245 lines
38 KiB
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245 lines
38 KiB
XML
<document ID-CLB-Dataset="129873" ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-PMC="PMC9848625" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-PubMed="36762355" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="210059B2AF7151B8821080CDE3E8D9AD" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupinella illustris Jirapatrasilp & Sutcharit & Panha 2022, comb. nov." docType="treatment" docVersion="6" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1732923707409" updateUser="ExternalLinkService">
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<mods:titleInfo id="B7B09D0A1C1FD2389866C016BBB4733F">
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<mods:title id="C5D2B78D313B9C0A4B89AE4B277B32CA">Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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<mods:name id="68F8F4C5D5ADB3C938B8DC82C33CB455" type="personal">
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<mods:namePart id="F5F2CA42165B423AB93A1E717D949E85">Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier id="880B4FD2AF81F3FFA136D7F9E247B2A2" type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation id="7698D95AAC53DBDEED1FED7E27020975">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="B2A7C3E8F54BEDEA600CCAD791201616" type="personal">
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<mods:namePart id="2705F44785CBC989F92FE28D4FB2A269">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="BE06F0C10D1BBF40380E4BC3BDEE3E7A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="2E99AE32F17A58496DB900E610CE0D87" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="A820EAF68051A51A401B550C707BDBE1">Panha, Somsak</mods:namePart>
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<mods:affiliation id="5A09FEC9F4778C1724FBA9616A2AE87D">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier id="21A2304147589230A86A3891BC23B657" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:typeOfResource id="7833BDC62E07D6220F9ECD0190B798B7">text</mods:typeOfResource>
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<mods:title id="6A33C4BE0D96000AFC3CCC71912989FA">ZooKeys</mods:title>
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<mods:part id="3EDB1C200ACA809A20D17FC736B28BDB">
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<mods:date id="7075138EF0288D2598AAB0A7E4AF73CB">2022</mods:date>
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<mods:number id="AB2A89CCA22B170C89E3E2C8C8A3F043">2022-08-25</mods:number>
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<mods:detail id="7E4608921238602939C069C245464AA6" type="volume">
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<mods:number id="A611ED3ECEF9C559D36EAE3603DBE9E0">1119</mods:number>
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<mods:start id="1D1B7752A7903279F9F7ED6801C7AC50">1</mods:start>
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<mods:end id="4368E5F6999897D12B07BE2FD7949771">115</mods:end>
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<mods:url id="18FDB8D5E3AAC156ED567C4341845B19">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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<mods:classification id="B97C3CFD20D96D14FF027E9DF5C82860">journal article</mods:classification>
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<mods:identifier id="A42F5139F2A64ED16E7A5F1C0E055FBC" type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier id="02D1155E601A8BFC104B516D2487E8AD" type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier id="26B268E7AA22C93D4570B04AF2AE955E" type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<treatment id="210059B2AF7151B8821080CDE3E8D9AD" ID-GBIF-Taxon="199839070" LSID="urn:lsid:plazi:treatment:210059B2AF7151B8821080CDE3E8D9AD" httpUri="http://treatment.plazi.org/id/210059B2AF7151B8821080CDE3E8D9AD" lastPageNumber="1" pageId="0" pageNumber="1" scope_class="Gastropoda" scope_family="Pupinidae" scope_phylum="Mollusca" scope_subClass="Caenogastropoda" scope_subFamily="Pupinellinae" scope_superFamily="Cyclophoroidea">
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<subSubSection id="3AC6F1282B02053DCBE80784C08D90FE" pageId="0" pageNumber="1" type="nomenclature">
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<paragraph id="09B0BE4D295104DDB60A767FE766FE96" pageId="0" pageNumber="1">
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<taxonomicName id="9FDA4E94DAE9FE7FA350FE84A23034CE" ID-CoL="9GG74" LSID="210059B2-AF71-51B8-8210-80CDE3E8D9AD" authority="(Mabille, 1887)" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris" status="comb. nov.">Pupinella illustris (Mabille, 1887)</taxonomicName>
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<taxonomicNameLabel id="6A48879850235C75C063C2B190BD4BA0" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="CA941408DDDB2BDA7BD71D6DFD81581B" pageId="0" pageNumber="1" type="description">
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<paragraph id="69EEE19F21C4E8866C303333B4D3C053" pageId="0" pageNumber="1">
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<figureCitation id="158093E7A974B6CC789494BAC8D6FBA0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">Figs 11I-L</figureCitation>
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<figureCitation id="8B7068674FA9E23CF9A10F312175C0EA" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">, 12D, E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="85C91D677366C08E1BA14C9EBDAC2745" pageId="0" pageNumber="1" type="reference_group">
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<paragraph id="77B2FEFDE2C7F391D4560A8210F6A948" pageId="0" pageNumber="1">
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<taxonomicName id="AD8E08333D99AD73802E27085EF30D97" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris" status="comb. nov.">Pupina illustris</taxonomicName>
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Mabille, 1887: 136, 137. Type locality: Tonkin.
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<bibRefCitation id="4084BDBACB7E8ADCCE335F2F8514CE1A" DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Imprimerie Dejussieu Pere et Fils, Autun" pageId="0" pageNumber="1" refId="B41" refString="Fischer, P, 1891. Catalogue et distribution geographique des Mollusques terrestres, fluviatilies & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des Mollusques terrestres, fluviatilies & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" year="1891">Fischer 1891</bibRefCitation>
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: 107.
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<bibRefCitation id="6F8B9C8509A3468A4DBD327E836FD1CF" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 431.
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</paragraph>
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<paragraph id="871ED7A02164C41DD229B6155BAE96A6" pageId="0" pageNumber="1">
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<taxonomicName id="0D15445421C449E4C14A150920084AED" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana" status="comb. nov.">Pupina tonkiniana</taxonomicName>
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Bavay & Dautzenberg, 1899: 54, 55, pl. 3, fig. 6, 6a (as
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<taxonomicName id="F01BA24E189ACB206F0A0E6F315EB6BF" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina tonkiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiana" status="comb. nov.">Pupina tonkiana</taxonomicName>
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in the original description). Type locality: Entre Lang-Son [Lang Son Province, Vietnam] et
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<normalizedToken id="181D6087F50AA51A2663C68705A60F15" originalValue="That-Khé">That-Khe</normalizedToken>
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[That Ke, Lang Son Province, Vietnam]. Syn. nov.
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</paragraph>
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<paragraph id="F108FE96C4333D3C2DB346639694D7F6" pageId="0" pageNumber="1">
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<taxonomicName id="E92CF99F3D996E914A1D1DD7605AE301" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris" status="comb. nov." subGenus="Tylotoechus">Pupina (Tylotoechus) illustris</taxonomicName>
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-
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<bibRefCitation id="FC01A05BF05753A437DE9E3060CCF3E0" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 314, 315.
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</paragraph>
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<paragraph id="628C0B5520BE4AB0DAC020320376C18B" pageId="0" pageNumber="1">
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<taxonomicName id="C8EDBBD8B2557ABA6DBDB8EFDD770475" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana" status="comb. nov." subGenus="Tylotoechus">Pupina (Tylotoechus) tonkiniana</taxonomicName>
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-
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<bibRefCitation id="2CD8F5E01148CF86236E00600FBFE457" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 323, 324.
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<bibRefCitation id="1AFB5A0303AA58ADF9B2986754AB6707" author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 56" refId="B198" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae. Archiv fuer Molluskenkunde 86: 41 - 56" title="Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae." volume="86" year="1957">Zilch 1957</bibRefCitation>
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: 48.
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</paragraph>
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<paragraph id="501DDF606099B63686E642466DA88313" pageId="0" pageNumber="1">
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<taxonomicName id="FA07F2F4BE2A0D83971D501C7B5F61D2" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana" status="comb. nov.">Pupina tonkiniana</taxonomicName>
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-
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<bibRefCitation id="8FD6037F83155FDA483F5C3DFC871D7C" author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 432.
