464 lines
54 KiB
XML
464 lines
54 KiB
XML
<document id="EBA9B1AAC96124CE86353A7BF3FAD1EC" ID-CLB-Dataset="47633" ID-DOI="10.5281/zenodo.197547" ID-GBIF-Dataset="70628c5e-0412-4a07-bf6e-7993256df56c" ID-ISSN="1175-5326" ID-Zenodo-Dep="197547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460272761230" checkinUser="plazi" docAuthor="Oliver, Paul M., Couper, Patrick & Amey, Andrew" docDate="2010" docId="1D4687A5D843FFEA55A2FC634185AA33" docLanguage="en" docName="zt02578p061.pdf" docOrigin="Zootaxa 2578" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pygopus robertsi Oliver, Couper & Amey, 2010, sp. nov." docType="treatment" docVersion="10" lastPageNumber="57" masterDocId="E17FFFDDD847FFE05535FF914012A814" masterDocTitle="A new species of Pygopus (Pygopodidae; Gekkota; Squamata) from north-eastern Queensland" masterLastPageNumber="61" masterPageNumber="47" pageNumber="51" updateTime="1732568760509" updateUser="ExternalLinkService">
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<mods:mods id="2E50AC39E964A61D9CF1123195844D42" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="10C8931468E77A77CE2C6C64E89C3156">
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<mods:title id="2D6C478733670E202EDDF15F9953740A">A new species of Pygopus (Pygopodidae; Gekkota; Squamata) from north-eastern Queensland</mods:title>
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</mods:titleInfo>
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<mods:name id="E66340A16FE50E8F1A5B3E2F786C3ADB" type="personal">
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<mods:role id="A5D7C15018B987C410DDB561335F9AA0">
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<mods:roleTerm id="8506E636DF29BFEE35556AA4DE379B95">Author</mods:roleTerm>
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<mods:namePart id="CA0C89194B11B61934A96B7D670E483E">Oliver, Paul M.</mods:namePart>
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</mods:name>
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<mods:name id="18E114D5F0E6F668D6735B6844F87863" type="personal">
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<mods:role id="89EB257F5144126D50F259B054323F30">
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<mods:roleTerm id="C5706BDC69DB03F1B6B56EB5FBDB48FF">Author</mods:roleTerm>
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<mods:namePart id="B1BDE612A82D66F05CC8436D67F6D9CE">Couper, Patrick</mods:namePart>
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</mods:name>
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<mods:name id="BF18B6F8A7057347F6BA552EB7109EB0" type="personal">
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<mods:roleTerm id="FD0627A05C4DF9FC6CEEBFAC446A868A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8118C811B940EFBF9957FBDDD23B0933">Amey, Andrew</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="F63C6B719C063FA3E9ED365C226AE277">text</mods:typeOfResource>
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<mods:titleInfo id="93BB184834B45EBE8793269E72E107BD">
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<mods:title id="058901B3303E2F86598D4B80B61862B1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="E235B6E77B87192783A24230568B0C42">
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<mods:date id="245EB9DEDB3942AB9CC55BCD65E85A79">2010</mods:date>
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<mods:detail id="3EE4C2F68007EFEFA89DE46629D60E69" type="volume">
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<mods:number id="47A59CF56BAA99E6B77918B7E1DC2019">2578</mods:number>
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</mods:detail>
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<mods:start id="BA28B20891FACA7EEA25EC1ABD361786">47</mods:start>
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<mods:classification id="01C2F902D1D24E95415047A872A21247">journal article</mods:classification>
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<mods:identifier id="0E1C90A873790F1243582BC4EBC9F822" type="CLB-Dataset">47633</mods:identifier>
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<mods:identifier id="2123261D2204DA920C803E2F63FDC2C6" type="DOI">10.5281/zenodo.197547</mods:identifier>
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<mods:identifier id="D2A7DD66AEDC7C0590E5C49971798F9E" type="GBIF-Dataset">70628c5e-0412-4a07-bf6e-7993256df56c</mods:identifier>
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<mods:identifier id="BFAE5C90934C3D28381650A1E5CD434E" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="8A7D9DDCE50241DE54A02145F79CDF13" type="Zenodo-Dep">197547</mods:identifier>
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<treatment id="1D4687A5D843FFEA55A2FC634185AA33" ID-DOI="http://doi.org/10.5281/zenodo.6196768" ID-GBIF-Taxon="119395437" ID-Zenodo-Dep="6196768" LSID="urn:lsid:plazi:treatment:1D4687A5D843FFEA55A2FC634185AA33" httpUri="http://treatment.plazi.org/id/1D4687A5D843FFEA55A2FC634185AA33" lastPageId="10" lastPageNumber="57" pageId="4" pageNumber="51" scope_class="Squamata" scope_family="Pygopodidae" scope_superFamily="Gekkota">
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<subSubSection id="DDF56538D843FFE455A2FC6341B0AC3B" pageId="4" pageNumber="51" type="nomenclature">
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<paragraph id="955036B3D843FFE455A2FC6341D4AC18" blockId="4.[151,454,1010,1071]" box="[151,454,1010,1036]" pageId="4" pageNumber="51">
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<heading id="CE1881DFD843FFE455A2FC6341D4AC18" bold="true" box="[151,454,1010,1036]" fontSize="11" level="1" pageId="4" pageNumber="51" reason="1">
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<emphasis id="A79BEAA1D843FFE455A2FC6341D4AC18" bold="true" box="[151,454,1010,1036]" pageId="4" pageNumber="51">
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<taxonomicName id="52EF4D30D843FFE455A2FC634173AC18" ID-CoL="4QPGS" box="[151,353,1010,1036]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
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<emphasis id="A79BEAA1D843FFE455A2FC634173AC18" bold="true" box="[151,353,1010,1036]" italics="true" pageId="4" pageNumber="51">Pygopus robertsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BCA857DAD843FFE4545DFC6341D4AC18" box="[360,454,1010,1036]" pageId="4" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="955036B3D843FFE455A2FB8441B0AC3B" blockId="4.[151,454,1010,1071]" box="[151,418,1045,1071]" pageId="4" pageNumber="51">
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<figureCitation id="0DD42A36D843FFE455A2FB8440F5AC3B" box="[151,231,1045,1071]" captionStart="FIGURE 1" captionStartId="5.[151,255,1879,1903]" captionTargetBox="[214,1337,192,1749]" captionTargetId="figure@5.[214,1337,171,1870]" captionTargetPageId="5" captionText="FIGURE 1. (A) Dorsal view of the head of Pygopus robertsi sp. nov. (holotype, QM J 47145). (B) Lateral view of the head of P. ro b e r t s i sp. nov. (holotype, QM J 47145). CF = caudal frontal, CS = Caudal supranasal, FN = frontonasals, IL = infralabial, L = loreals, M = mental, P = parietals, PF = prefrontal, PN = post nasal, PO = preoculars, PT = pretemporal, R = rostral, RF = rostral frontal, RS = rostral supranasals, SC = supraciliaries, SL = supralabials, SO = supraoculars, SUB = suboculars, T = upper temporals." httpUri="https://zenodo.org/record/197548/files/figure.png" pageId="4" pageNumber="51">Figs. 1</figureCitation>
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,
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<figureCitation id="0DD42A36D843FFE455C0FB844116AC3B" box="[245,260,1045,1071]" captionStart="FIGURE 2" captionStartId="6.[151,259,1474,1498]" captionTargetBox="[216,1367,188,1461]" captionTargetId="figure@6.[211,1376,182,1466]" captionTargetPageId="6" captionText="FIGURE 2. Upper row: degree of keeling on dorsal body scales. A) weak on body scales of P. robertsi sp. nov. (holotype, QM J 47145). B) strong on body scales of P. lepidopodus (QM J 47934) from south-eastern Queensland. Middle row: Scalation of supraciliary / supraocular junction. C) supraciliaries of P. robertsi sp. nov. (holotype, QM J 47145) in a single, continuous row. D) supraciliaries of P. lepidopodus (QM J 47934) in two rows along the posterior portion of the orbit. Lower row: Condition of scales on temporal region. E) smooth for P. robertsi sp. nov. (holotype, QM J 47145). F) weakly keeled for P. lepidopodus (QM J 47934) from south-eastern Queensland." httpUri="https://zenodo.org/record/197549/files/figure.png" pageId="4" pageNumber="51">2</figureCitation>
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A,C,E, 4A,B.
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</paragraph>
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</subSubSection>
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<subSubSection id="DDF56538D843FFE455A2FBCE4532AE4B" pageId="4" pageNumber="51" type="materials_examined">
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<paragraph id="955036B3D843FFE455A2FBCE4265AC8B" blockId="4.[151,1437,1119,1967]" pageId="4" pageNumber="51">
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<emphasis id="A79BEAA1D843FFE455A2FBCE4103AC6D" bold="true" box="[151,273,1119,1145]" pageId="4" pageNumber="51">
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<typeStatus id="4A548811D843FFE455A2FBCE411EAC6D" box="[151,268,1119,1145]" pageId="4" pageNumber="51" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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QM
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<accessionNumber id="8ABCAB50D843FFE45464FBF141BBAC6E" box="[337,425,1120,1146]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="4" pageNumber="51" type="EnaNcbi">J47145</accessionNumber>
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, male, Little Forks, Shipton's Flat, via Cooktown,
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<geoCoordinate id="F0DB5074D843FFE45137FBF14463AC6E" box="[1026,1137,1120,1146]" direction="south" orientation="latitude" pageId="4" pageNumber="51" precision="925" value="-15.783334">15° 47' S</geoCoordinate>
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,
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<geoCoordinate id="F0DB5074D843FFE4514AFBF144EDAC6E" box="[1151,1279,1120,1146]" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="925" value="145.25">145° 15' E</geoCoordinate>
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, collected by Lewis Roberts (registered
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<date id="E1511073D843FFE454F8FB14427DAC8B" box="[461,623,1157,1183]" pageId="4" pageNumber="51" value="1987-08-28">28 Aug. 1987</date>
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)
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</paragraph>
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<paragraph id="955036B3D843FFE455F0FB3B40E6AD1B" blockId="4.[151,1437,1119,1967]" pageId="4" pageNumber="51">
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<emphasis id="A79BEAA1D843FFE455F0FB3B4158ACD0" bold="true" box="[197,330,1194,1220]" pageId="4" pageNumber="51">
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<typeStatus id="4A548811D843FFE455F0FB3B4157ACD0" box="[197,325,1194,1220]" pageId="4" pageNumber="51" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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QM J3899, sex not determined, Yungaburra,
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<geoCoordinate id="F0DB5074D843FFE45657FB3B43E2ACD0" box="[866,1008,1194,1220]" direction="south" orientation="latitude" pageId="4" pageNumber="51" precision="15" value="-17.266666">17° 16' 0" S</geoCoordinate>
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,
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<geoCoordinate id="F0DB5074D843FFE456C8FB3B448CACD0" box="[1021,1182,1194,1220]" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="15" value="145.58333">145° 35' 0" E</geoCoordinate>
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; QM
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<accessionNumber id="8ABCAB50D843FFE451D4FB3B452AACD0" box="[1249,1336,1194,1220]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J57778" pageId="4" pageNumber="51" type="EnaNcbi">J57778</accessionNumber>
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, female, Heathlands area,
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<geoCoordinate id="F0DB5074D843FFE45454FB4141FCACFE" box="[353,494,1232,1258]" direction="south" orientation="latitude" pageId="4" pageNumber="51" precision="15" value="-11.733334">11° 44' 0" S</geoCoordinate>
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,
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<geoCoordinate id="F0DB5074D843FFE454CEFB414289ACFE" box="[507,667,1232,1258]" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="15" value="142.53334">142° 32' 0" E</geoCoordinate>
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; QM
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<accessionNumber id="8ABCAB50D843FFE457EBFB414327ACFE" box="[734,821,1232,1258]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J58986" pageId="4" pageNumber="51" type="EnaNcbi">J58986</accessionNumber>
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, male,
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<quantity id="52179B56D843FFE456BCFB4143F0ACFE" box="[905,994,1232,1258]" metricMagnitude="4" metricUnit="m" metricValue="1.05" pageId="4" pageNumber="51" unit="km" value="10.5">10.5km</quantity>
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NE of Heathlands,
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<geoCoordinate id="F0DB5074D843FFE451FCFB414544ACFE" box="[1225,1366,1232,1258]" direction="south" orientation="latitude" pageId="4" pageNumber="51" precision="15" value="-11.683333">11° 41' 0" S</geoCoordinate>
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,
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<geoCoordinate id="F0DB5074D843FFE45051FB4140E2AD1B" direction="east" orientation="longitude" pageId="4" pageNumber="51" precision="15" value="142.65">142° 39' 0"E</geoCoordinate>
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.
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</paragraph>
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<paragraph id="955036B3D843FFE455F0FA8B4190ADB0" blockId="4.[151,1437,1119,1967]" pageId="4" pageNumber="51">
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<emphasis id="A79BEAA1D843FFE455F0FA8B41B9AD20" bold="true" box="[197,427,1306,1332]" pageId="4" pageNumber="51">Generic diagnosis.</emphasis>
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A species in the genus
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<taxonomicName id="52EF4D30D843FFE457F4FA8A4334AD20" box="[705,806,1307,1332]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="51" phylum="Chordata" rank="genus">
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<emphasis id="A79BEAA1D843FFE457F4FA8A4334AD20" box="[705,806,1307,1332]" italics="true" pageId="4" pageNumber="51">Pygopus</emphasis>
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</taxonomicName>
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as defined by the following combination of character states: dorsal surface of head covered with large and small scales, 3 to 5 postmental scales, 21 or more midbody scale rows, dorsal body scales usually keeled and adults of both sexes with 9 or more precloacal pores (Kluge 1974).
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</paragraph>
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<paragraph id="955036B3D843FFE455F0FA3E4532AE4B" blockId="4.[151,1437,1119,1967]" pageId="4" pageNumber="51">
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<emphasis id="A79BEAA1D843FFE455F0FA3E41BCADDD" bold="true" box="[197,430,1455,1481]" pageId="4" pageNumber="51">Specific diagnosis.</emphasis>
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<taxonomicName id="52EF4D30D843FFE45483FA214291ADDD" box="[438,643,1456,1481]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="51" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
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<emphasis id="A79BEAA1D843FFE45483FA214291ADDD" box="[438,643,1456,1481]" italics="true" pageId="4" pageNumber="51">Pygopus robertsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="BCA857DAD843FFE457BEFA3E42F9ADDE" box="[651,747,1455,1482]" pageId="4" pageNumber="51" rank="species">
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<emphasis id="A79BEAA1D843FFE457BEFA3E42F1ADDD" bold="true" box="[651,739,1455,1481]" pageId="4" pageNumber="51">sp. nov</emphasis>
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.
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</taxonomicNameLabel>
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is a large, robust pygopodid (maximum SVL ~
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<quantity id="52179B56D843FFE4501AFA214586ADDE" box="[1327,1428,1456,1482]" metricMagnitude="-1" metricUnit="m" metricValue="2.2399999999999998" pageId="4" pageNumber="51" unit="mm" value="224.0">224 mm</quantity>
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) with a rounded snout, precloacal pores> 10, ventrals <100, midbody scales in 21 rows, no dark collar on the head or neck, no dark patch beneath the nostril, dark pigmentation around the margins of the ventral scales (at least posteriorly), approximately nine rows of weakly keeled dorsal scales and supraciliaries in a single, continuous row. Further specific morphological comparisons are given in the results section above.
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</paragraph>
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</subSubSection>
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<subSubSection id="DDF56538D843FFE755F0F9FB4416AFFF" lastPageId="7" lastPageNumber="54" pageId="4" pageNumber="51" type="description">
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<paragraph id="955036B3D843FFE455F0F9FB420FAEE0" blockId="4.[151,1437,1119,1967]" pageId="4" pageNumber="51">
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<emphasis id="A79BEAA1D843FFE455F0F9FB4179AE90" bold="true" box="[197,363,1642,1668]" pageId="4" pageNumber="51">Description.</emphasis>
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Sample size = 4 (unless otherwise stated). SVL =
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<quantity id="52179B56D843FFE456C8F9FB44B7AE90" box="[1021,1189,1642,1668]" metricMagnitude="-1" metricUnit="m" metricValue="1.73" metricValueMax="2.2399999999999998" metricValueMin="1.22" pageId="4" pageNumber="51" unit="mm" value="173.0" valueMax="224.0" valueMin="122.0">122–224 mm</quantity>
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(mean =
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<quantity id="52179B56D843FFE45014F9FB4583AE90" box="[1313,1425,1642,1668]" metricMagnitude="-1" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="51" unit="mm" value="187.0">187 mm</quantity>
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). Proportions as %SVL: Tail (no original tails in sample but one tail with a regrown tip)> 176; HL = 7.4–9.5 (mean = 8.1); HW = 4.1–6.0 (n = 3, mean = 5.2); S = 3.1–4.2 (mean = 3.4); EE = 3.4–4.0 (mean = 3.6); hindlimb = 3.4–5.3 (mean = 4.7).
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</paragraph>
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<paragraph id="955036B3D843FFE655F3F8914420AFF8" blockId="4.[151,1437,1119,1967]" lastBlockId="6.[151,1436,1704,2028]" lastPageId="6" lastPageNumber="53" pageId="4" pageNumber="51">
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<emphasis id="A79BEAA1D843FFE455F3F891411CAF0D" box="[198,270,1792,1817]" italics="true" pageId="4" pageNumber="51">Head.</emphasis>
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Bullet-shaped with a rounded snout; rostral barely projecting between rostral supranasals; rostral supranasals in narrow to point contact with the first supralabial, anterior to the nostril; caudal supranasals present, in narrow contact with dorsal edge of nostril; frontonasal divided; rostral frontal and caudal frontal in broad contact; parietals in broad contact behind caudal frontal and bordered posteriorly by a small, centrally placed occipital scale (in three specimens – not present in
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<typeStatus id="4A548811D843FFE45675F80443BEAFBB" box="[832,940,1941,1967]" pageId="4" pageNumber="51" type="holotype">holotype</typeStatus>
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); a single postnasal in broad contact with first supralabial; loreals, left side 10–12 (mean = 11.25, mode = 12), right side 8–12 (mean = 10.5, mode = 11), heterogenous; preoculars, left side 8–13 (mean = 11.25, mode = 12), right side 9–14 (mean = 11, mode = 9); suboculars 3; supraoculars 2, first longest; supraciliaries in a single row (
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<figureCitation id="0DD42A36D841FFE6511DF963447BAF18" box="[1064,1129,1778,1804]" captionStart="FIGURE 2" captionStartId="6.[151,259,1474,1498]" captionTargetBox="[216,1367,188,1461]" captionTargetId="figure@6.[211,1376,182,1466]" captionTargetPageId="6" captionText="FIGURE 2. Upper row: degree of keeling on dorsal body scales. A) weak on body scales of P. robertsi sp. nov. (holotype, QM J 47145). B) strong on body scales of P. lepidopodus (QM J 47934) from south-eastern Queensland. Middle row: Scalation of supraciliary / supraocular junction. C) supraciliaries of P. robertsi sp. nov. (holotype, QM J 47145) in a single, continuous row. D) supraciliaries of P. lepidopodus (QM J 47934) in two rows along the posterior portion of the orbit. Lower row: Condition of scales on temporal region. E) smooth for P. robertsi sp. nov. (holotype, QM J 47145). F) weakly keeled for P. lepidopodus (QM J 47934) from south-eastern Queensland." httpUri="https://zenodo.org/record/197549/files/figure.png" pageId="6" pageNumber="53">Fig 2</figureCitation>
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C, thus only 5 interorbital scales present), left side 3–4 (mean = 3.75, mode = 4), right side 4, usually
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<quantity id="52179B56D841FFE6513CF8894424AF26" box="[1033,1078,1816,1842]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="6" pageNumber="53" unit="in" value="2.0">2 in</quantity>
|
||
contact with first supraocular; supralabials, left side 7, right side 7 or 8 (mean = 7.25, mode = 7), fourth below centre of eye; infralabials 6 or 7 (mean = 6.75, mode = 7), first infralabials well separated by a single scale; second infralabials separated by 3–4 scale rows (mean = 3.25, mode = 3); third infralabials separated by 5–8 scales rows (mean = 6.75, mode = 7); pretemporal single; upper temporals, left side 1 or 2 (mean = 1.75, mode = 2), right side 2; nuchal scales 16–20 (mean = 17.5, mode 16); gular scales 16–18 (mean = 17.3, mode = 18).
|
||
</paragraph>
|
||
<caption id="C190663BD842FFE555A2F8C642CEAFFA" httpUri="https://zenodo.org/record/197548/files/figure.png" pageId="5" pageNumber="52" targetBox="[214,1337,192,1749]" targetPageId="5">
|
||
<paragraph id="955036B3D842FFE555A2F8C642CEAFFA" blockId="5.[151,1437,1879,2031]" pageId="5" pageNumber="52">
|
||
<emphasis id="A79BEAA1D842FFE555A2F8C64101AF7B" bold="true" box="[151,275,1879,1903]" pageId="5" pageNumber="52">FIGURE 1</emphasis>
|
||
. (A) Dorsal view of the head of
|
||
<taxonomicName id="52EF4D30D842FFE5574DF8C64322AF7A" box="[632,816,1879,1902]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="52" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D842FFE5574DF8C64322AF7A" box="[632,816,1879,1902]" italics="true" pageId="5" pageNumber="52">Pygopus robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD842FFE5560DF8C6439DAF7B" box="[824,911,1879,1903]" pageId="5" pageNumber="52" rank="species">
|
||
<emphasis id="A79BEAA1D842FFE5560DF8C6439DAF7B" bold="true" box="[824,911,1879,1903]" pageId="5" pageNumber="52">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype, QM
|
||
<accessionNumber id="8ABCAB50D842FFE5510BF8C64482AF7A" box="[1086,1168,1879,1902]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="5" pageNumber="52" type="EnaNcbi">J47145</accessionNumber>
|
||
). (B) Lateral view of the head of
|
||
<emphasis id="A79BEAA1D842FFE555D9F8E64144AF9A" box="[236,342,1911,1934]" italics="true" pageId="5" pageNumber="52">P. ro b e r t s i</emphasis>
|
||
<emphasis id="A79BEAA1D842FFE55455F8E641A7AF9B" bold="true" box="[352,437,1911,1935]" pageId="5" pageNumber="52">sp. nov.</emphasis>
|
||
(holotype, QM
|
||
<accessionNumber id="8ABCAB50D842FFE55755F8E642A0AF9A" box="[608,690,1911,1934]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="5" pageNumber="52" type="EnaNcbi">J47145</accessionNumber>
|
||
).
|
||
<emphasis id="A79BEAA1D842FFE557F6F8E642F4AF9B" bold="true" box="[707,742,1911,1935]" pageId="5" pageNumber="52">CF</emphasis>
|
||
= caudal frontal,
|
||
<emphasis id="A79BEAA1D842FFE55696F8E643D7AF9B" bold="true" box="[931,965,1911,1935]" pageId="5" pageNumber="52">CS</emphasis>
|
||
= Caudal supranasal,
|
||
<emphasis id="A79BEAA1D842FFE55184F8E644C7AF9B" bold="true" box="[1201,1237,1911,1935]" pageId="5" pageNumber="52">FN</emphasis>
|
||
= frontonasals,
|
||
<emphasis id="A79BEAA1D842FFE550B5F8E6458FAF9B" bold="true" box="[1408,1437,1911,1935]" pageId="5" pageNumber="52">IL</emphasis>
|
||
= infralabial,
|
||
<emphasis id="A79BEAA1D842FFE55403F806415AAFBB" bold="true" box="[310,328,1943,1967]" pageId="5" pageNumber="52">L</emphasis>
|
||
= loreals,
|
||
<emphasis id="A79BEAA1D842FFE554F2F80641F2AFBB" bold="true" box="[455,480,1943,1967]" pageId="5" pageNumber="52">M</emphasis>
|
||
= mental,
|
||
<emphasis id="A79BEAA1D842FFE5576AF806427DAFBB" bold="true" box="[607,623,1943,1967]" pageId="5" pageNumber="52">P</emphasis>
|
||
= parietals,
|
||
<emphasis id="A79BEAA1D842FFE55636F8064334AFBB" bold="true" box="[771,806,1943,1967]" pageId="5" pageNumber="52">PF</emphasis>
|
||
= prefrontal,
|
||
<emphasis id="A79BEAA1D842FFE556FDF80643FCAFBB" bold="true" box="[968,1006,1943,1967]" pageId="5" pageNumber="52">PN</emphasis>
|
||
= post nasal,
|
||
<emphasis id="A79BEAA1D842FFE551A4F80644AAAFBB" bold="true" box="[1169,1208,1943,1967]" pageId="5" pageNumber="52">PO</emphasis>
|
||
= preoculars,
|
||
<emphasis id="A79BEAA1D842FFE55054F8064597AFBB" bold="true" box="[1377,1413,1943,1967]" pageId="5" pageNumber="52">PT</emphasis>
|
||
= pretemporal,
|
||
<emphasis id="A79BEAA1D842FFE5541CF826412EAFDB" bold="true" box="[297,316,1975,1999]" pageId="5" pageNumber="52">R</emphasis>
|
||
= rostral,
|
||
<emphasis id="A79BEAA1D842FFE5549AF82641C1AFDB" bold="true" box="[431,467,1975,1999]" pageId="5" pageNumber="52">RF</emphasis>
|
||
= rostral frontal,
|
||
<emphasis id="A79BEAA1D842FFE557A3F82642ABAFDB" bold="true" box="[662,697,1975,1999]" pageId="5" pageNumber="52">RS</emphasis>
|
||
= rostral supranasals,
|
||
<emphasis id="A79BEAA1D842FFE55685F82643C1AFDB" bold="true" box="[944,979,1975,1999]" pageId="5" pageNumber="52">SC</emphasis>
|
||
= supraciliaries,
|
||
<emphasis id="A79BEAA1D842FFE551A4F82644A0AFDB" bold="true" box="[1169,1202,1975,1999]" pageId="5" pageNumber="52">SL</emphasis>
|
||
= supralabials,
|
||
<emphasis id="A79BEAA1D842FFE55054F8264597AFDB" bold="true" box="[1377,1413,1975,1999]" pageId="5" pageNumber="52">SO</emphasis>
|
||
= supraoculars,
|
||
<emphasis id="A79BEAA1D842FFE5541EF846414DAFFB" bold="true" box="[299,351,2007,2031]" pageId="5" pageNumber="52">SUB</emphasis>
|
||
= suboculars,
|
||
<emphasis id="A79BEAA1D842FFE554CFF846421EAFFB" bold="true" box="[506,524,2007,2031]" pageId="5" pageNumber="52">T</emphasis>
|
||
= upper temporals.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C190663BD841FFE655A2FA53447FAE6D" httpUri="https://zenodo.org/record/197549/files/figure.png" pageId="6" pageNumber="53" targetBox="[216,1367,188,1461]" targetPageId="6">
|
||
<paragraph id="955036B3D841FFE655A2FA53447FAE6D" blockId="6.[151,1437,1474,1657]" pageId="6" pageNumber="53">
|
||
<emphasis id="A79BEAA1D841FFE655A2FA534133ADCE" bold="true" box="[151,289,1474,1498]" pageId="6" pageNumber="53">FIGURE 2.</emphasis>
|
||
Upper row: degree of keeling on dorsal body scales. A) weak on body scales of
|
||
<taxonomicName id="52EF4D30D841FFE651F7FA53452AADCD" box="[1218,1336,1474,1497]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D841FFE651F7FA53452AADCD" box="[1218,1336,1474,1497]" italics="true" pageId="6" pageNumber="53">P. robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD841FFE65074FA53458EADCE" box="[1345,1436,1474,1498]" pageId="6" pageNumber="53" rank="species">
|
||
<emphasis id="A79BEAA1D841FFE65074FA53458EADCE" bold="true" box="[1345,1436,1474,1498]" pageId="6" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype, QM
|
||
<accessionNumber id="8ABCAB50D841FFE65470FA734189ADED" box="[325,411,1506,1529]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="6" pageNumber="53" type="EnaNcbi">J47145</accessionNumber>
|
||
). B) strong on body scales of
|
||
<taxonomicName id="52EF4D30D841FFE657D9FA734382ADED" box="[748,912,1506,1529]" class="Reptilia" family="Bipedidae" genus="Bipes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="lepidopodus">
|
||
<emphasis id="A79BEAA1D841FFE657D9FA734382ADED" box="[748,912,1506,1529]" italics="true" pageId="6" pageNumber="53">P. lepidopodus</emphasis>
|
||
</taxonomicName>
|
||
(QM
|
||
<accessionNumber id="8ABCAB50D841FFE656E0FA73443AADED" box="[981,1064,1506,1529]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47934" pageId="6" pageNumber="53" type="EnaNcbi">J47934</accessionNumber>
|
||
) from south-eastern Queensland. Middle row: Scalation of supraciliary/supraocular junction. C) supraciliaries of
|
||
<taxonomicName id="52EF4D30D841FFE6512CF993449FAE0D" box="[1049,1165,1538,1561]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D841FFE6512CF993449FAE0D" box="[1049,1165,1538,1561]" italics="true" pageId="6" pageNumber="53">P. robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD841FFE651A1F99344FCAE0E" box="[1172,1262,1538,1562]" pageId="6" pageNumber="53" rank="species">
|
||
<emphasis id="A79BEAA1D841FFE651A1F99344FCAE0E" bold="true" box="[1172,1262,1538,1562]" pageId="6" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype, QM
|
||
<accessionNumber id="8ABCAB50D841FFE655A2F9B340F5AE2D" box="[151,231,1570,1593]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="6" pageNumber="53" type="EnaNcbi">J47145</accessionNumber>
|
||
) in a single, continuous row. D) supraciliaries of
|
||
<taxonomicName id="52EF4D30D841FFE65631F9B343B6AE2D" box="[772,932,1570,1593]" class="Reptilia" family="Bipedidae" genus="Bipes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="lepidopodus">
|
||
<emphasis id="A79BEAA1D841FFE65631F9B343B6AE2D" box="[772,932,1570,1593]" italics="true" pageId="6" pageNumber="53">P. lepidopodus</emphasis>
|
||
</taxonomicName>
|
||
(QM
|
||
<accessionNumber id="8ABCAB50D841FFE656DDF9B3442AAE2D" box="[1000,1080,1570,1593]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47934" pageId="6" pageNumber="53" type="EnaNcbi">J47934</accessionNumber>
|
||
) in two rows along the posterior portion of the orbit. Lower row: Condition of scales on temporal region. E) smooth for
|
||
<taxonomicName id="52EF4D30D841FFE65105F9D3448CAE4D" box="[1072,1182,1602,1625]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D841FFE65105F9D3448CAE4D" box="[1072,1182,1602,1625]" italics="true" pageId="6" pageNumber="53">P. robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD841FFE65196F9D344EAAE4E" box="[1187,1272,1602,1626]" pageId="6" pageNumber="53" rank="species">
|
||
<emphasis id="A79BEAA1D841FFE65196F9D344EAAE4E" bold="true" box="[1187,1272,1602,1626]" pageId="6" pageNumber="53">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(holotype, QM
|
||
<accessionNumber id="8ABCAB50D841FFE655A2F9F340FAAE6D" box="[151,232,1634,1657]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="6" pageNumber="53" type="EnaNcbi">J47145</accessionNumber>
|
||
). F) weakly keeled for
|
||
<taxonomicName id="52EF4D30D841FFE654EBF9F34269AE6D" box="[478,635,1634,1657]" class="Reptilia" family="Bipedidae" genus="Bipes" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="53" phylum="Chordata" rank="species" species="lepidopodus">
|
||
<emphasis id="A79BEAA1D841FFE654EBF9F34269AE6D" box="[478,635,1634,1657]" italics="true" pageId="6" pageNumber="53">P. lepidopodus</emphasis>
|
||
</taxonomicName>
|
||
(QM
|
||
<accessionNumber id="8ABCAB50D841FFE65789F9F34319AE6D" box="[700,779,1634,1657]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47934" pageId="6" pageNumber="53" type="EnaNcbi">J47934</accessionNumber>
|
||
) from south-eastern Queensland.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="C190663BD840FFE755A2FB5A4377AD17" httpUri="https://zenodo.org/record/197550/files/figure.png" pageId="7" pageNumber="54" targetBox="[192,1396,180,1215]" targetPageId="7">
|
||
<paragraph id="955036B3D840FFE755A2FB5A4377AD17" blockId="7.[151,1436,1227,1283]" pageId="7" pageNumber="54">
|
||
<emphasis id="A79BEAA1D840FFE755A2FB5A410BACF7" bold="true" box="[151,281,1227,1251]" pageId="7" pageNumber="54">FIGURE 3.</emphasis>
|
||
Consensus topology of relationships amongst
|
||
<taxonomicName id="52EF4D30D840FFE7563AFB5D4379ACF7" box="[783,875,1228,1251]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="A79BEAA1D840FFE7563AFB5D4379ACF7" box="[783,875,1228,1251]" italics="true" pageId="7" pageNumber="54">Pygopus</emphasis>
|
||
</taxonomicName>
|
||
specimens sampled in this study with ML bootstrap supports and Bayesian Posterior probabilities shown for key nodes.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="955036B3D840FFE755F3FAA04508AD81" blockId="7.[151,1437,1329,2027]" pageId="7" pageNumber="54">
|
||
<emphasis id="A79BEAA1D840FFE755F3FAA0411BAD5E" box="[198,265,1329,1354]" italics="true" pageId="7" pageNumber="54">Body.</emphasis>
|
||
Weak to very weak unicarinate keeling (
|
||
<figureCitation id="0DD42A36D840FFE757D6FAA0433AAD5F" box="[739,808,1329,1355]" captionStart="FIGURE 2" captionStartId="6.[151,259,1474,1498]" captionTargetBox="[216,1367,188,1461]" captionTargetId="figure@6.[211,1376,182,1466]" captionTargetPageId="6" captionText="FIGURE 2. Upper row: degree of keeling on dorsal body scales. A) weak on body scales of P. robertsi sp. nov. (holotype, QM J 47145). B) strong on body scales of P. lepidopodus (QM J 47934) from south-eastern Queensland. Middle row: Scalation of supraciliary / supraocular junction. C) supraciliaries of P. robertsi sp. nov. (holotype, QM J 47145) in a single, continuous row. D) supraciliaries of P. lepidopodus (QM J 47934) in two rows along the posterior portion of the orbit. Lower row: Condition of scales on temporal region. E) smooth for P. robertsi sp. nov. (holotype, QM J 47145). F) weakly keeled for P. lepidopodus (QM J 47934) from south-eastern Queensland." httpUri="https://zenodo.org/record/197549/files/figure.png" pageId="7" pageNumber="54">Fig. 2</figureCitation>
|
||
A) extending over 9 dorsal scale rows, weakest on the outermost row; midbody scale rows 21; ventral scales 93–96 (mean = 93.8, mode = 93); ventral body scales paired, transversely enlarged (BBS 71-76, mean = 73.25); precloacal pores 14; precloacal scales 3.
|
||
</paragraph>
|
||
<paragraph id="955036B3D840FFE755F3FA3044B1ADAF" blockId="7.[151,1437,1329,2027]" box="[198,1187,1441,1467]" pageId="7" pageNumber="54">
|
||
<emphasis id="A79BEAA1D840FFE755F3FA30412EADAE" box="[198,316,1441,1466]" italics="true" pageId="7" pageNumber="54">Hindlimb.</emphasis>
|
||
5–6 scales present on ventral margin of hindlimb (mean = 5.8, mode = 6).
|
||
</paragraph>
|
||
<paragraph id="955036B3D840FFE755F3FA56415EAEF1" blockId="7.[151,1437,1329,2027]" pageId="7" pageNumber="54">
|
||
<emphasis id="A79BEAA1D840FFE755F3FA564205ADF4" box="[198,535,1479,1504]" italics="true" pageId="7" pageNumber="54">Colouration in preservative.</emphasis>
|
||
The dorsal surface of the body and tail is light to mid-grey and unmarked. The lower flanks are heavily patterned with dark stippling and larger white spots. The ventral scales are white to cream with a faint grey edging; this increases in intensity, becoming dark grey to black on the posterior third to half of the body. Some specimens have a series of faint grey chin straps and there is a dark, inconspicuous to prominent bar running vertically below the orbit, through the fourth supralabial, and extending to the infralabials or 1 to 2 scale rows beneath the infralabials. Photographs of
|
||
<taxonomicName id="52EF4D30D840FFE75196F9104509AE8E" box="[1187,1307,1665,1690]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="54" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D840FFE75196F9104509AE8E" box="[1187,1307,1665,1690]" italics="true" pageId="7" pageNumber="54">P. robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD840FFE75014F911456CAE8E" box="[1313,1406,1664,1690]" pageId="7" pageNumber="54" rank="species">
|
||
<emphasis id="A79BEAA1D840FFE75014F911456CAE8E" bold="true" box="[1313,1406,1664,1690]" pageId="7" pageNumber="54">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in life (
|
||
<figureCitation id="0DD42A36D840FFE755E5F937410BAED4" box="[208,281,1702,1728]" captionStart="FIGURE 4. A" captionStartId="8.[151,255,1906,1930]" captionTargetBox="[154,1436,223,1867]" captionTargetId="figure@8.[151,1436,194,1885]" captionTargetPageId="8" captionText="FIGURE 4. A) Holotype of P. robertsi sp. nov., QM J 47145, in life (photograph by Steve Wilson). B) Pygopus robertsi sp. nov. in life, non-vouchered specimen from the lower slopes of Mt Sampson 15 ° 48 ' S, 145 ° 11 ' E (photograph by Lewis Roberts)." httpUri="https://zenodo.org/record/197551/files/figure.png" pageId="7" pageNumber="54">Fig. 4</figureCitation>
|
||
) show a pinkish to rufous tinge to the dorsum and scattered pink scales on the lower flanks and ventral surface.
|
||
</paragraph>
|
||
<paragraph id="955036B3D840FFE755F0F9614416AFFF" blockId="7.[151,1437,1329,2027]" pageId="7" pageNumber="54">
|
||
<emphasis id="A79BEAA1D840FFE755F0F9614235AF1E" bold="true" box="[197,551,1776,1802]" pageId="7" pageNumber="54">
|
||
Summary data for
|
||
<typeStatus id="4A548811D840FFE75481F9614230AF1E" box="[436,546,1776,1802]" pageId="7" pageNumber="54" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(QM
|
||
<accessionNumber id="8ABCAB50D840FFE7575AF96042D7AF1F" box="[623,709,1777,1803]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="7" pageNumber="54" type="EnaNcbi">J47145</accessionNumber>
|
||
): SVL =
|
||
<quantity id="52179B56D840FFE75600F9604388AF1F" box="[821,922,1777,1803]" metricMagnitude="-1" metricUnit="m" metricValue="1.86" pageId="7" pageNumber="54" unit="mm" value="186.0">186 mm</quantity>
|
||
; Tail =
|
||
<quantity id="52179B56D840FFE756C6F9604445AF1F" box="[1011,1111,1777,1803]" metricMagnitude="-1" metricUnit="m" metricValue="3.27" pageId="7" pageNumber="54" unit="mm" value="327.0">327 mm</quantity>
|
||
, with regrown tip (
|
||
<quantity id="52179B56D840FFE75003F960459CAF1F" box="[1334,1422,1777,1803]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="7" pageNumber="54" unit="mm" value="36.0">36 mm</quantity>
|
||
); HL =
|
||
<quantity id="52179B56D840FFE755E8F887415BAF24" box="[221,329,1814,1840]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="7" pageNumber="54" unit="mm" value="14.7">14.7 mm</quantity>
|
||
; HW =
|
||
<quantity id="52179B56D840FFE75493F8874203AF24" box="[422,529,1814,1840]" metricMagnitude="-3" metricUnit="m" metricValue="9.64" pageId="7" pageNumber="54" unit="mm" value="9.64">9.64 mm</quantity>
|
||
; S =
|
||
<quantity id="52179B56D840FFE75778F88742ABAF24" box="[589,697,1814,1840]" metricMagnitude="-3" metricUnit="m" metricValue="6.22" pageId="7" pageNumber="54" unit="mm" value="6.22">6.22 mm</quantity>
|
||
; EE =
|
||
<quantity id="52179B56D840FFE7563DF8874366AF24" box="[776,884,1814,1840]" metricMagnitude="-3" metricUnit="m" metricValue="6.27" pageId="7" pageNumber="54" unit="mm" value="6.27">6.27 mm</quantity>
|
||
; hindlimb =
|
||
<quantity id="52179B56D840FFE7513FF8874464AF24" box="[1034,1142,1814,1840]" metricMagnitude="-3" metricUnit="m" metricValue="9.84" pageId="7" pageNumber="54" unit="mm" value="9.84">9.84 mm</quantity>
|
||
. Loreals 11; preoculars 8 (L) or 9 (R); suboculars 3; supraciliaries 3 (L) or 4 (R); supraoculars 2; supralabials 7, fourth below eye; infralabials 7 (L) or 6 (R); single scale row between first infralabials; 4 scale rows between second infralabials; 7 scale rows between third infralabials; pretemporal single; temporals 1 (L) or 2 (R); nuchals 20; gulars 18; midbody scale rows 21; ventrals 93; BBS 72; precloacals 3; very weak keels present on 9 dorsal scale rows; precloacal pores 14; 6 scales along ventral margin of hindlimb.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="C190663BD84FFFE855A2F8E34150AFDE" httpUri="https://zenodo.org/record/197551/files/figure.png" pageId="8" pageNumber="55" targetBox="[154,1436,223,1867]" targetPageId="8">
|
||
<paragraph id="955036B3D84FFFE855A2F8E34150AFDE" blockId="8.[151,1436,1906,1994]" pageId="8" pageNumber="55">
|
||
<emphasis id="A79BEAA1D84FFFE855A2F8E34100AF9E" bold="true" box="[151,274,1906,1930]" pageId="8" pageNumber="55">FIGURE 4</emphasis>
|
||
. A) Holotype of
|
||
<taxonomicName id="52EF4D30D84FFFE854FEF8E2422BAF9E" box="[459,569,1907,1930]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="55" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84FFFE854FEF8E2422BAF9E" box="[459,569,1907,1930]" italics="true" pageId="8" pageNumber="55">P. robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD84FFFE8570AF8E34284AF9E" box="[575,662,1906,1930]" pageId="8" pageNumber="55" rank="species">
|
||
<emphasis id="A79BEAA1D84FFFE8570AF8E34284AF9E" bold="true" box="[575,662,1906,1930]" pageId="8" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,QM
|
||
<accessionNumber id="8ABCAB50D84FFFE857E0F8E24336AF9E" box="[725,804,1907,1930]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="8" pageNumber="55" type="EnaNcbi">J47145</accessionNumber>
|
||
, in life (photograph by Steve Wilson). B)
|
||
<taxonomicName id="52EF4D30D84FFFE851D3F8E2458EAF9E" box="[1254,1436,1907,1930]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="55" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84FFFE851D3F8E2458EAF9E" box="[1254,1436,1907,1930]" italics="true" pageId="8" pageNumber="55">Pygopus robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD84FFFE855A2F80340FDAFBE" box="[151,239,1938,1962]" pageId="8" pageNumber="55" rank="species">
|
||
<emphasis id="A79BEAA1D84FFFE855A2F80340FDAFBE" bold="true" box="[151,239,1938,1962]" pageId="8" pageNumber="55">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in life, non-vouchered specimen from the lower slopes of Mt Sampson 15° 48' S, 145° 11' E (photograph by Lewis Roberts).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="DDF56538D84EFFEA55F0FF064498A986" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56" type="distribution">
|
||
<paragraph id="955036B3D84EFFE955F0FF0641DEA9C8" blockId="9.[151,1436,151,476]" pageId="9" pageNumber="56">
|
||
<emphasis id="A79BEAA1D84EFFE955F0FF064175A8A5" bold="true" box="[197,359,151,177]" pageId="9" pageNumber="56">Distribution.</emphasis>
|
||
<taxonomicName id="52EF4D30D84EFFE9545AFF094228A8A5" box="[367,570,152,177]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84EFFE9545AFF094228A8A5" box="[367,570,152,177]" italics="true" pageId="9" pageNumber="56">Pygopus robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD84EFFE95777FF0642B3A8A5" box="[578,673,151,177]" pageId="9" pageNumber="56" rank="species">
|
||
<emphasis id="A79BEAA1D84EFFE95777FF0642B3A8A5" bold="true" box="[578,673,151,177]" pageId="9" pageNumber="56">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is only known from a few scattered localities in two apparently disjunct areas of north-eastern Queensland (
|
||
<figureCitation id="0DD42A36D84EFFE957ABFF2C42F4A8C3" box="[670,742,189,215]" captionStart="FIGURE 5" captionStartId="9.[151,256,1654,1678]" captionTargetBox="[155,1432,558,1607]" captionTargetId="figure@9.[151,1436,530,1633]" captionTargetPageId="9" captionText="FIGURE 5. Map of Australia showing the localities for museum records of Pygopus robertsi sp. nov. (triangles) and Pygopus lepidopodus (circles). Grey symbols denote localities for specimens genetically typed in this study. Locality records are taken from the collections of the Australian Museum, Western Australian Museum, South Australian Museum, Museum Victoria, Queensland Museum and Australian National Wildlife collection." httpUri="https://zenodo.org/record/197552/files/figure.png" pageId="9" pageNumber="56">Fig. 5</figureCitation>
|
||
). The northernmost records are of two specimens from the Heathlands area (
|
||
<geoCoordinate id="F0DB5074D84EFFE9545EFF7341CEA8E8" box="[363,476,226,252]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="-11.683333">11° 41' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F0DB5074D84EFFE954DFFF73427FA8E8" box="[490,621,226,252]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="142.65">142° 39' E</geoCoordinate>
|
||
). The remaining specimens range from the Shipton's Flat area near Cooktown (
|
||
<geoCoordinate id="F0DB5074D84EFFE95411FE994186A936" box="[292,404,264,290]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="-15.783334">15° 47' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F0DB5074D84EFFE95497FE994231A936" box="[418,547,264,290]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="145.25">145° 15' E</geoCoordinate>
|
||
), south to the Atherton Tableland (Yungaburra,
|
||
<geoCoordinate id="F0DB5074D84EFFE95157FE9944C3A936" box="[1122,1233,264,290]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="-17.266666">17° 16' S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F0DB5074D84EFFE951D5FE994570A936" box="[1248,1378,264,290]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="145.58333">145° 35' E</geoCoordinate>
|
||
). An additional
|
||
<taxonomicName id="52EF4D30D84EFFE95426FEBC416AA952" box="[275,376,301,326]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="A79BEAA1D84EFFE95426FEBC416AA952" box="[275,376,301,326]" italics="true" pageId="9" pageNumber="56">Pygopus</emphasis>
|
||
</taxonomicName>
|
||
specimen (QM J4635) from Irvinebank, Mt Misery (
|
||
<geoCoordinate id="F0DB5074D84EFFE956D2FEBC444AA953" box="[999,1112,301,327]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="-17.466667">17° 28’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F0DB5074D84EFFE95150FEBC44F7A953" box="[1125,1253,301,327]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="145.23334">145° 14’ E</geoCoordinate>
|
||
) was present in the Queensland Museum but was destroyed in 1964. If this specimen was also assignable to
|
||
<emphasis id="A79BEAA1D84EFFE951C1FEC3457FA97F" box="[1268,1389,338,363]" italics="true" pageId="9" pageNumber="56">P. ro b e r t s i</emphasis>
|
||
<emphasis id="A79BEAA1D84EFFE9504DFEC340D9A985" bold="true" pageId="9" pageNumber="56">sp. nov.</emphasis>
|
||
it extends the distribution of this species a little to the SW of the Yungaburra record. A local naturalist has also seen another
|
||
<taxonomicName id="52EF4D30D84EFFE954A0FE0C41E8A9A2" box="[405,506,413,438]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="56" phylum="Chordata" rank="genus">
|
||
<emphasis id="A79BEAA1D84EFFE954A0FE0C41E8A9A2" box="[405,506,413,438]" italics="true" pageId="9" pageNumber="56">Pygopus</emphasis>
|
||
</taxonomicName>
|
||
specimen (
|
||
<figureCitation id="0DD42A36D84EFFE957B5FE0C42D4A9A3" box="[640,710,413,439]" captionStart="FIGURE 4. A" captionStartId="8.[151,255,1906,1930]" captionTargetBox="[154,1436,223,1867]" captionTargetId="figure@8.[151,1436,194,1885]" captionTargetPageId="8" captionText="FIGURE 4. A) Holotype of P. robertsi sp. nov., QM J 47145, in life (photograph by Steve Wilson). B) Pygopus robertsi sp. nov. in life, non-vouchered specimen from the lower slopes of Mt Sampson 15 ° 48 ' S, 145 ° 11 ' E (photograph by Lewis Roberts)." httpUri="https://zenodo.org/record/197551/files/figure.png" pageId="9" pageNumber="56">Fig. 4</figureCitation>
|
||
B) on the lower slopes of Mt Sampson (
|
||
<geoCoordinate id="F0DB5074D84EFFE951A2FE0C451AA9A3" box="[1175,1288,413,439]" direction="south" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="-15.8">15° 48’ S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F0DB5074D84EFFE95020FE0C4581A9A3" box="[1301,1427,413,439]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="145.18333">145° 11’ E</geoCoordinate>
|
||
) (L. Roberts, pers. comm.).
|
||
</paragraph>
|
||
<caption id="C190663BD84EFFE955A2F9E74494AEF9" httpUri="https://zenodo.org/record/197552/files/figure.png" pageId="9" pageNumber="56" targetBox="[155,1432,558,1607]" targetPageId="9">
|
||
<paragraph id="955036B3D84EFFE955A2F9E74494AEF9" blockId="9.[151,1437,1654,1773]" pageId="9" pageNumber="56">
|
||
<emphasis id="A79BEAA1D84EFFE955A2F9E74106AE9A" bold="true" box="[151,276,1654,1678]" pageId="9" pageNumber="56">FIGURE 5</emphasis>
|
||
. Map of Australia showing the localities for museum records of
|
||
<taxonomicName id="52EF4D30D84EFFE956E9F9E64484AE9A" box="[988,1174,1655,1678]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84EFFE956E9F9E64484AE9A" box="[988,1174,1655,1678]" italics="true" pageId="9" pageNumber="56">Pygopus robertsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A79BEAA1D84EFFE951ABF9E744E4AE9A" bold="true" box="[1182,1270,1654,1678]" pageId="9" pageNumber="56">
|
||
<taxonomicNameLabel id="BCA857DAD84EFFE951ABF9E744E4AE9A" box="[1182,1270,1654,1678]" pageId="9" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(triangles) and
|
||
<taxonomicName id="52EF4D30D84EFFE955A2F9064193AEBA" box="[151,385,1687,1710]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="lepidopodus">
|
||
<emphasis id="A79BEAA1D84EFFE955A2F9064193AEBA" box="[151,385,1687,1710]" italics="true" pageId="9" pageNumber="56">Pygopus lepidopodus</emphasis>
|
||
</taxonomicName>
|
||
(circles). Grey symbols denote localities for specimens genetically typed in this study. Locality records are taken from the collections of the Australian Museum, Western Australian Museum, South Australian Museum, Museum Victoria, Queensland Museum and Australian National Wildlife collection.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="955036B3D84EFFEA55F0F88A43BBA8C3" blockId="9.[151,1436,1819,2032]" lastBlockId="10.[151,1437,152,551]" lastPageId="10" lastPageNumber="57" pageId="9" pageNumber="56">
|
||
<emphasis id="A79BEAA1D84EFFE955F0F88A42A9AF21" bold="true" box="[197,699,1819,1845]" pageId="9" pageNumber="56">Natural history and conservation status.</emphasis>
|
||
All records of the southern population of
|
||
<taxonomicName id="52EF4D30D84EFFE9519CF88D4560AF21" box="[1193,1394,1820,1845]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="56" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84EFFE9519CF88D4560AF21" box="[1193,1394,1820,1845]" italics="true" pageId="9" pageNumber="56">Pygopus robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD84EFFE9504CF88A40D8AF4F" pageId="9" pageNumber="56" rank="species">
|
||
<emphasis id="A79BEAA1D84EFFE9504CF88A40D8AF4F" bold="true" pageId="9" pageNumber="56">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
for which habitat data are available are from relatively dry woodland along the western edge of the main Wet Tropics rainforest block. The
|
||
<typeStatus id="4A548811D84EFFE95718F8F74286AF94" box="[557,660,1894,1920]" pageId="9" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
(QM
|
||
<accessionNumber id="8ABCAB50D84EFFE957E8F8F74327AF94" box="[733,821,1894,1920]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J47145" pageId="9" pageNumber="56" type="EnaNcbi">J47145</accessionNumber>
|
||
) was collected at Little Forks on a ridge adjacent to the Annan River at around
|
||
<quantity id="52179B56D84EFFE954EAF81D4239AFB2" box="[479,555,1932,1958]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="56" unit="m" value="300.0">300 m</quantity>
|
||
elevation in open bloodwood forest (
|
||
<taxonomicName id="52EF4D30D84EFFE956C4F81D44F6AFB1" box="[1009,1252,1932,1957]" class="Magnoliopsida" family="Myrtaceae" genus="Corymbia" higherTaxonomySource="CoL" kingdom="Animalia" order="Myrtales" pageId="9" pageNumber="56" phylum="Tracheophyta" rank="species" species="nesophila">
|
||
<emphasis id="A79BEAA1D84EFFE956C4F81D44F6AFB1" box="[1009,1252,1932,1957]" italics="true" pageId="9" pageNumber="56">Corymbia nesophila</emphasis>
|
||
</taxonomicName>
|
||
), with a sparse understorey of
|
||
<taxonomicName id="52EF4D30D84EFFE95472F82041F9AFDE" box="[327,491,1969,1994]" class="Liliopsida" family="Xanthorrhoeaceae" genus="Xanthorrhoea" kingdom="Plantae" order="Asparagales" pageId="9" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="A79BEAA1D84EFFE95472F82041F9AFDE" box="[327,491,1969,1994]" italics="true" pageId="9" pageNumber="56">Xanthorrhoea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="52EF4D30D84EFFE95711F82042B2AFDE" box="[548,672,1969,1994]" class="Magnoliopsida" family="Casuarinaceae" genus="Casuarina" kingdom="Plantae" order="Fagales" pageId="9" pageNumber="56" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="A79BEAA1D84EFFE95711F82042B2AFDE" box="[548,672,1969,1994]" italics="true" pageId="9" pageNumber="56">Casuarina</emphasis>
|
||
</taxonomicName>
|
||
above a grassy ground cover (
|
||
<figureCitation id="0DD42A36D84EFFE95136F8204459AFDF" box="[1027,1099,1969,1995]" captionStart="FIGURE 6" captionStartId="10.[151,258,1579,1603]" captionTargetBox="[151,1436,607,1571]" captionTargetId="figure@10.[151,1436,607,1571]" captionTargetPageId="10" captionText="FIGURE 6. Open forest habitat at the type locality of Pygopus robertsi sp. nov. - Little Forks, Shipton's Flat, via Cooktown, Queensland (photograph by Jeanette Covacevich)." httpUri="https://zenodo.org/record/197553/files/figure.png" pageId="9" pageNumber="56">Fig. 6</figureCitation>
|
||
), and the specimen observed at Mt Sampson came from similar habitat. The single specimen of the northern population for which habitat data is available (
|
||
<typeStatus id="4A548811D84DFFEA5454FF0941DAA8A6" box="[353,456,152,178]" pageId="10" pageNumber="57" type="paratype">paratype</typeStatus>
|
||
QM
|
||
<accessionNumber id="8ABCAB50D84DFFEA5730FF094249A8A6" box="[517,603,152,178]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/J57778" pageId="10" pageNumber="57" type="EnaNcbi">J57778</accessionNumber>
|
||
) was collected near the road approaching the Heathlands ranger station in low (approx
|
||
<quantity id="52179B56D84DFFEA547FFF2C416BA8C3" box="[330,377,189,215]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="57" unit="m" value="2.0">2 m</quantity>
|
||
) regenerating heath (Barry Lyon, pers. comm.).
|
||
</paragraph>
|
||
<paragraph id="955036B3D84DFFEA55F0FF734498A986" blockId="10.[151,1437,152,551]" pageId="10" pageNumber="57">
|
||
Despite its large size and moderately wide distribution, spanning a well-surveyed region,
|
||
<taxonomicName id="52EF4D30D84DFFEA5007FF7340E1A935" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="57" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84DFFEA5007FF7340E1A935" italics="true" pageId="10" pageNumber="57">Pygopus robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD84DFFEA55CEFE964148A935" box="[251,346,263,289]" pageId="10" pageNumber="57" rank="species">
|
||
<emphasis id="A79BEAA1D84DFFEA55CEFE964148A935" bold="true" box="[251,346,263,289]" pageId="10" pageNumber="57">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
remains poorly known and rare in museum collections. Whilst this may reflect true rarity it could also result from cryptic behaviour. Apparently suitable habitat (open woodland and heath) is moderately widespread, but has not received the same survey attention as nearby patches of rainforest. On this basis we suggest that
|
||
<taxonomicName id="52EF4D30D84DFFEA541CFEE941B0A985" box="[297,418,376,401]" class="Reptilia" family="Pygopodidae" genus="Pygopus" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="57" phylum="Chordata" rank="species" species="robertsi" status="sp. nov.">
|
||
<emphasis id="A79BEAA1D84DFFEA541CFEE941B0A985" box="[297,418,376,401]" italics="true" pageId="10" pageNumber="57">P. robertsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BCA857DAD84DFFEA549CFEE64215A985" box="[425,519,375,401]" pageId="10" pageNumber="57" rank="species">
|
||
<emphasis id="A79BEAA1D84DFFEA549CFEE64215A985" bold="true" box="[425,519,375,401]" pageId="10" pageNumber="57">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
be classified as data deficient, pending further studies.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DDF56538D84DFFEA55F0FE0C4185AA33" pageId="10" pageNumber="57" type="etymology">
|
||
<paragraph id="955036B3D84DFFEA55F0FE0C4185AA33" blockId="10.[151,1437,152,551]" pageId="10" pageNumber="57">
|
||
<emphasis id="A79BEAA1D84DFFEA55F0FE0C4144A9A3" bold="true" box="[197,342,413,439]" pageId="10" pageNumber="57">Etymology.</emphasis>
|
||
Named for Charles George Roberts who, with his brother Lewis, has assisted zoologists, botanists and ecologists with their field studies in north-east Queensland for over four decades. The Roberts brothers are recognised experts on the flora and fauna of the Cooktown area and Honorary Consultants to the Queensland Museum.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |