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<document id="8B2E001F3F65D280D83F204D34C3FE37" ID-CLB-Dataset="46415" ID-DOI="10.5281/zenodo.204428" ID-GBIF-Dataset="b27a74a0-2042-4720-839a-c5176cb48a38" ID-ISSN="1175-5326" ID-Zenodo-Dep="204428" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460385671311" checkinUser="plazi" docAuthor="Bellis, Glenn &amp; Dyce, Alan" docDate="2011" docId="1D6FB737FF9EFFABFF2FFEC850E3F957" docLanguage="en" docName="zt03014p058.pdf" docOrigin="Zootaxa 3014" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Culicoides zentae Bellis &amp; Dyce, sp. nov." docType="treatment" docVersion="11" lastPageNumber="46" masterDocId="E156CF4FFF97FFA0FFB8FFEA5414FFA8" masterDocTitle="Marksomyia, a new subgenus of Culicoides Latreille (Diptera: Ceratopogonidae) from the Australasian biogeographic region with descriptions of two new species" masterLastPageNumber="38" masterPageNumber="35" pageNumber="44" updateTime="1698297491458" updateUser="ExternalLinkService">
<mods:mods id="5F796C2FD02DBB80E036F9D6658A2C8C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7D5834C15317E414A06C9216732603D4">
<mods:title id="4FC5054CC1A8E121E702DD7FC5949896">Marksomyia, a new subgenus of Culicoides Latreille (Diptera: Ceratopogonidae) from the Australasian biogeographic region with descriptions of two new species</mods:title>
</mods:titleInfo>
<mods:name id="7CED2EE1E8E7AFF66255E3F1BC187083" type="personal">
<mods:role id="8CD28F15AEB96534C38BE3A785B0A6AF">
<mods:roleTerm id="6C57AD69737CC466A21642D74F77F50C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B39E2AFD0FE3B0D7E808AB5B2B4AA3F">Bellis, Glenn</mods:namePart>
</mods:name>
<mods:name id="4CE17E52334074600FEF8ADBFB9FB102" type="personal">
<mods:role id="2527C57C2AB97D123D15F035392C7493">
<mods:roleTerm id="83575887E2E938C4BE8F0EEB055E9BFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F160338DB80BD026DEF5B019CFA4C3E">Dyce, Alan</mods:namePart>
</mods:name>
<mods:typeOfResource id="D4083C94DD78FFF3B1E10F912C92DF6F">text</mods:typeOfResource>
<mods:relatedItem id="FA48A37128CC2B184D2AD0E1303EC5A6" type="host">
<mods:titleInfo id="930F950C02B0CCC11381A9BB628B9219">
<mods:title id="A226932F1F8B1516AC6DA0BAFE630DAE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C5FA8FC7DEF945D799B64689070C862C">
<mods:date id="416D5BEEF3A47E372C120A36E0667785">2011</mods:date>
<mods:detail id="33BF68CEDBBE1673EE0BD51AC1161B0E" type="volume">
<mods:number id="CC2BC0D38265D10252656BFA6F505E9A">3014</mods:number>
</mods:detail>
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<mods:start id="8B955B2A9CF5EBFD2DF9A158C9E7ADB7">35</mods:start>
<mods:end id="AECA531EF2C1C85C73FA8AFAB44A32D7">38</mods:end>
</mods:extent>
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</mods:relatedItem>
<mods:classification id="28B38F28FFAA0A4B7009788C628D9C3A">journal article</mods:classification>
<mods:identifier id="2D4932EC7E6C454B1236B70C262B5D4D" type="CLB-Dataset">46415</mods:identifier>
<mods:identifier id="F2DA71ED1B5D492C2BF36A774A577276" type="DOI">10.5281/zenodo.204428</mods:identifier>
<mods:identifier id="BD4ED05A79525C447206E820E26DC4B4" type="GBIF-Dataset">b27a74a0-2042-4720-839a-c5176cb48a38</mods:identifier>
<mods:identifier id="FEFB79CD15B2CB5EA8C8FCE2AE5A6638" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1FC836BFF91C21687F1D230B2F0DBA35" type="Zenodo-Dep">204428</mods:identifier>
</mods:mods>
<treatment id="1D6FB737FF9EFFABFF2FFEC850E3F957" ID-DOI="http://doi.org/10.5281/zenodo.5673953" ID-GBIF-Taxon="119415362" ID-Zenodo-Dep="5673953" LSID="urn:lsid:plazi:treatment:1D6FB737FF9EFFABFF2FFEC850E3F957" httpUri="http://treatment.plazi.org/id/1D6FB737FF9EFFABFF2FFEC850E3F957" lastPageId="11" lastPageNumber="46" pageId="9" pageNumber="44">
<subSubSection id="DDDC55AAFF9EFFA9FF2FFEC85624FEF5" pageId="9" pageNumber="44" type="nomenclature">
<paragraph id="95790621FF9EFFA9FF2FFEC8566AFE94" blockId="9.[151,638,290,349]" box="[151,638,290,316]" pageId="9" pageNumber="44">
<heading id="CE31B14DFF9EFFA9FF2FFEC8566AFE94" bold="true" box="[151,638,290,316]" fontSize="11" level="1" pageId="9" pageNumber="44" reason="1">
<emphasis id="A7B2DA33FF9EFFA9FF2FFEC8566AFE94" bold="true" box="[151,638,290,316]" pageId="9" pageNumber="44">
<taxonomicName id="52C67DA2FF9EFFA9FF2FFEC8560EFE94" ID-CoL="8BP2N" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[151,538,290,316]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF9EFFA9FF2FFEC8557DFE94" bold="true" box="[151,361,290,316]" italics="true" pageId="9" pageNumber="44">Culicoides zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<taxonomicNameLabel id="BC816748FF9EFFA9FD99FEC8566AFE94" box="[545,638,290,316]" pageId="9" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="95790621FF9EFFA9FF2FFEAF5624FEF5" blockId="9.[151,638,290,349]" box="[151,560,325,349]" pageId="9" pageNumber="44">
(
<figureCitation id="0DFD1AA4FF9EFFA9FF27FEAF54F3FEF5" box="[159,231,325,349]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1399,1422]" captionTargetBox="[187,1398,218,1321]" captionTargetId="figure@4.[151,1436,193,1378]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Eye separation of female Culicoides (Marksomyia) species. 1. C. marksi Lee &amp; Reye. 2. C. pseudostigmaticus Tokunaga. 3. C. zentae Bellis &amp; Dyce sp. nov. 4. C. kayi Bellis &amp; Dyce sp. nov. 5. C. parvimaculatus Lee &amp; Reye. 6. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204429/files/figure.png" pageId="9" pageNumber="44">Figs 3</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FF4BFEAF5517FEF5" box="[243,259,325,349]" captionStart="FIGURES 7 – 12" captionStartId="5.[151,264,1892,1915]" captionTargetBox="[195,1309,231,1843]" captionTargetId="figure@5.[151,1436,193,1871]" captionTargetPageId="5" captionText="FIGURES 7 – 12. Female Culicoides (Marksomyia) species. 