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<bibRefCitation id="5012AA67ADEE44DDA0802155497B810D" author="Fischer-Piette, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="149 - 180" refId="B47" refString="Fischer-Piette, E, 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180" title="Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette 1950</bibRefCitation>
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: 167.
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<bibRefCitation id="89820A757DE5D49ECFD5C240FAD96F24" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
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: 126, fig. 6b, Son La Province, Vietnam.
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<bibRefCitation id="C4AD5F2EBDFEABE96C60AB09F24FD961" author="Raheem, DC" journalOrPublisher="The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" pageId="0" pageNumber="1" refId="B137" refString="Raheem, DC, Backeljau, T, Pearce-Kelly, P, Taylor, H, Fenn, J, Sutcharit, C, Panha, S, von, Oheimb KCM, von, Oheimb PV, Ikebe, C, Pall-Gergely, B, Gargominy, O, Hao, LV, Sang, PV, Tu, DV, Phong, DT, Naggs, M, Ablett, J, Dodds, JM, Wade, CM, Naggs, F, 2017. An Illustrated Guide to the Land Snails and Slugs of Vietnam. The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" title="An Illustrated Guide to the Land Snails and Slugs of Vietnam." year="2017">Raheem et al. 2017</bibRefCitation>
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: 5 (plate figure).
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</paragraph>
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</subSubSection>
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<subSubSection id="87AC8519BDD69A6EA11D4C3E179FB7F1" pageId="0" pageNumber="1" type="type material">
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<paragraph id="02AAAEA1B09A281DA8A9D1DFD93F2470" pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph id="3311967B4A20FCCD5EE62C6CB2EC308F" pageId="0" pageNumber="1">
|
|
<emphasis id="6C1CF0C8C8805BA2C9D1CE5942FB0008" bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
|
|
of
|
|
<taxonomicName id="E363A3D97D56DDC1431F4407E24A0E3D" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
|
|
<emphasis id="102C322C2375AFB305EA709C9104AEE7" italics="true" pageId="0" pageNumber="1">Pupina illustris</emphasis>
|
|
</taxonomicName>
|
|
MNHN-IM-2000-35842 (9 shells; Figs
|
|
<figureCitation id="2103ABEC2DBDEE25EB2A24B6B7E1ED7D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11I, J</figureCitation>
|
|
,
|
|
<figureCitation id="660F3EE64938B04D66B8529AFE1CE1BE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">12D</figureCitation>
|
|
) from Tonkin.
|
|
<emphasis id="D2F46F8540F60B74DDEFE1EC2294D4C5" bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
|
|
of
|
|
<taxonomicName id="C1DAB62D74BE455543282EB4386541E7" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="E7211A14ADABF95105036DB7E95912BF" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
MNHN-IM-2000-35838 (Fig.
|
|
<figureCitation id="C27BA61572829CB2ABD7D496A6000B21" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11K</figureCitation>
|
|
) from Lang-Son et
|
|
<normalizedToken id="DF0A13AD6065342E4F704F4AA5EFF2EA" originalValue="That-Khé">That-Khe</normalizedToken>
|
|
. Paralectotypes of
|
|
<taxonomicName id="780AA0998069BA7A7A3A56BD24CB249F" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="32730A9B0D76FE547C803D43BBF14172" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
SMF 109932/10 (10 shells; Figs
|
|
<figureCitation id="872960B385C62ED831C583E1253826A0" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11L</figureCitation>
|
|
,
|
|
<figureCitation id="AA3A061CA7A1521C1897B0E786CDA0A6" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">12E</figureCitation>
|
|
) from Tonkin:
|
|
<normalizedToken id="73C00542EB9276389082F27CB3E0C45C" originalValue="That-khé">That-khe</normalizedToken>
|
|
. Paralectotypes of
|
|
<taxonomicName id="5C9A1455345812899C9A8C6D6B8A31DA" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="7220C1E732B74E38165BD8CF1893EBCC" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
RBINS MT976/2 (14 shells) from Lang Son et
|
|
<normalizedToken id="62B018D4FADD6BCBC346557A21873A73" originalValue="That-khé">That-khe</normalizedToken>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="74139871820E53CDB236B1FF141A5039" pageId="0" pageNumber="1" type="diagnosis">
|
|
<paragraph id="CF9B3335C55D4A4CDB3186249AF1B003" pageId="0" pageNumber="1">Diagnosis.</paragraph>
|
|
<paragraph id="F7B9040587DD271AC803D80DD3CDD95F" pageId="0" pageNumber="1">Shell elongate fusiform; last whorl ca. 55-60% of shell height. Apertural lip expanded and slightly reflected; apertural lip when observed from lateral view almost straight. Parietal callus absent. Parietal tooth pointily sharp, located next to wide posterior canal. Anterior canal funnel-like. Umbilicus closed.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="4BB6B6702CF12883B2BDF85814356BEA" pageId="0" pageNumber="1" type="differential diagnosis">
|
|
<paragraph id="A8D714FA6A1BADC3F57F815DF55C92CC" pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
|
|
<paragraph id="EC45D816526CC85F074B685E8EE2B8B6" pageId="0" pageNumber="1">
|
|
<taxonomicName id="91E1D5442758260B5C7767F79D12C538" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
|
|
<emphasis id="6EE854F4F55D7BF206F2A71720B747CC" italics="true" pageId="0" pageNumber="1">Pupinella illustris</emphasis>
|
|
</taxonomicName>
|
|
can be distinguished from all other species in mainland Southeast Asia by an elongate fusiform shell shape, an absence of parietal callus and a pointily sharp parietal tooth located next to a wide posterior canal.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="0156985B417C0486116BCCD2FF691BD8" pageId="0" pageNumber="1" type="distribution">
|
|
<paragraph id="604342F267209EA11CB9175F841187E2" pageId="0" pageNumber="1">Distribution.</paragraph>
|
|
<paragraph id="EF588D0238EB69B1C7FA92E51ECCCC47" pageId="0" pageNumber="1">
|
|
Northern Vietnam (
|
|
<bibRefCitation id="B4FC2B36C6EC091540F788081A1D1596" author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="BC6569988447E06AE17AD80DB1816AE8" author="Raheem, DC" journalOrPublisher="The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" pageId="0" pageNumber="1" refId="B137" refString="Raheem, DC, Backeljau, T, Pearce-Kelly, P, Taylor, H, Fenn, J, Sutcharit, C, Panha, S, von, Oheimb KCM, von, Oheimb PV, Ikebe, C, Pall-Gergely, B, Gargominy, O, Hao, LV, Sang, PV, Tu, DV, Phong, DT, Naggs, M, Ablett, J, Dodds, JM, Wade, CM, Naggs, F, 2017. An Illustrated Guide to the Land Snails and Slugs of Vietnam. The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" title="An Illustrated Guide to the Land Snails and Slugs of Vietnam." year="2017">Raheem et al. 2017</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="5F4AA442F5C20FC6171AA00352C6F712" pageId="0" pageNumber="1" type="remarks">
|
|
<paragraph id="91096B0B30DA7EB854DC3EC3B9F0EF62" pageId="0" pageNumber="1">Remarks.</paragraph>
|
|
<paragraph id="2E65CBE1C233A62ACB9A0284E114860E" pageId="0" pageNumber="1">
|
|
This taxon is allocated to the genus
|
|
<taxonomicName id="065A22B2CF6B0465781DBEAE092FB597" authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
|
|
<emphasis id="C27443CAE1620759A73CCB30C0CA38B7" italics="true" pageId="0" pageNumber="1">Pupinella</emphasis>
|
|
</taxonomicName>
|
|
due to the presence of a funnel-like anterior canal, which is the diagnostic character of this genus. In the original description of
|
|
<taxonomicName id="6C9DFA05A15589AE0C36A5902324FDF2" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="4020CE49D84369FA5D19E0C1A044996C" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
, two ways of spelling were shown: the spelling '
|
|
<taxonomicName id="D16A3082B7D439EA2BB1FDC29B6D27E2" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiana">
|
|
<emphasis id="4AA3B7A3A669E509AF8E75CB84D0776A" italics="true" pageId="0" pageNumber="1">Pupina tonkiana</emphasis>
|
|
</taxonomicName>
|
|
' in the description, and '
|
|
<taxonomicName id="309356D2370A3F37C0132CE7D7701106" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="9C2C76D78A5C830D20E04FD682936C7E" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
' in the plate caption. Later,
|
|
<bibRefCitation id="0C3733732B4BE02A4BE3C44723C91138" author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt (1902)</bibRefCitation>
|
|
acted as the First Reviser (
|
|
<bibRefCitation id="C3B5A3102FE110E5053B8B06CAC8F2BB" author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
|
|
: Art. 24.2.3) in selecting '
|
|
<taxonomicName id="9FD040B538F3753A220FC4E59560461B" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="24CFF761D07EDFF07F0A5E355E16C1E4" italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
' as the correct original spelling. As the original description did not explicitly state that the description of
|
|
<taxonomicName id="0E8A4FD922213A1FFDB954E98BFCE3C8" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="6EAA39E8641D603FEF9B359FB98B3057" italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
was based on a single specimen (nor could this be inferred), the designation of a holotype by
|
|
<bibRefCitation id="4BA4346600AE6FAB4D1EB3D932EBDED8" author="Fischer-Piette, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="149 - 180" refId="B47" refString="Fischer-Piette, E, 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180" title="Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette (1950)</bibRefCitation>
|
|
in fact constitutes a lectotype designation (
|
|
<bibRefCitation id="6B3BE5C6AAB8BDE37E6C7DD0A5016A7A" author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
|
|
: Art. 74.6).
|
|
</paragraph>
|
|
<paragraph id="D602CCC6FB815C9CA00A9BA547187F86" pageId="0" pageNumber="1">
|
|
Upon examining the type specimens of both
|
|
<taxonomicName id="572A45CC4C598C3BDB70F10067D3BE68" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
|
|
<emphasis id="C4DDE193DCB2C987436AED68AD90C5FF" italics="true" pageId="0" pageNumber="1">P. illustris</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="7F6E5154104CB373D82F911E519AA2C0" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="8F79F52CCE6113F349FE587F3CE9B1DF" italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
, the type series of
|
|
<taxonomicName id="0C6052EDD940B911B74EEDAF2483032C" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="7A107D22F3AD4A8E32FF7F1D790EE320" italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
agree well with all the syntypes of
|
|
<taxonomicName id="ABAA14B1A50426CFB4B5703E0BE1D472" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
|
|
<emphasis id="D0E729E294155F4D6C7B5D4FC5C4168E" italics="true" pageId="0" pageNumber="1">P. illustris</emphasis>
|
|
</taxonomicName>
|
|
in having an elongate fusiform shell shape, an expanded and slightly reflected apertural lip without a parietal callus, and a sharp, tooth-like, parietal tooth located next to a wide posterior canal. Thus,
|
|
<taxonomicName id="01A785581D0B252C3AD69DF24343FBAF" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
|
|
<emphasis id="9B406CC503DEF0DBE26C703FF509BF40" italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
is regarded herein as a junior subjective synonym of
|
|
<taxonomicName id="3F602FA6DDEFED239E4A541F0CA0DA41" class="Gastropoda" family="Pupinidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="P. illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
|
|
<emphasis id="94127B3BEC608E1D89167A8A8A01C037" italics="true" pageId="0" pageNumber="1">P. illustris</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |