BFAF277BAFA74FD5E" box="[1333,1419,727,752]" captionStart="TABLE 1" captionStartId="5.[151,239,324,347]" captionText="TABLE 1. Loadings of the variables on the first three principal components of a PCA on the log-transformed measurements of specimens of Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) and P. philander (n = 21). The highlighted results indicate the characters with the highest loadings on PC 2 and PC 3. These characters are eye diameter, interorbital width, head width, anal fin base, caudal peduncle length and caudal peduncle depth." pageId="4" pageNumber="185">Table 1</tableCitation>
|
||
). Comparisons of morphometric ratios between
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFD6F7790FC8FFCBA" box="[680,880,764,788]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFD6F7790FC8FFCBA" box="[680,880,764,788]" italics="true" pageId="4" pageNumber="185">P. pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBAFF9BFCBF7791FC2BFCBA" box="[888,980,764,788]" pageId="4" pageNumber="185" rank="species">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFCBF7791FC2BFCBA" bold="true" box="[888,980,764,788]" pageId="4" pageNumber="185">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFBCA7790FB67FCBA" box="[1037,1176,764,788]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFBCA7790FB67FCBA" box="[1037,1176,764,788]" italics="true" pageId="4" pageNumber="185">P. philander</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="E138035AFFBAFF9BFB6F7791FB00FCBA" box="[1192,1279,764,789]" captionStart="TABLE 2" captionStartId="6.[119,195,560,580]" captionText="TABLE 2. Morphometric ratios for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 2 1) from Lake Mweru. Ranges of P pyrrhocaudalis incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (73 mm SL), inđicate the relative contribution of the highlighteđ characters to the đifferences between the two species. These characters are eye điameter, interorbital wiđth," pageId="4" pageNumber="185">Table 2</tableCitation>
|
||
) indicate that the characters responsible for the observed morphological differences are eye diameter as a proportion of head length (ED_HL), interorbital width as a proportion of head length (IOW_HL) and as a proportion of head width (IOW_HW), head width as a proportion of head length (HW_HL), caudal peduncle depth as a proportion of standard length (CPD_ SL) and as a proportion of caudal peduncle length (CPD_ CPL). Comparisons of meristic counts between
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFE8D76DCFDEFFC66" box="[330,528,944,968]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFE8D76DCFDEFFC66" box="[330,528,944,968]" italics="true" pageId="4" pageNumber="185">P. pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBAFF9BFDD176DDFD90FC66" box="[534,623,944,968]" pageId="4" pageNumber="185" rank="species">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFDD176DDFD90FC66" bold="true" box="[534,623,944,968]" pageId="4" pageNumber="185">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFD6376DCFCD4FC66" box="[676,811,944,968]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFD6376DCFCD4FC66" box="[676,811,944,968]" italics="true" pageId="4" pageNumber="185">P. philander</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<tableCitation id="E138035AFFBAFF9BFCFE76C2FC72FC66" box="[825,909,943,968]" captionStart="TABLE 3" captionStartId="7.[125,201,546,566]" captionText="TABLE 3. Meristic counts for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 21) from Lake Mweru. Summary values incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (" pageId="4" pageNumber="185">Table 3</tableCitation>
|
||
) indicate that the characters responsible for the differences are number of upper jaw teeth (UJT), number of upper jaw inner rows (UJIR), dorsal spiny rays (Dspiny), anal soft rays (Asoft) and number of scales around the caudal peduncle (CP). The columns with Mann Whitney U test results (probabilities) in both
|
||
<tableCitation id="E138035AFFBAFF9BFD577171FD11FB9A" box="[656,750,1052,1077]" captionStart="TABLE 2" captionStartId="6.[119,195,560,580]" captionText="TABLE 2. Morphometric ratios for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 2 1) from Lake Mweru. Ranges of P pyrrhocaudalis incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (73 mm SL), inđicate the relative contribution of the highlighteđ characters to the đifferences between the two species. These characters are eye điameter, interorbital wiđth," pageId="4" pageNumber="185">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="E138035AFFBAFF9BFCE37171FCCDFB9A" box="[804,818,1052,1076]" captionStart="TABLE 3" captionStartId="7.[125,201,546,566]" captionText="TABLE 3. Meristic counts for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 21) from Lake Mweru. Summary values incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (" pageId="4" pageNumber="185">3</tableCitation>
|
||
were generated using