7. C. marksi Lee &amp; Reye, mandibular teeth and palp. 8. C. pseudostigmaticus Tokunaga, mandibular teeth and palp. 9. C. zentae Bellis &amp; Dyce sp. nov., mandibular teeth and palp. 10. C. kayi Bellis &amp; Dyce sp. nov., mandibular teeth and palp. 11. C. parvimaculatus Lee &amp; Reye, mandibular teeth and palp. 12. C. dycei Lee &amp; Reye, mandibular teeth and palp." httpUri="https://zenodo.org/record/204430/files/figure.png" pageId="9" pageNumber="44">9</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FEB7FEAF5539FEF5" box="[271,301,325,349]" captionStart="FIGURES 13 – 18" captionStartId="6.[151,264,1104,1127]" captionTargetBox="[202,1435,214,1036]" captionTargetId="figure@6.[151,1436,193,1083]" captionTargetPageId="6" captionText="FIGURES 13 – 18. Antennae of female Culicoides (Marksomyia) species. 13. C. marksi Lee &amp; Reye, antennal segment 6 and antenna. 14. C. pseudostigmaticus Tokunaga. 15. C. zentae Bellis &amp; Dyce sp. nov. 16. C. kayi Bellis &amp; Dyce sp. nov. 17. C. parvimaculatus Lee &amp; Reye. 18. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204431/files/figure.png" pageId="9" pageNumber="44">15</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FE81FEAF5543FEF5" box="[313,343,325,349]" captionStart="FIGURES 19 – 24" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[187,1404,215,1824]" captionTargetId="figure@7.[151,1436,193,1871]" captionTargetPageId="7" captionText="FIGURES 19 – 24. Female Culicoides (Marksomyia) species 19. C. marksi Lee &amp; Reye, legs and tibial comb. 20. C. pseudostigmaticus Tokunaga, legs and tibial comb. 21. C. zentae Bellis &amp; Dyce sp. nov., legs and tibial comb. 22. C. kayi Bellis &amp; Dyce sp. nov., legs and tibial comb. 23. C. parvimaculatus Lee &amp; Reye, legs and tibial comb. 24. C. dycei Lee &amp; Reye, legs and tibial comb." httpUri="https://zenodo.org/record/204432/files/figure.png" pageId="9" pageNumber="44">21</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FEDBFEAF5596FEF5" box="[355,386,325,349]" captionStart="FIGURES 25 – 32" captionStartId="8.[151,264,1616,1639]" captionTargetBox="[160,1418,200,1568]" captionTargetId="figure@8.[151,1436,178,1608]" captionTargetPageId="8" captionText="FIGURES 25 – 32. Wings of female Culicoides (Marksomyia) species. 25. C. marksi Lee &amp; Reye. 26. C. marksi Lee &amp; Reye, rare aberrant wing pattern. 27. C. pseudostigmaticus Tokunaga. 28. C. zentae Bellis &amp; Dyce sp. nov. 29. C. kayi Bellis &amp; Dyce sp. nov. 30. C. parvimaculatus Lee &amp; Reye. 31.. C. dycei Lee &amp; Reye 32. C. dycei Lee &amp; Reye, rare aberrant wing pattern." httpUri="https://zenodo.org/record/204433/files/figure.png" pageId="9" pageNumber="44">28</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FE35FEAF55BFFEF5" box="[397,427,325,349]" captionStart="FIGURES 33 – 38" captionStartId="10.[151,264,1892,1915]" captionTargetBox="[186,1399,217,1834]" captionTargetId="figure@10.[151,1436,193,1871]" captionTargetPageId="10" captionText="FIGURES 33 – 38. Female Culicoides (Marksomyia) species. 33. C. marksi Lee &amp; Reye, spermathecae and genital sclerotisation. 34. C. pseudostigmaticus Tokunaga, spermathecae and genital sclerotisation. 35. C. zentae Bellis &amp; Dyce sp. nov., spermathecae and genital sclerotisation. 36. C. kayi Bellis &amp; Dyce sp. nov., spermathecae and genital sclerotisation. 37. C. parvimaculatus Lee &amp; Reye, spermathecae and genital sclerotisation. 38. C. dycei Lee &amp; Reye, spermathecae and genital sclerotisation." httpUri="https://zenodo.org/record/204434/files/figure.png" pageId="9" pageNumber="44">35</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FE0FFEAF55C1FEF5" box="[439,469,325,349]" captionStart="FIGURES 39 – 44" captionStartId="11.[151,264,1151,1174]" captionTargetBox="[193,1391,223,1085]" captionTargetId="figure@11.[151,1436,193,1130]" captionTargetPageId="11" captionText="FIGURES 39 – 44. Antennae of male Culicoides (Marksomyia) species. Plume aristae are not shown although attachment bases are indicated where present. 39. C. marksi Lee &amp; Reye. 40. C. pseudostigmaticus Tokunaga. 41. C. zentae Bellis &amp; Dyce sp. nov. 42. C. kayi Bellis &amp; Dyce sp. nov. 43. C. parvimaculatus Lee &amp; Reye. 44. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204435/files/figure.png" pageId="9" pageNumber="44">41</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FE59FEAF55EBFEF5" box="[481,511,325,349]" captionStart="FIGURES 45 – 50" captionStartId="13.[151,264,1876,1899]" captionTargetBox="[299,1291,193,1847]" captionTargetId="figure@13.[298,1292,193,1847]" captionTargetPageId="13" captionText="FIGURES 45 – 50. Male Culicoides (Marksomyia) species 45. C. marksi Lee &amp; Reye, hypopygium, parameres and aedeagus. 46. C. pseudostigmaticus Tokunaga, hypopygium, parameres and aedeagus. 47. C. zentae Bellis &amp; Dyce sp. nov., hypopygium, parameres and aedeagus. 48. C. kayi Bellis &amp; Dyce sp. nov., hypopygium, parameres and aedeagus. 49. C. parvimaculatus Lee &amp; Reye, hypopygium, parameres and aedeagus. 50. C. dycei Lee &amp; Reye, hypopygium, parameres and aedeagus." httpUri="https://zenodo.org/record/204436/files/figure.png" pageId="9" pageNumber="44">47</figureCitation>
,
<figureCitation id="0DFD1AA4FF9EFFA9FDB3FEAF563CFEF5" box="[523,552,325,349]" captionStart="FIGURES 51 – 56" captionStartId="14.[151,264,1155,1178]" captionTargetBox="[151,1425,200,1083]" captionTargetId="figure@14.[151,1436,193,1134]" captionTargetPageId="14" captionText="FIGURES 51 – 56. Wings of male Culicoides (Marksomyia) species. 51. C. marksi Lee &amp; Reye. 52. C. pseudostigmaticus Tokunaga. 53. C. zentae Bellis &amp; Dyce sp. nov. 54. C. kayi Bellis &amp; Dyce sp. nov. 55. C. parvimaculatus Lee &amp; Reye. 56. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204437/files/figure.png" pageId="9" pageNumber="44">53</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9EFFA9FF2FFE625176FD8D" pageId="9" pageNumber="44" type="materials_examined">
<paragraph id="95790621FF9EFFA9FF2FFE625176FD8D" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF2FFE625557FE08" bold="true" box="[151,323,391,416]" pageId="9" pageNumber="44">
<typeStatus id="4A7DB883FF9EFFA9FF2FFE6254C4FE08" box="[151,208,392,416]" pageId="9" pageNumber="44">Type</typeStatus>
material.
</emphasis>
<typeStatus id="4A7DB883FF9EFFA9FEF5FE6D55A1FE37" box="[333,437,391,415]" pageId="9" pageNumber="44" type="holotype">Holotype</typeStatus>
female,
<emphasis id="A7B2DA33FF9EFFA9FDA0FE6D56D1FE08" bold="true" box="[536,709,391,416]" pageId="9" pageNumber="44">
<collectingCountry id="EDD146B1FF9EFFA9FDA0FE6D569CFE08" box="[536,648,391,416]" name="Australia" pageId="9" pageNumber="44">Australia</collectingCountry>
Qld,
</emphasis>
Gilruth, Bore drain, sweep net, sunset,
<date id="E17820E1FF9EFFA9FB3EFE6D5113FE37" box="[1158,1287,391,415]" pageId="9" pageNumber="44" value="1963-07-26">26.vii.1963</date>
, A.L. Dyce (female,
<collectionCode id="F3D79EE4FF9EFFA9FF4AFE43552FFE69" LSID="urn:lsid:biocol.org:col:32981" box="[242,315,425,449]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="44">ANIC</collectionCode>
).
<emphasis id="A7B2DA33FF9EFFA9FEF6FE4355D2FE69" bold="true" box="[334,454,425,449]" pageId="9" pageNumber="44">
<typeStatus id="4A7DB883FF9EFFA9FEF6FE4355D2FE69" box="[334,454,425,449]" pageId="9" pageNumber="44" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<emphasis id="A7B2DA33FF9EFFA9FE6EFE42569DFE69" bold="true" box="[470,649,424,449]" pageId="9" pageNumber="44">
<collectingCountry id="EDD146B1FF9EFFA9FE6EFE42565CFE69" box="[470,584,424,449]" name="Australia" pageId="9" pageNumber="44">Australia</collectingCountry>
, Qld,
</emphasis>
Gilruth, Light trap, Bore drain,
<date id="E17820E1FF9EFFA9FC4EFE43506CFE69" box="[1014,1144,425,449]" pageId="9" pageNumber="44" value="1963-07-26">26.vii.1963</date>
, A.L. Dyce &amp; M.D. Murray (
<specimenCount id="83C0CDA8FF9EFFA9FF75FE205529FE4A" box="[205,317,458,482]" pageId="9" pageNumber="44" type="female">9 females</specimenCount>
<specimenCount id="83C0CDA8FF9EFFA9FEFFFE205581FE4A" box="[327,405,458,482]" pageId="9" pageNumber="44" type="male">1 male</specimenCount>
<collectionCode id="F3D79EE4FF9EFFA9FE27FE2055FCFE4A" LSID="urn:lsid:biocol.org:col:32981" box="[415,488,458,482]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="44">ANIC</collectionCode>
); Charleville,
<date id="E17820E1FF9EFFA9FD34FE205715FE4A" box="[652,769,458,482]" pageId="9" pageNumber="44" value="1969-02-20">20.ii.1969</date>
, Truck trap run 18.30–19.00 hrs, I. Fanning &amp; B. Kay (
<specimenCount id="83C0CDA8FF9EFFA9FA36FE2054CCFDAC" pageId="9" pageNumber="44" type="male">8 males</specimenCount>
<collectionCode id="F3D79EE4FF9EFFA9FF59FE06553FFDAC" LSID="urn:lsid:biocol.org:col:32981" box="[225,299,492,516]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="44">ANIC</collectionCode>
); Charleville,
<date id="E17820E1FF9EFFA9FE73FE06562AFDAC" box="[459,574,492,516]" pageId="9" pageNumber="44" value="1969-02-18">18.ii.1969</date>
, Wallal Bore, A.L. Dyce, (
<specimenCount id="83C0CDA8FF9EFFA9FCCBFE0657F0FDAC" box="[883,996,492,516]" pageId="9" pageNumber="44" type="female">5 females</specimenCount>
,
<specimenCount id="83C0CDA8FF9EFFA9FC49FE06502AFDAC" box="[1009,1086,492,516]" pageId="9" pageNumber="44" type="male">1 male</specimenCount>
and associated pupae,
<collectionCode id="F3D79EE4FF9EFFA9FAFEFE065184FDAC" LSID="urn:lsid:biocol.org:col:32981" box="[1350,1424,492,516]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="9" pageNumber="44">ANIC</collectionCode>
);
<emphasis id="A7B2DA33FF9EFFA9FF2FFDE754A8FD8D" bold="true" box="[151,188,525,549]" pageId="9" pageNumber="44">NT</emphasis>
, Vic River Research Station,
<geoCoordinate id="F0F260E6FF9EFFA9FDBFFDE75671FD8D" box="[519,613,525,549]" direction="south" orientation="latitude" pageId="9" pageNumber="44" precision="925" value="-16.4">16°24’S</geoCoordinate>
,
<geoCoordinate id="F0F260E6FF9EFFA9FDC9FDE756CBFD8D" box="[625,735,525,549]" direction="east" orientation="longitude" pageId="9" pageNumber="44" precision="925" value="131.01666">131°01’E</geoCoordinate>
,
<date id="E17820E1FF9EFFA9FD53FDE75749FD8D" box="[747,861,525,549]" pageId="9" pageNumber="44" value="2003-10-28">28.x.2003</date>
, D. Cherry L.T. 03-1615 (
<specimenCount id="83C0CDA8FF9EFFA9FB3CFDE750F7FD8D" box="[1156,1251,525,549]" pageId="9" pageNumber="44" type="female">1 female</specimenCount>
AQISNT).
