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<document id="5F2C58A4BB77F49D5039422DB615199E" ID-DOI="10.18590/euscorpius.2004.vol2004.iss15.1" ID-ISSN="1536-9307" ID-Zenodo-Dep="12781607" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721358433569" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2004" docId="1E6687C0FFF0FF934D74FC23A49A194F" docLanguage="en" docName="Euscorpius.15.1-60.pdf" docOrigin="Euscorpius 15 (15)" docSource="https://mds.marshall.edu/euscorpius/vol2004/iss15/1/" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Heterometrus mysorensis Kovařík, 2004, sp. n." docType="treatment" docVersion="2" lastPageNumber="29" masterDocId="E25FFFB8FFEEFF8D4C47FF8BA0111F63" masterDocTitle="A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae)" masterLastPageNumber="60" masterPageNumber="1" pageNumber="29" updateTime="1721381062215" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="A0A42067FECA464B5ED6F0ACF2D0FD60">A review of the genus Heterometrus Ehrenberg, 1828, with descriptions of seven new species (Scorpiones, Scorpionidae)</mods:title>
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<mods:namePart id="7F5BB1D4F7F5B62DEB5191A8DDE1BB06">Kovařík, František</mods:namePart>
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<mods:title id="2AEF421074EAC008483A13062C91B8D2">Euscorpius</mods:title>
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<mods:date id="3D8306DB2F40BE0E0A6E9A51B34947D2">2004</mods:date>
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<mods:number id="DFAB4AE68FBE0CF89F6698C902B52982">2004-12-31</mods:number>
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<mods:classification id="4B8C0B0B3A65C06C52C3F45FE34D0F67">journal article</mods:classification>
<mods:identifier id="C0D9A6F15B24DC1E806DC1D052878624" type="DOI">10.18590/euscorpius.2004.vol2004.iss15.1</mods:identifier>
<mods:identifier id="051104B670313F9F80433834288E1AEE" type="ISSN">1536-9307</mods:identifier>
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<treatment id="1E6687C0FFF0FF934D74FC23A49A194F" ID-DOI="http://doi.org/10.5281/zenodo.12781437" ID-Zenodo-Dep="12781437" LSID="urn:lsid:plazi:treatment:1E6687C0FFF0FF934D74FC23A49A194F" httpUri="http://treatment.plazi.org/id/1E6687C0FFF0FF934D74FC23A49A194F" lastPageNumber="29" pageId="30" pageNumber="29">
<subSubSection id="DED5655DFFF0FF934D74FC23A2BE1CA1" box="[307,687,936,962]" pageId="30" pageNumber="29" type="nomenclature">
<paragraph id="967036D6FFF0FF934D74FC23A2BE1CA1" blockId="30.[307,687,936,995]" box="[307,687,936,962]" pageId="30" pageNumber="29">
<heading id="CD3881BAFFF0FF934D74FC23A2BE1CA1" box="[307,687,936,962]" centered="true" fontSize="10" level="2" pageId="30" pageNumber="29" reason="4">
<emphasis id="A4BBEAC4FFF0FF934D74FC23A2BE1CA1" bold="true" box="[307,687,936,962]" pageId="30" pageNumber="29">
<taxonomicName id="51CF4D55FFF0FF934D74FC23A2771CA1" authorityName="Kovařík" authorityYear="2004" box="[307,614,936,962]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="mysorensis" status="sp. nov.">
<emphasis id="A4BBEAC4FFF0FF934D74FC23A2771CA1" bold="true" box="[307,614,936,962]" italics="true" pageId="30" pageNumber="29">Heterometrus mysorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8857BFFFF0FF934E2AFC23A2BE1CA1" box="[621,687,936,962]" pageId="30" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DED5655DFFF0FF934DFAFC42A2351C80" box="[445,548,969,995]" pageId="30" pageNumber="29" type="description">
<paragraph id="967036D6FFF0FF934DFAFC42A2351C80" blockId="30.[307,687,936,995]" box="[445,548,969,995]" pageId="30" pageNumber="29">
(
<figureCitation id="0EF42A53FFF0FF934D81FC42A20A1C80" box="[454,539,969,995]" captionStart="Figure 21" captionStartId="31.[192,268,1852,1875]" captionTargetBox="[268,1387,226,1788]" captionTargetId="figure-17@31.[273,1383,230,1783]" captionTargetPageId="31" captionText="Figure 21: Heterometrus mysorensis, sp. nov., dorsal view, male holotype." figureDoi="http://doi.org/10.5281/zenodo.12781651" httpUri="https://zenodo.org/record/12781651/files/figure.png" pageId="30" pageNumber="29">Fig. 21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DED5655DFFF0FF934C87FB82A2F91B1F" pageId="30" pageNumber="29" type="materials_examined">
<paragraph id="967036D6FFF0FF934C87FB82A20D1B5C" blockId="30.[192,802,1033,1148]" pageId="30" pageNumber="29">
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<typeStatus id="49748874FFF0FF934C87FB82A0E91B43" box="[192,248,1033,1056]" pageId="30" pageNumber="29">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934D45FB86A17C1B43" baselines="1051" box="[258,365,1037,1056]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="locality" pageId="30" pageNumber="29">LOCALITY</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934D31FB86A1B51B43" baselines="1051" box="[374,420,1037,1056]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="30" pageNumber="29">AND</smallCapsWord>
<materialsCitation id="26A73C8BFFF0FF934DEAFB86A1D61B5C" country="India" elevation="900" location="Mysore" municipality="Maddur" pageId="30" pageNumber="29" specimenCount="1" typeStatus="holotype">
<smallCapsWord id="9096A00AFFF0FF934DEAFB86A1F31B43" baselines="1051" box="[429,482,1037,1056]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="type" pageId="30" pageNumber="29">
<typeStatus id="49748874FFF0FF934DEAFB86A1F31B43" box="[429,482,1037,1056]" pageId="30" pageNumber="29">TYPE</typeStatus>
</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934DABFB86A2611B43" baselines="1051" box="[492,624,1037,1056]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="repository" pageId="30" pageNumber="29">REPOSITORY</smallCapsWord>
.
<collectingCountry id="EED87646FFF0FF934EC6FB82A2AA1B43" box="[641,699,1033,1056]" name="India" pageId="30" pageNumber="29">India</collectingCountry>
,
<location id="9310600DFFF0FF934E8EFB82A30C1B43" LSID="urn:lsid:plazi:treatment:1E6687C0FFF0FF934D74FC23A49A194F:9310600DFFF0FF934E8EFB82A30C1B43" box="[713,797,1033,1056]" country="India" municipality="Maddur" name="Mysore" pageId="30" pageNumber="29">Mysore</location>
,
<collectingMunicipality id="7614ACACFFF0FF934C87FBA3A1041B5C" box="[192,277,1064,1087]" pageId="30" pageNumber="29">Maddur</collectingMunicipality>
env., cca
<quantity id="51379B33FFF0FF934DC7FBA3A1D61B5C" box="[384,455,1064,1087]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="29" unit="m" value="900.0">
<elevation id="1DE2D1E5FFF0FF934DC7FBA3A1D61B5C" box="[384,455,1064,1087]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="29" unit="m" value="900.0">900 m</elevation>
</quantity>
</materialsCitation>
;
<collectionCode id="F0DEAE13FFF0FF934D96FBA3A2061B5C" box="[465,535,1064,1087]" pageId="30" pageNumber="29">FKCP</collectionCode>
.
</paragraph>
<paragraph id="967036D6FFF0FF934C87FBCDA2F91B1F" blockId="30.[192,802,1033,1148]" pageId="30" pageNumber="29">
<materialsCitation id="26A73C8BFFF0FF934C87FBCDA2F51B1F" collectionCode="FKCP" country="India" elevation="900" location="Mysore" municipality="Maddur" pageId="30" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<smallCapsWord id="9096A00AFFF0FF934C87FBCDA0E91B3E" baselines="1111,1112" box="[192,248,1094,1117]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Type" pageId="30" pageNumber="29">TYPE</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934D45FBC1A1611B3E" baselines="1112" box="[258,368,1098,1117]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="material" pageId="30" pageNumber="29">MATERIAL</smallCapsWord>
.
<emphasis id="A4BBEAC4FFF0FF934DC6FBCDA1AF1B3E" bold="true" box="[385,446,1094,1117]" pageId="30" pageNumber="29">
<collectingCountry id="EED87646FFF0FF934DC6FBCDA1AF1B3E" box="[385,446,1094,1117]" name="India" pageId="30" pageNumber="29">India</collectingCountry>
</emphasis>
,
<emphasis id="A4BBEAC4FFF0FF934D97FBCDA2301B3E" bold="true" box="[464,545,1094,1117]" italics="true" pageId="30" pageNumber="29">
<location id="9310600DFFF0FF934D97FBCDA2301B3E" LSID="urn:lsid:plazi:treatment:1E6687C0FFF0FF934D74FC23A49A194F:9310600DFFF0FF934D97FBCDA2301B3E" box="[464,545,1094,1117]" country="India" municipality="Maddur" name="Mysore" pageId="30" pageNumber="29">Mysore</location>
</emphasis>
,
<collectingMunicipality id="7614ACACFFF0FF934E74FBCDA2961B3E" box="[563,647,1094,1117]" pageId="30" pageNumber="29">Maddur</collectingMunicipality>
env., ca.
<quantity id="51379B33FFF0FF934EBEFBCDA0C61B1F" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="29" unit="m" value="900.0">
<elevation id="1DE2D1E5FFF0FF934EBEFBCDA0C61B1F" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="30" pageNumber="29" unit="m" value="900.0">900 m</elevation>
</quantity>
, 1983,
<specimenCount id="80C9FD5FFFF0FF934D62FBEEA1571B1F" box="[293,326,1125,1148]" count="1" pageId="30" pageNumber="29" type="male">1♂</specimenCount>
(
<typeStatus id="49748874FFF0FF934D13FBEEA1A61B1F" box="[340,439,1125,1148]" pageId="30" pageNumber="29" type="holotype">holotype</typeStatus>
), collector unknown,
<collectionCode id="F0DEAE13FFF0FF934ED9FBEEA2F51B1F" box="[670,740,1125,1148]" pageId="30" pageNumber="29">FKCP</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DED5655DFFF0FF934C87FB29A2CA1BDA" box="[192,731,1186,1209]" pageId="30" pageNumber="29" type="etymology">
<paragraph id="967036D6FFF0FF934C87FB29A2CA1BDA" blockId="30.[192,731,1186,1209]" box="[192,731,1186,1209]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934C87FB29A15B1BDA" baselines="1203,1204" box="[192,330,1186,1209]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Etymology" pageId="30" pageNumber="29">ETYMOLOGY</smallCapsWord>
. Named after the state of occurrence.
</paragraph>
</subSubSection>
<subSubSection id="DED5655DFFF0FF934C87FB56A1C91926" pageId="30" pageNumber="29" type="diagnosis">
<paragraph id="967036D6FFF0FF934C87FB56A1C91926" blockId="30.[192,809,1245,1605]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934C87FB56A1271B90" baselines="1262,1262" box="[192,310,1245,1268]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Diagnosis" pageId="30" pageNumber="29">DIAGNOSIS</smallCapsWord>
. Adult male
<quantity id="51379B33FFF0FF934D80FB56A2311B97" box="[455,544,1245,1268]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="30" pageNumber="29" unit="mm" value="118.0">118 mm</quantity>
long. Female unknown. Color uniformly reddish brown, only telson and leg tarsomeres reddish yellow. Pectinal teeth number 16 and 17. Male assumed to have femur, patella and chela of pedipalp longer than female. Chela slightly lobed and strongly hirsute, its length to width ratio 2.8. Dorsal surface of pedipalp manus bears rounded granules of varying size that form elevated rows. Patella of pedipalp without conspicuous internal tubercle. Disc of carapace smooth and glossy, margins finely granulose and anteriorly slightly tuberculate. Telson hirsute, with vesicle as long as aculeus.
</paragraph>
</subSubSection>
<subSubSection id="DED5655DFFF0FF934C87F9E3A49A194F" pageId="30" pageNumber="29" type="description">
<paragraph id="967036D6FFF0FF934C87F9E3A1A21817" blockId="30.[192,802,1640,1908]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934C87F9E3A15F191C" baselines="1657,1658" box="[192,334,1640,1663]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Description" pageId="30" pageNumber="29">DESCRIPTION</smallCapsWord>
: The male
<typeStatus id="49748874FFF0FF934D90F9E3A224191C" box="[471,565,1640,1663]" pageId="30" pageNumber="29" type="holotype">holotype</typeStatus>
is an adult
<quantity id="51379B33FFF0FF934E83F9E3A333191C" box="[708,802,1640,1663]" metricMagnitude="-1" metricUnit="m" metricValue="1.18" pageId="30" pageNumber="29" unit="mm" value="118.0">118 mm</quantity>
long. Measurements of the carapace, telson, segments of the metasoma and segments of the pedipalps, and numbers of pectinal teeth are given in
<tableCitation id="DB4D036DFFF0FF934E34F94FA2D819B8" box="[627,713,1732,1755]" captionStart="Table 1" captionStartId="8.[192,257,1076,1099]" captionText="Table 1: Measurements (in milimetres) of type specimens of seven new Heterometrus species." pageId="30" pageNumber="29">Table 1</tableCitation>
. Sexual dimorphism is presumably expressed in the length of the femur, patella and chela of pedipalp, and probably also in the width of the chela. Although the female is not known, morphology of other species makes this assumption fairly safe.
</paragraph>
<paragraph id="967036D6FFF0FF934F79FF4CA39E1E59" blockId="30.[830,1441,199,1326]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934F79FF4CA3DD1FBD" baselines="216,217" box="[830,972,199,222]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Coloration" pageId="30" pageNumber="29">COLORATION</smallCapsWord>
: The color is uniformly reddish brown, only the telson and leg tarsomeres are reddish yellow. Also the sternites are lighter colored, and the pectens are yellow.
</paragraph>
<paragraph id="967036D6FFF0FF934F79FEC9A48D1E91" blockId="30.[830,1441,199,1326]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934F79FEC9A3AD1E3B" baselines="339,339" box="[830,956,322,345]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Mesosoma" pageId="30" pageNumber="29">MESOSOMA</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934F8CFECEA3E81E3B" baselines="339" box="[971,1017,325,344]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="30" pageNumber="29">AND</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF93484FFECEA46B1E3B" baselines="339" box="[1032,1146,325,344]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="carapace" pageId="30" pageNumber="29">CARAPACE</smallCapsWord>
: The mesosoma lacks a carina. Both the mesosoma and carapace are smooth except for minute granules on the margins and low tubercles on the anterior part of the carapace. The sternites are smooth, entirely without granules. The pectinal teeth number 16 and 17.
</paragraph>
<paragraph id="967036D6FFF0FF934F79FE71A3871C01" blockId="30.[830,1441,199,1326]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934F79FE71A3AC1D73" baselines="523,523" box="[830,957,506,529]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Metasoma" pageId="30" pageNumber="29">METASOMA</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934F82FE76A3E21D73" baselines="523" box="[965,1011,509,528]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="and" pageId="30" pageNumber="29">AND</smallCapsWord>
<smallCapsWord id="9096A00AFFF0FF934FBCFE76A45C1D73" baselines="523" box="[1019,1101,509,528]" lowerCaseFontSize="8" mainFontSize="10" normCase="lower" normString="telson" pageId="30" pageNumber="29">TELSON</smallCapsWord>
: The metasoma is smooth and hirsute. The first to fourth segments bear eight carinae, of which the ventral are smooth and the dorsal consist of minute, rounded granules. The fifth segment bears five complete carinae and two additional lateral carinae developed only in the anterior half. The three ventral carinae of this segment consist of pointed granules and the spaces between them are finely granulate. The dorsal carinae consist only of minute, rounded granules. The telson is hirsute, dorsally without granules, and ventrally with four rows of granules. The vesicle is as long as the aculeus.
</paragraph>
<paragraph id="967036D6FFF0FF934F79FCE1A52A1B92" blockId="30.[830,1441,199,1326]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934F79FCE1A3A11CE3" baselines="891,891" box="[830,944,874,897]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Pedipalps" pageId="30" pageNumber="29">PEDIPALPS</smallCapsWord>
: The femur has granulose carinae and its dorsal surface is slightly granulated. The patella is smooth, dorsally and externally slightly tuberculate, with four smooth carinae and on the internal surface with scattered pointed granules substituting for two additional carinae; they do not form a conspicuous internal tubercle otherwise characteristic for the genus. The chela is slightly lobed and strongly hirsute. The dorsal side of the manus bears rounded granules of varying size, which do not cover the entire surface and form elevated rows. The external surface of the chela is smooth, with two long carinae. The movable fingers bear six straight rows of granules without internal and external granules.
</paragraph>
<paragraph id="967036D6FFF0FF934F79FB73A5341A4D" blockId="30.[830,1441,199,1326]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934F79FB73A3661A6C" baselines="1289,1290" box="[830,887,1272,1295]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Legs" pageId="30" pageNumber="29">LEGS</smallCapsWord>
: The legs bear solitary long and short setae. The third and fourth leg spine formula is 5–6:6–7.
</paragraph>
<paragraph id="967036D6FFF0FF934F79FAD6A49A194F" blockId="30.[830,1441,1373,1580]" pageId="30" pageNumber="29">
<smallCapsWord id="9096A00AFFF0FF934F79FAD6A3A51A17" baselines="1390,1391" box="[830,948,1373,1396]" lowerCaseFontSize="8" mainFontSize="10" normCase="title" normString="Affinities" pageId="30" pageNumber="29">AFFINITIES</smallCapsWord>
. The described features distinguish
<taxonomicName id="51CF4D55FFF0FF9349C1FAD6A3A41AF0" authorityName="Kovařík" authorityYear="2004" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="mysorensis" status="sp. n.">
<emphasis id="A4BBEAC4FFF0FF9349C1FAD6A3A41AF0" italics="true" pageId="30" pageNumber="29">H. mysorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8857BFFFF0FF934F86FAF7A3EE1AF0" box="[961,1023,1404,1427]" pageId="30" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
from all other species of the genus. They are recounted in the key below.
<taxonomicName id="51CF4D55FFF0FF9348A7FA10A5681AD1" authorityName="Kovařík" authorityYear="2004" box="[1248,1401,1435,1458]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="mysorensis" status="sp. n.">
<emphasis id="A4BBEAC4FFF0FF9348A7FA10A5681AD1" box="[1248,1401,1435,1458]" italics="true" pageId="30" pageNumber="29">H. mysorensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BF8857BFFFF0FF9349C5FA10A3421AB3" pageId="30" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
is close to
<emphasis id="A4BBEAC4FFF0FF934F91FA32A44D1AB2" box="[982,1116,1465,1489]" italics="true" pageId="30" pageNumber="29">
<taxonomicName id="51CF4D55FFF0FF934F91FA32A4491AB2" baseAuthorityName="Pocock" baseAuthorityYear="1893" box="[982,1112,1465,1489]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="phipsoni">H. phipsoni</taxonomicName>
,
</emphasis>
but has conspicuous rows of granules (carinae) on the chela and only slightly tuberculate patella of pedipalp, whereas in
<taxonomicName id="51CF4D55FFF0FF934967FA7CA5B1196D" baseAuthorityName="Pocock" baseAuthorityYear="1893" box="[1312,1440,1526,1550]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="30" pageNumber="29" phylum="Arthropoda" rank="species" species="phipsoni">
<emphasis id="A4BBEAC4FFF0FF934967FA7CA5B1196D" box="[1312,1440,1526,1550]" italics="true" pageId="30" pageNumber="29">H. phipsoni</emphasis>
</taxonomicName>
the chela is strongly granulose.
</paragraph>
</subSubSection>
</treatment>
</document>
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