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFBEB7170FB0DFB9A" box="[1068,1266,1052,1076]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFBEB7170FB0DFB9A" box="[1068,1266,1052,1076]" italics="true" pageId="4" pageNumber="185">P. pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBAFF9BFB3F7171FAADFB9A" box="[1272,1362,1052,1076]" pageId="4" pageNumber="185" rank="species">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFB3F7171FAADFB9A" bold="true" box="[1272,1362,1052,1076]" pageId="4" pageNumber="185">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFA407170FEFBFBF6" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFA407170FEFBFBF6" italics="true" pageId="4" pageNumber="185">P. philander</emphasis>
|
||
</taxonomicName>
|
||
specimens of similar size class (
|
||
<quantity id="6B429B04FFBAFF9BFDB5712DFD1AFBF6" box="[626,741,1088,1112]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" metricValueMax="7.3" metricValueMin="4.4" pageId="4" pageNumber="185" unit="mm" value="58.5" valueMax="73.0" valueMin="44.0">44–73mm</quantity>
|
||
SL). These results (in the last column of each of
|
||
<tableCitation id="E138035AFFBAFF9BFAC87152FA92FBF6" box="[1295,1389,1087,1112]" captionStart="TABLE 2" captionStartId="6.[119,195,560,580]" captionText="TABLE 2. Morphometric ratios for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 2 1) from Lake Mweru. Ranges of P pyrrhocaudalis incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (73 mm SL), inđicate the relative contribution of the highlighteđ characters to the đifferences between the two species. These characters are eye điameter, interorbital wiđth," pageId="4" pageNumber="185">Tables 2</tableCitation>
|
||
and
|
||
<tableCitation id="E138035AFFBAFF9BFF507109FF5AFBD2" box="[151,165,1124,1148]" captionStart="TABLE 3" captionStartId="7.[125,201,546,566]" captionText="TABLE 3. Meristic counts for Pseudocrenilabrus pyrrhocaudalis sp. nov. (n = 31) anđ P. philander (n = 21) from Lake Mweru. Summary values incluđe holotype. The highlighteđ results inđicate the characters which are đifferent between the two. The Mann Whitney U-test results (probabilities) in the last column which were baseđ on specimens within a similar size class (44 (" pageId="4" pageNumber="185">3</tableCitation>
|
||
) indicate the relative contribution of the highlighted characters to the differences between the two species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="F8C56669FFBAFF9BFF5072D1FBA5F85F" httpUri="https://zenodo.org/record/322769/files/figure.png" pageId="4" pageNumber="185" targetBox="[229,1353,1192,1944]" targetPageId="4">
|
||
<paragraph id="AC0536E1FFBAFF9BFF5072D1FBA5F85F" blockId="4.[151,1436,1980,2033]" pageId="4" pageNumber="185">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFF5072D1FEECF87C" bold="true" box="[151,275,1980,2002]" pageId="4" pageNumber="185">FIGURE 4.</emphasis>
|
||
Scatter plot of the specimens of
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFDA472D1FC3DF87C" box="[611,962,1980,2002]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFDA472D1FC3DF87C" box="[611,962,1980,2002]" italics="true" pageId="4" pageNumber="185">Pseudocrenilabrus pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBAFF9BFC0E72D1FBE5F87C" box="[969,1050,1980,2002]" pageId="4" pageNumber="185" rank="species">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFC0E72D1FBE5F87C" bold="true" box="[969,1050,1980,2002]" pageId="4" pageNumber="185">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(n = 31) and
|
||
<taxonomicName id="6BBA4D62FFBAFF9BFB6172D3FADCF87C" box="[1190,1315,1980,2003]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="185" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBAFF9BFB6172D3FADCF87C" box="[1190,1315,1980,2003]" italics="true" pageId="4" pageNumber="185">P. philander</emphasis>
|
||
</taxonomicName>
|
||
(n = 21) on the first and second axes of a Principal Component analysis on log-transformed measurements.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="E4A0656AFFBBFF9AFF0075F5FB19FD61" pageId="5" pageNumber="186" type="etymology">
|
||
<paragraph id="AC0536E1FFBBFF9AFF0075F5FC97FEB3" blockId="5.[151,1436,151,285]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF0075F5FEB1FF1E" bold="true" box="[199,334,152,176]" pageId="5" pageNumber="186">Etymology.</emphasis>
|
||
Named
|
||
<taxonomicName id="6BBA4D62FFBBFF9AFE7375F5FCC2FF1E" box="[436,829,152,176]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="186" phylum="Chordata" rank="species" species="pyrrhocaudalis">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFE7375F5FCC2FF1E" box="[436,829,152,176]" italics="true" pageId="5" pageNumber="186">Pseudocrenilabrus pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
because this species has a bright orange tail which resembles a flame of fire. The common name fire-tailed
|
||
<taxonomicName id="6BBA4D62FFBBFF9AFCD775D1FC22FF7B" box="[784,989,188,213]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="186" phylum="Chordata" rank="genus">Pseudocrenilabrus</taxonomicName>
|
||
is proposed for this species.