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9EFFA9FF7FFDC455D3FCA6" pageId="9" pageNumber="44" type="diagnosis">
<paragraph id="95790621FF9EFFA9FF7FFDC455D3FCA6" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7FFDC45555FDEF" bold="true" box="[199,321,558,583]" pageId="9" pageNumber="44">Diagnosis.</emphasis>
Wing with 2 pale spots in cell M4, anterior spot rounded or triangular. Female with SCo distribution 3, (4–6),7,(8),9,11–15, six SCh on 4–10, two to three on 11–15; spermathecae with unsclerotised, pear-shaped body and short, sclerotised necks. Male with SCo distribution 3, (4),13–15; STl distribution 3–6; STc distribution 4–6; six or seven SCh on 14; apical half of parameres strongly curved laterad and with triangular excavation medially, ventral membrane of ninth sternite with spicules present laterally, medially bare. Pupal abdomen with paired lateral spines on
<emphasis id="A7B2DA33FF9EFFA9FEE3FD3F5591FD45" box="[347,389,725,749]" italics="true" pageId="9" pageNumber="44">lpm</emphasis>
,
<emphasis id="A7B2DA33FF9EFFA9FE2EFD3F55DFFD45" box="[406,459,725,749]" italics="true" pageId="9" pageNumber="44">lasm</emphasis>
and
<emphasis id="A7B2DA33FF9EFFA9FDBEFD3F5655FD45" box="[518,577,725,749]" italics="true" pageId="9" pageNumber="44">dasm</emphasis>
, thorax with paired lateral spines on
<emphasis id="A7B2DA33FF9EFFA9FC4AFD3F5022FD45" box="[1010,1078,725,749]" italics="true" pageId="9" pageNumber="44">di. dii</emphasis>
and
<emphasis id="A7B2DA33FF9EFFA9FBC9FD3F5087FD45" box="[1137,1171,725,749]" italics="true" pageId="9" pageNumber="44">div</emphasis>
; prothoracic horn with scales covering basal third.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9EFFABFF7FFCF257EBFA93" lastPageId="11" lastPageNumber="46" pageId="9" pageNumber="44" type="description">
<paragraph id="95790621FF9EFFA9FF7FFCF25131FCDA" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7FFCF25560FC98" bold="true" box="[199,372,792,816]" pageId="9" pageNumber="44">Female. Head.</emphasis>
Eyes separated by a distance of about one facet (
<figureCitation id="0DFD1AA4FF9EFFA9FC1AFCFD57F4FC87" box="[930,992,791,815]" captionStart="FIGURES 1 – 6" captionStartId="4.[151,264,1399,1422]" captionTargetBox="[187,1398,218,1321]" captionTargetId="figure@4.[151,1436,193,1378]" captionTargetPageId="4" captionText="FIGURES 1 – 6. Eye separation of female Culicoides (Marksomyia) species. 1. C. marksi Lee &amp; Reye. 2. C. pseudostigmaticus Tokunaga. 3. C. zentae Bellis &amp; Dyce sp. nov. 4. C. kayi Bellis &amp; Dyce sp. nov. 5. C. parvimaculatus Lee &amp; Reye. 6. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204429/files/figure.png" pageId="9" pageNumber="44">Fig 3</figureCitation>
); proboscis short. Palpus (fig 9) brown with five segments, segment 3 expanding apically then abruptly narrowed beyond a round, shallow sensory pit with several protruding capitate sensilla. Antennomeres (fig 15) 4–10 barrel shaped, 11–15 cylindrical, short.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7DFC965063FC74" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7DFC965530FC3C" bold="true" box="[197,292,892,916]" pageId="9" pageNumber="44">Thorax.</emphasis>
Legs (fig 21) dark brown with dark knees, all femora pale from base to about midlength and with pale subapical band, all tibiae with pale sub basal band and broadly pale subapically (less obvious on mid tibiae), tarsi pale. Wing (fig 28) strongly patterned with two ovoid to triangular pale spots in cell M4.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7DFC0D556DFBEF" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7DFC0D5555FBA8" bold="true" box="[197,321,999,1024]" pageId="9" pageNumber="44">Abdomen.</emphasis>
Three developed, pear-shaped spermathecae, unsclerotised and difficult to see in most specimens but necks strongly sclerotised and conspicuous (fig 35), ducts short, about as long as spermathecae; sclerotised ring tapering posteriorly.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7FFBBE5744FAFD" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7FFBBE554DFBC4" bold="true" box="[199,345,1108,1132]" pageId="9" pageNumber="44">Male. Head.</emphasis>
Eyes bare. Palpus similar to female with shallow pit on segment 3. Antenna (
<figureCitation id="0DFD1AA4FF9EFFA9FB7DFBBE5104FBC4" box="[1221,1296,1108,1132]" captionStart="FIGURES 39 – 44" captionStartId="11.[151,264,1151,1174]" captionTargetBox="[193,1391,223,1085]" captionTargetId="figure@11.[151,1436,193,1130]" captionTargetPageId="11" captionText="FIGURES 39 – 44. Antennae of male Culicoides (Marksomyia) species. Plume aristae are not shown although attachment bases are indicated where present. 39. C. marksi Lee &amp; Reye. 40. C. pseudostigmaticus Tokunaga. 41. C. zentae Bellis &amp; Dyce sp. nov. 42. C. kayi Bellis &amp; Dyce sp. nov. 43. C. parvimaculatus Lee &amp; Reye. 44. C. dycei Lee &amp; Reye." httpUri="https://zenodo.org/record/204435/files/figure.png" pageId="9" pageNumber="44">Fig 41</figureCitation>
) with single row of plume verticils on antennomeres 3–12; antennomeres 13–15 elongate subcylindrical and narrow.
<emphasis id="A7B2DA33FF9EFFA9FF2FFB7C5516FB07" bold="true" box="[151,258,1174,1199]" italics="true" pageId="9" pageNumber="44">Genitalia</emphasis>
. (fig 47) Ninth tergite with caudal margin slightly convex. Ninth sternite with deep, U shaped caudomedial excavation, spicules confined to two posterior lateral patches. Gonocoxite short and broad, longer than wide, dorsal root long and simple; ventral root reduced to rounded point. Gonostylus slender, distally curving gently to a pointed apex. Aedeagus with distal process short with a slight apical rounded expansion. Parameres separate, with triangular excavation near base, basal arms roughly parallel, stem weakly swollen narrowing gradually to simple sharp-tipped distal portion curved lateroventrad to almost 90°.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7FFAB5560AFADE" blockId="9.[151,1437,391,1989]" box="[199,542,1374,1399]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7FFAB55546FADF" bold="true" box="[199,338,1375,1399]" pageId="9" pageNumber="44">Immatures.</emphasis>
Larvae: unknown
</paragraph>
<paragraph id="95790621FF9EFFA9FF7FFA6A5500FA72" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7FFA6A554EFA30" bold="true" box="[199,346,1408,1432]" pageId="9" pageNumber="44">Pupa. Head.</emphasis>
(fig 60): Setae
<emphasis id="A7B2DA33FF9EFFA9FDB2FA6B563FFA30" box="[522,555,1409,1432]" italics="true" pageId="9" pageNumber="44">vm</emphasis>
and
<emphasis id="A7B2DA33FF9EFFA9FDDAFA6A5662FA30" box="[610,630,1408,1432]" italics="true" pageId="9" pageNumber="44">vl</emphasis>
with each group consisting of a long and a short seta;
<emphasis id="A7B2DA33FF9EFFA9FB66FA6B5114FA30" box="[1246,1280,1409,1432]" italics="true" pageId="9" pageNumber="44">am</emphasis>
(fig 61) large tubercule bearing a blunt spur, a stout seta and a sensillum;
<emphasis id="A7B2DA33FF9EFFA9FCF2FA4B5772FA11" box="[842,870,1441,1465]" italics="true" pageId="9" pageNumber="44">ad</emphasis>
(fig 61) a large tubercule with a pair of unequal, stout setae.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7FFA0954D6F9F7" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7FFA095542FA54" bold="true" box="[199,342,1507,1532]" pageId="9" pageNumber="44">Operculum.</emphasis>
(fig 61): A single row of bluntly pointed thorns along lateral margins posterior to lateral angles, extending to about ¾ of operculum length; two groups of thorns either side of midline posterior to each
<emphasis id="A7B2DA33FF9EFFA9FA86F9EC5174F9B5" box="[1342,1376,1542,1565]" italics="true" pageId="9" pageNumber="44">am</emphasis>
, narrowly connected anteriorly; oval structures between lateral rows of thorns extending between
<emphasis id="A7B2DA33FF9EFFA9FB09F9C250C7F997" box="[1201,1235,1576,1599]" italics="true" pageId="9" pageNumber="44">am</emphasis>
’s to anterior margin.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7DF9805068F90D" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7DF9805531F92A" bold="true" box="[197,293,1642,1666]" pageId="9" pageNumber="44">Thorax.</emphasis>
(fig 57): Tubercules
<emphasis id="A7B2DA33FF9EFFA9FDABF9835640F929" box="[531,596,1641,1665]" italics="true" pageId="9" pageNumber="44">di, dii</emphasis>
, and
<emphasis id="A7B2DA33FF9EFFA9FD2BF98356A1F929" box="[659,693,1641,1665]" italics="true" pageId="9" pageNumber="44">div</emphasis>
with two pointed lateral spines and a stout seta;
<emphasis id="A7B2DA33FF9EFFA9FB62F98350EBF929" box="[1242,1279,1641,1665]" italics="true" pageId="9" pageNumber="44">diii</emphasis>
ridged with a small seta;
<emphasis id="A7B2DA33FF9EFFA9FEADF9675523F90D" box="[277,311,1677,1701]" italics="true" pageId="9" pageNumber="44">dm</emphasis>
a long, thin seta;
<emphasis id="A7B2DA33FF9EFFA9FE46F9675600F90D" box="[510,532,1677,1701]" italics="true" pageId="9" pageNumber="44">dl</emphasis>
long thin seta and short stout seta on protuberant base.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7DF95B55D2F8B9" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7DF95B5589F962" bold="true" box="[197,413,1713,1738]" pageId="9" pageNumber="44">Prothoracic horn.</emphasis>
(fig 62): Pigmented throughout but darker near base, on lateral tubercules and in distal third; folds and sharply pointed scales on basal third. Four to six (mean 5.18) terminal and one to three (mean 2) lateral spiracular openings.
</paragraph>
<paragraph id="95790621FF9EFFA9FF7DF8F75682F86D" blockId="9.[151,1437,391,1989]" pageId="9" pageNumber="44">
<emphasis id="A7B2DA33FF9EFFA9FF7DF8F75555F89E" bold="true" box="[197,321,1821,1846]" pageId="9" pageNumber="44">Abdomen.</emphasis>
(fig 58): Setae
<emphasis id="A7B2DA33FF9EFFA9FE54F8F75658F89D" box="[492,588,1821,1845]" italics="true" pageId="9" pageNumber="44">dpm i, iv</emphasis>
and
<emphasis id="A7B2DA33FF9EFFA9FD39F8F45699F89D" box="[641,653,1822,1845]" italics="true" pageId="9" pageNumber="44">v</emphasis>
with paired spines and stout setae, seta on
<emphasis id="A7B2DA33FF9EFFA9FBD2F8F45066F89D" box="[1130,1138,1822,1845]" italics="true" pageId="9" pageNumber="44">i</emphasis>
much shorter than on
<emphasis id="A7B2DA33FF9EFFA9FAD2F8F4516AF89D" box="[1386,1406,1822,1845]" italics="true" pageId="9" pageNumber="44">iv</emphasis>
or
<emphasis id="A7B2DA33FF9EFFA9FF2FF8A854B7F8F1" box="[151,163,1858,1881]" italics="true" pageId="9" pageNumber="44">v</emphasis>
;
<emphasis id="A7B2DA33FF9EFFA9FF0AF8A854D5F8F1" box="[178,193,1858,1881]" italics="true" pageId="9" pageNumber="44">ii</emphasis>
and
<emphasis id="A7B2DA33FF9EFFA9FF41F8A85504F8F1" box="[249,272,1858,1881]" italics="true" pageId="9" pageNumber="44">iii</emphasis>
ridged tubercules with one rounded spine and no setae.
<emphasis id="A7B2DA33FF9EFFA9FC32F8AB57A0F8F1" box="[906,948,1857,1881]" italics="true" pageId="9" pageNumber="44">lpm</emphasis>
all with paired sharply pointed spurs, i and
<emphasis id="A7B2DA33FF9EFFA9FF2FF88C54BBF8D5" box="[151,175,1894,1917]" italics="true" pageId="9" pageNumber="44">iii</emphasis>
with short stout seta,
<emphasis id="A7B2DA33FF9EFFA9FE13F88C55AEF8D5" box="[427,442,1894,1917]" italics="true" pageId="9" pageNumber="44">ii</emphasis>
with long thin seta;
<emphasis id="A7B2DA33FF9EFFA9FD1EF88C56A3F8D5" box="[678,695,1894,1917]" italics="true" pageId="9" pageNumber="44">V</emphasis>
ridged tubercules, i with short seta,
<emphasis id="A7B2DA33FF9EFFA9FBE0F88C5073F8D5" box="[1112,1127,1894,1917]" italics="true" pageId="9" pageNumber="44">ii</emphasis>
and
<emphasis id="A7B2DA33FF9EFFA9FB19F88C50ACF8D5" box="[1185,1208,1894,1917]" italics="true" pageId="9" pageNumber="44">iii</emphasis>
with long thin seta;
<emphasis id="A7B2DA33FF9EFFA9FF2FF86354C6F809" box="[151,210,1929,1953]" italics="true" pageId="9" pageNumber="44">dasm</emphasis>
with paired rounded spines and long (
<emphasis id="A7B2DA33FF9EFFA9FD39F860569DF809" box="[641,649,1930,1953]" italics="true" pageId="9" pageNumber="44">i</emphasis>
) or short (
<emphasis id="A7B2DA33FF9EFFA9FCB8F860571BF809" box="[768,783,1930,1953]" italics="true" pageId="9" pageNumber="44">ii</emphasis>
) seta;
<emphasis id="A7B2DA33FF9EFFA9FCE3F8635784F809" box="[859,912,1929,1953]" italics="true" pageId="9" pageNumber="44">lasm</emphasis>
similar to
<emphasis id="A7B2DA33FF9EFFA9FBB0F8635026F809" box="[1032,1074,1929,1953]" italics="true" pageId="9" pageNumber="44">lpm</emphasis>
. Anterior band of spicules present dorsally and ventrally but absent laterally.
</paragraph>
<caption id="C1B956A9FF9DFFAAFF2FF88E54EAF85D" httpUri="https://zenodo.org/record/204434/files/figure.png" pageId="10" pageNumber="45" targetBox="[186,1399,217,1834]" targetPageId="10">
<paragraph id="95790621FF9DFFAAFF2FF88E54EAF85D" blockId="10.[151,1436,1892,2037]" pageId="10" pageNumber="45">
<emphasis id="A7B2DA33FF9DFFAAFF2FF88E5541F8D2" bold="true" box="[151,341,1892,1915]" pageId="10" pageNumber="45">FIGURES 33–38.</emphasis>
Female
<taxonomicName id="52C67DA2FF9DFFAAFE15F88F56A5F8D2" authority="Marksomyia" authorityName="Marksomyia" box="[429,689,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF9DFFAAFE15F88F560DF8D3" box="[429,537,1893,1915]" italics="true" pageId="10" pageNumber="45">Culicoides</emphasis>
(
<emphasis id="A7B2DA33FF9DFFAAFD92F88F56BCF8D3" box="[554,680,1893,1915]" italics="true" pageId="10" pageNumber="45">Marksomyia</emphasis>
)
</taxonomicName>
species. 33.
<taxonomicName id="52C67DA2FF9DFFAAFC8EF88F500BF8D2" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[822,1055,1893,1915]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF9DFFAAFC8EF88F578DF8D3" box="[822,921,1893,1915]" italics="true" pageId="10" pageNumber="45">C. marksi</emphasis>
Lee &amp; Reye
</taxonomicName>
, spermathecae and genital sclerotisation. 34.
<taxonomicName id="52C67DA2FF9DFFAAFF4BF86E562DF831" authority="Tokunaga" authorityName="Tokunaga" box="[243,569,1923,1945]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF9DFFAAFF4BF86E55D8F831" box="[243,460,1923,1945]" italics="true" pageId="10" pageNumber="45">C. pseudostigmaticus</emphasis>
Tokunaga
</taxonomicName>
, spermathecae and genital sclerotisation. 35.
<taxonomicName id="52C67DA2FF9DFFAAFBB2F86E5117F831" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[1034,1283,1923,1945]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF9DFFAAFBB2F86E507DF831" box="[1034,1129,1924,1945]" italics="true" pageId="10" pageNumber="45">C. zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF9DFFAAFAB2F86E5148F831" bold="true" box="[1290,1372,1924,1945]" pageId="10" pageNumber="45">
<taxonomicNameLabel id="BC816748FF9DFFAAFAB2F86E5148F831" box="[1290,1372,1924,1945]" pageId="10" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, spermathecae and genital sclerotisation. 36.
<taxonomicName id="52C67DA2FF9DFFAAFDFDF849572DF810" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[581,825,1954,1976]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="kayi" status="sp. nov.">
<emphasis id="A7B2DA33FF9DFFAAFDFDF8495686F810" box="[581,658,1954,1976]" italics="true" pageId="10" pageNumber="45">C. kayi</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF9DFFAAFCFCF848578DF81F" bold="true" box="[836,921,1954,1975]" pageId="10" pageNumber="45">
<taxonomicNameLabel id="BC816748FF9DFFAAFCFCF848578DF81F" box="[836,921,1954,1975]" pageId="10" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, spermathecae and genital sclerotisation. 37.
<taxonomicName id="52C67DA2FF9DFFAAFA3CF84955ACF87E" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF9DFFAAFA3CF8495520F87E" italics="true" pageId="10" pageNumber="45">C. parvimaculatus</emphasis>
Lee &amp; Reye
</taxonomicName>
, spermathecae and genital sclerotisation. 38.
<taxonomicName id="52C67DA2FF9DFFAAFC3CF82B504FF87E" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[900,1115,1984,2006]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF9DFFAAFC3CF82B57C3F87E" box="[900,983,1984,2006]" italics="true" pageId="10" pageNumber="45">C. dycei</emphasis>
Lee &amp; Reye
</taxonomicName>
, spermathecae and genital sclerotisation.
</paragraph>
</caption>
<caption id="C1B956A9FF9CFFABFF2FFB95508FFB7A" httpUri="https://zenodo.org/record/204435/files/figure.png" pageId="11" pageNumber="46" targetBox="[193,1391,223,1085]" targetPageId="11">
<paragraph id="95790621FF9CFFABFF2FFB95508FFB7A" blockId="11.[151,1436,1151,1235]" pageId="11" pageNumber="46">
<emphasis id="A7B2DA33FF9CFFABFF2FFB955547FB3D" bold="true" box="[151,339,1151,1174]" pageId="11" pageNumber="46">FIGURES 39–44.</emphasis>
Antennae of male
<taxonomicName id="52C67DA2FF9CFFABFDAAFB955700FB3D" authority="Marksomyia" authorityName="Marksomyia" box="[530,788,1151,1173]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="genus">
<emphasis id="A7B2DA33FF9CFFABFDAAFB95566AFB3D" box="[530,638,1151,1173]" italics="true" pageId="11" pageNumber="46">Culicoides</emphasis>
(
<emphasis id="A7B2DA33FF9CFFABFD35FB955718FB3D" box="[653,780,1151,1173]" italics="true" pageId="11" pageNumber="46">Marksomyia</emphasis>
)
</taxonomicName>
species. Plume aristae are not shown although attachment bases are indicated where present. 39.
<taxonomicName id="52C67DA2FF9CFFABFE5DFB7556C7FB1C" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." box="[485,723,1182,1204]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="marksi">
<emphasis id="A7B2DA33FF9CFFABFE5DFB75565DFB1C" box="[485,585,1182,1204]" italics="true" pageId="11" pageNumber="46">C. marksi</emphasis>
Lee &amp; Reye.
</taxonomicName>
40.
<taxonomicName id="52C67DA2FF9CFFABFCBAFB75505AFB1C" authority="Tokunaga." authorityName="Tokunaga." box="[770,1102,1182,1204]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="pseudostigmaticus">
<emphasis id="A7B2DA33FF9CFFABFCBAFB7557C8FB1C" box="[770,988,1182,1204]" italics="true" pageId="11" pageNumber="46">C. pseudostigmaticus</emphasis>
Tokunaga.
</taxonomicName>
41.
<taxonomicName id="52C67DA2FF9CFFABFBC4FB755163FB1C" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[1148,1399,1182,1204]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="zentae" status="sp. nov.">
<emphasis id="A7B2DA33FF9CFFABFBC4FB7550C8FB1C" box="[1148,1244,1183,1204]" italics="true" pageId="11" pageNumber="46">C. zentae</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF9CFFABFAC5FB7554D7FB7A" bold="true" pageId="11" pageNumber="46">
<taxonomicNameLabel id="BC816748FF9CFFABFAC5FB7554D7FB7A" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
42.
<taxonomicName id="52C67DA2FF9CFFABFF57FB5755DBFB7A" authority="Bellis &amp; Dyce" authorityName="Bellis &amp; Dyce" box="[239,463,1212,1235]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="kayi" status="sp. nov.">
<emphasis id="A7B2DA33FF9CFFABFF57FB575523FB7B" box="[239,311,1213,1235]" italics="true" pageId="11" pageNumber="46">C. kayi</emphasis>
Bellis &amp; Dyce
</taxonomicName>
<emphasis id="A7B2DA33FF9CFFABFE6CFB575631FB7A" bold="true" box="[468,549,1213,1234]" pageId="11" pageNumber="46">
<taxonomicNameLabel id="BC816748FF9CFFABFE6CFB575631FB7A" box="[468,549,1213,1234]" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
43.
<taxonomicName id="52C67DA2FF9CFFABFDEAFB575781FB7A" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." box="[594,917,1213,1235]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF9CFFABFDEAFB575718FB7B" box="[594,780,1213,1235]" italics="true" pageId="11" pageNumber="46">C. parvimaculatus</emphasis>
Lee &amp; Reye.
</taxonomicName>
44.
<taxonomicName id="52C67DA2FF9CFFABFC78FB57508FFB7A" authority="Lee &amp; Reye." authorityName="Lee &amp; Reye." box="[960,1179,1213,1235]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="dycei">
<emphasis id="A7B2DA33FF9CFFABFC78FB575007FB7B" box="[960,1043,1213,1235]" italics="true" pageId="11" pageNumber="46">C. dycei</emphasis>
Lee &amp; Reye.
</taxonomicName>
</paragraph>
</caption>
<paragraph id="95790621FF9CFFABFF7DFB1557EBFA93" blockId="11.[151,1436,1279,1792]" pageId="11" pageNumber="46">
<emphasis id="A7B2DA33FF9CFFABFF7DFB155598FAB0" bold="true" box="[197,396,1279,1304]" pageId="11" pageNumber="46">Caudal segment.</emphasis>
(fig 59): Encircling band of spicules anteriorly; median group of v-shaped spicules dorsally; apicolateral processes pigmented distally with spicules scattered over surface.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9CFFABFF7DFAA257BDFA2B" pageId="11" pageNumber="46" type="etymology">
<paragraph id="95790621FF9CFFABFF7DFAA257BDFA2B" blockId="11.[151,1436,1279,1792]" pageId="11" pageNumber="46">
<emphasis id="A7B2DA33FF9CFFABFF7DFAA25558FAC8" bold="true" box="[197,332,1352,1376]" pageId="11" pageNumber="46">Etymology.</emphasis>
This species is named for the late Ms. Zenta Liepa, formerly of CSIRO. Division of Entomology, in recognition for her assistance with the curation of
<taxonomicName id="52C67DA2FF9CFFABFD5EFA81572EFA2B" box="[742,826,1387,1411]" kingdom="Plantae" order="Diptera" pageId="11" pageNumber="46" rank="order">Diptera</taxonomicName>
at ANIC.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9CFFABFF7FFA6550ABFA0F" box="[199,1215,1423,1448]" pageId="11" pageNumber="46" type="distribution">
<paragraph id="95790621FF9CFFABFF7FFA6550ABFA0F" blockId="11.[151,1436,1279,1792]" box="[199,1215,1423,1448]" pageId="11" pageNumber="46">
<emphasis id="A7B2DA33FF9CFFABFF7FFA655574FA00" bold="true" box="[199,352,1423,1448]" pageId="11" pageNumber="46">Distribution.</emphasis>
(fig 65) Northern and eastern
<collectingCountry id="EDD146B1FF9CFFABFD0CFA65570AFA0F" box="[692,798,1423,1447]" name="Australia" pageId="11" pageNumber="46">Australia</collectingCountry>
, more common in drier, inland areas.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9CFFABFF7FFA5A506FF9A3" pageId="11" pageNumber="46" type="biology_ecology">
<paragraph id="95790621FF9CFFABFF7FFA5A506FF9A3" blockId="11.[151,1436,1279,1792]" pageId="11" pageNumber="46">
<emphasis id="A7B2DA33FF9CFFABFF7FFA5A5533FA61" bold="true" box="[199,295,1456,1481]" pageId="11" pageNumber="46">Biology.</emphasis>
Pupae examined in this study were collected from the muddy margins of a
<date id="E17820E1FF9CFFABFBCFFA5A5087FA60" box="[1143,1171,1456,1480]" pageId="11" pageNumber="46">5x</date>
<quantity id="523EABC4FF9CFFABFB2BFA5A50A3FA60" box="[1171,1207,1456,1480]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="46" unit="m" value="3.0">3m</quantity>
pond constantly fed by warm, subterranean water. Adults are often collected in light traps suggesting a crepuscular or nocturnal activity.
<bibRefCitation id="F1577BD0FF9CFFABFF2FFA195567F9A3" box="[151,371,1523,1548]" pageId="11" pageNumber="46" refString="Muller, M. J., Murray, M. D. &amp; Edwards, J. A. (1981) Blood-sucking midges and mosquitoes feeding on mammals at Beatrice Hill, N. T. Australian Journal of Zoology, 29, 573 - 588." type="journal article">
Muller
<emphasis id="A7B2DA33FF9CFFABFF52FA1F5536F9A4" box="[234,290,1524,1548]" italics="true" pageId="11" pageNumber="46">et al.</emphasis>
(1981)
</bibRefCitation>
reported this species (as C. McMaster sp121) feeding on marsupials.
</paragraph>
</subSubSection>
<subSubSection id="DDDC55AAFF9CFFABFF7FF9FF50E3F957" pageId="11" pageNumber="46" type="discussion">
<paragraph id="95790621FF9CFFABFF7FF9FF50C7F9E6" blockId="11.[151,1436,1279,1792]" pageId="11" pageNumber="46">
<emphasis id="A7B2DA33FF9CFFABFF7FF9FF552FF985" bold="true" box="[199,315,1557,1581]" pageId="11" pageNumber="46">Remarks.</emphasis>
. The adult morphology of this species is very similar to that of
<taxonomicName id="52C67DA2FF9CFFABFBA1F9FC50FEF985" box="[1049,1258,1557,1581]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="parvimaculatus">
<emphasis id="A7B2DA33FF9CFFABFBA1F9FC50FEF985" box="[1049,1258,1557,1581]" italics="true" pageId="11" pageNumber="46">C. parvimaculatus</emphasis>
</taxonomicName>
but pupal morphology is markedly different indicating the usefulness of pupal morphology in defining species.
</paragraph>
<paragraph id="95790621FF9CFFABFF7DF9BD5005F974" blockId="11.[151,1436,1279,1792]" pageId="11" pageNumber="46">
Males of this species can be distinguished from those of other members of
<emphasis id="A7B2DA33FF9CFFABFBB4F9BD508CF9C7" box="[1036,1176,1623,1647]" italics="true" pageId="11" pageNumber="46">Marksomyia</emphasis>
by the triangular excavation on the medial part of the parameres. Females can be distinguished by the distribution of antennal SCo. Pupae can be distinguished from those of other species of
<emphasis id="A7B2DA33FF9CFFABFCF7F97557CFF91F" box="[847,987,1695,1719]" italics="true" pageId="11" pageNumber="46">Marksomyia</emphasis>
by the paired, sharp lateral spines on abdominal tubercules
<emphasis id="A7B2DA33FF9CFFABFE36F92E560FF974" box="[398,539,1732,1756]" italics="true" pageId="11" pageNumber="46">dasm, dpmiv</emphasis>
and
<emphasis id="A7B2DA33FF9CFFABFDEAF92E569BF974" box="[594,655,1732,1756]" italics="true" pageId="11" pageNumber="46">dpmv</emphasis>
and thoracic tubercules
<emphasis id="A7B2DA33FF9CFFABFC18F92E57A2F974" box="[928,950,1732,1756]" italics="true" pageId="11" pageNumber="46">di</emphasis>
and
<emphasis id="A7B2DA33FF9CFFABFC54F92E5005F974" box="[1004,1041,1732,1756]" italics="true" pageId="11" pageNumber="46">dii.</emphasis>
</paragraph>
<paragraph id="95790621FF9CFFABFF7DF90D50E3F957" blockId="11.[151,1436,1279,1792]" box="[197,1271,1767,1792]" pageId="11" pageNumber="46">
The wing of this species was illustrated by
<bibRefCitation id="F1577BD0FF9CFFABFD10F902577FF957" box="[680,875,1767,1792]" pageId="11" pageNumber="46" refString="Dyce, A. L., Bellis, G. A. &amp; Muller, M. J. (2007) Pictorial atlas of Australasian Culicoides wings (Diptera: Ceratopogonidae). Australian Biological Resources Study, Canberra, 88 pp." type="book">Dyce et al (2007)</bibRefCitation>
under the name
<taxonomicName id="52C67DA2FF9CFFABFB9EF90D5061F957" box="[1062,1141,1767,1791]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="57" phylum="Arthropoda" rank="species" species="marksi">Marksi</taxonomicName>
gp sp no 1.
</paragraph>
</subSubSection>
</treatment>
</document>
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