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF0075B2FEA6FF56" bold="true" box="[199,345,223,248]" pageId="5" pageNumber="186">Distribution</emphasis>
|
||
. Probably endemic to Lake Mweru where it has been found near the beaches at Kalobwa, Kabuta, Ntoto and Kashikishi and at the Mwatishi River estuary (
|
||
<figureCitation id="34812A64FFBBFF9AFCD17469FCA5FEB3" box="[790,858,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,913,935]" captionTargetBox="[261,1327,201,873]" captionTargetId="figure@8.[247,1335,196,878]" captionTargetPageId="8" captionText="FIGURE 5. Known distribution of Pseudocrenilabrus pyrrhocaudalis sp. nov. in Lake Mweru and the location of the study area within Africa." httpUri="https://zenodo.org/record/322770/files/figure.png" pageId="5" pageNumber="186">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="F8C56669FFBBFF9AFF507429FB6BFE11" pageId="5" pageNumber="186">
|
||
<paragraph id="AC0536E1FFBBFF9AFF507429FB6BFE11" blockId="5.[151,1436,324,1350]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF507429FEF0FEF5" bold="true" box="[151,271,324,347]" pageId="5" pageNumber="186">TABLE 1.</emphasis>
|
||
Loadings of the variables on the first three principal components of a PCA on the log-transformed measurements of specimens of
|
||
<taxonomicName id="6BBA4D62FFBBFF9AFE327409FC95FED2" box="[501,874,356,380]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="186" phylum="Chordata" rank="species" species="pyrrhocaudalis" status="sp. nov.">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFE327409FC95FED2" box="[501,874,356,380]" italics="true" pageId="5" pageNumber="186">Pseudocrenilabrus pyrrhocaudalis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="85FD5788FFBBFF9AFCB27408FC30FED2" box="[885,975,357,380]" pageId="5" pageNumber="186" rank="species">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFCB27408FC30FED2" bold="true" box="[885,975,357,380]" pageId="5" pageNumber="186">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(n = 31) and
|
||
<taxonomicName id="6BBA4D62FFBBFF9AFBB47408FB07FED2" box="[1139,1272,356,380]" class="Actinopterygii" family="Cichlidae" genus="Pseudocrenilabrus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="186" phylum="Chordata" rank="species" species="philander">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFBB47408FB07FED2" box="[1139,1272,356,380]" italics="true" pageId="5" pageNumber="186">P. philander</emphasis>
|
||
</taxonomicName>
|
||
(n = 21). The highlighted results indicate the characters with the highest loadings on PC2 and PC3. These characters are eye diameter, interorbital width, head width, anal fin base, caudal peduncle length and caudal peduncle depth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5874B4FB2CFE41" blockId="5.[151,1436,324,1350]" box="[159,1235,473,495]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF5874B4FB2CFE41" bold="true" box="[159,1235,473,495]" pageId="5" pageNumber="186">Variable PC1 PC2 PC3</emphasis>
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587769FB19FDB4" blockId="5.[151,1436,324,1350]" box="[159,1254,516,538]" pageId="5" pageNumber="186">Lachrymal depth 0.851 0.142 -0.207</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587744FB19FD91" blockId="5.[151,1436,324,1350]" box="[159,1254,553,575]" pageId="5" pageNumber="186">Snout length 0.893 0.241 -0.044</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587720FB22FDCC" blockId="5.[151,1436,324,1350]" box="[159,1245,588,610]" pageId="5" pageNumber="186">Lower jaw length 0.872 -0.250 0.249</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF58771CFB22FD29" blockId="5.[151,1436,324,1350]" box="[159,1245,625,647]" pageId="5" pageNumber="186">Premaxillary pedicel length 0.958 0.025 0.056</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5877F9FB22FD04" blockId="5.[151,1436,324,1350]" box="[159,1245,660,682]" pageId="5" pageNumber="186">Cheek depth 0.934 -0.034 0.061</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5877D4FB19FD61" blockId="5.[151,1436,324,1350]" box="[159,1254,697,719]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF5877D4FED0FD61" bold="true" box="[159,303,697,719]" pageId="5" pageNumber="186">Eye diameter</emphasis>
|
||
0.613
|
||
<emphasis id="9ECEEAF3FFBBFF9AFC6277D4FB19FD61" bold="true" box="[933,1254,697,719]" pageId="5" pageNumber="186">-0.465 -0.596</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E4A0656AFFBBFF9AFF5877B1FB22FAE8" pageId="5" pageNumber="186" type="description">
|
||
<paragraph id="AC0536E1FFBBFF9AFF5877B1FB22FD5C" blockId="5.[151,1436,324,1350]" box="[159,1245,732,754]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF5877B1FE9DFD5C" bold="true" box="[159,354,732,754]" pageId="5" pageNumber="186">Interorbital width</emphasis>
|
||
0.810
|
||
<emphasis id="9ECEEAF3FFBBFF9AFC6277B1FC21FD5C" bold="true" box="[933,990,732,754]" pageId="5" pageNumber="186">0.485</emphasis>
|
||
0.043
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587664FB19FCB1" blockId="5.[151,1436,324,1350]" box="[159,1254,777,799]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF587664FEE2FCB1" bold="true" box="[159,285,777,799]" pageId="5" pageNumber="186">Head width</emphasis>
|
||
0.905
|
||
<emphasis id="9ECEEAF3FFBBFF9AFC627664FC21FCB1" bold="true" box="[933,990,777,799]" pageId="5" pageNumber="186">0.310</emphasis>
|
||
-0.007
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587658FB19FCE5" blockId="5.[151,1436,324,1350]" box="[159,1254,821,843]" pageId="5" pageNumber="186">Head length 0.983 0.023 -0.030</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF58760EFB22FCD7" blockId="5.[151,1436,324,1350]" box="[159,1245,867,889]" pageId="5" pageNumber="186">Standard length 0.978 -0.124 0.062</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5876FCFB22FC09" blockId="5.[151,1436,324,1350]" box="[159,1245,913,935]" pageId="5" pageNumber="186">Body depth 0.952 0.006 0.106</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5876D0FB23FC7D" blockId="5.[151,1436,324,1350]" box="[159,1244,957,979]" pageId="5" pageNumber="186">Dorsal fin base length 0.964 -0.081 0.118</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587684FB22FC51" blockId="5.[151,1436,324,1350]" box="[159,1245,1001,1023]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF587684FE88FC51" bold="true" box="[159,375,1001,1023]" pageId="5" pageNumber="186">Anal fin base length</emphasis>
|
||
0.824
|
||
<emphasis id="9ECEEAF3FFBBFF9AFC627684FB22FC51" bold="true" box="[933,1245,1001,1023]" pageId="5" pageNumber="186">-0.316 0.323</emphasis>
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587163FB19FB8A" blockId="5.[151,1436,324,1350]" box="[159,1254,1038,1060]" pageId="5" pageNumber="186">Predorsal distance 0.972 0.040 -0.064</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF58715FFB22FBE6" blockId="5.[151,1436,324,1350]" box="[159,1245,1074,1096]" pageId="5" pageNumber="186">Prepelvic distance 0.933 0.024 0.012</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF58713BFB19FBC2" blockId="5.[151,1436,324,1350]" box="[159,1254,1110,1132]" pageId="5" pageNumber="186">Preventral distance 0.976 -0.023 -0.064</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5871EDFB19FB38" blockId="5.[151,1436,324,1350]" box="[159,1254,1152,1174]" pageId="5" pageNumber="186">Pre-anal distance 0.972 0.044 -0.057</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5871C1FB22FB6C" blockId="5.[151,1436,324,1350]" box="[159,1245,1196,1218]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF5871C1FE60FB6C" bold="true" box="[159,415,1196,1218]" pageId="5" pageNumber="186">Caudal peduncle length</emphasis>
|
||
0.685
|
||
<emphasis id="9ECEEAF3FFBBFF9AFC6271C1FC18FB6C" bold="true" box="[933,999,1196,1218]" pageId="5" pageNumber="186">-0.579</emphasis>
|
||
0.041
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF5871B5FB19FB40" blockId="5.[151,1436,324,1350]" box="[159,1254,1240,1262]" pageId="5" pageNumber="186">
|
||
<emphasis id="9ECEEAF3FFBBFF9AFF5871B5FE66FB40" bold="true" box="[159,409,1240,1262]" pageId="5" pageNumber="186">Caudal peduncle depth</emphasis>
|
||
0.888
|
||
<emphasis id="9ECEEAF3FFBBFF9AFC6271B5FC21FB40" bold="true" box="[933,990,1240,1262]" pageId="5" pageNumber="186">0.280</emphasis>
|
||
-0.166
|
||
</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF587069FB22FAB4" blockId="5.[151,1436,324,1350]" box="[159,1245,1284,1306]" pageId="5" pageNumber="186">Explained variance 15.333 1.231 0.646</paragraph>
|
||
<paragraph id="AC0536E1FFBBFF9AFF58705DFB22FAE8" blockId="5.[151,1436,324,1350]" box="[159,1245,1328,1350]" pageId="5" pageNumber="186">Proportion of total variance 0.807 0.065 0.034</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |