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treatments-xml/data/18/48/87/184887E9FFF5BD5E1F9CACE091B70776.xml
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<document id="4F7F0C0BFD3DADD5436293EAAC66AF8B" ID-CLB-Dataset="55566" ID-DOI="10.5281/zenodo.5305057" ID-GBIF-Dataset="c7ccba22-dcb8-4e60-b5fb-a8439522e453" ID-ISSN="0253-116X" ID-Zenodo-Dep="5305057" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630111141298" checkinUser="marcus" docAuthor="Assing, V." docDate="2014" docId="184887E9FFF5BD5E1F9CACE091B70776" docLanguage="en" docName="LinzerbiolBeitr.2014.46.1.461-479.pdf" docOrigin="Linzer biologische Beiträge 46 (1)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Orsunius granulosus Assing 2014, nov.sp." docType="treatment" docVersion="9" lastPageNumber="472" masterDocId="E471FF91FFFFBD551F2EA97490250160" masterDocTitle="On Orsunius II. Eight new species and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina)" masterLastPageNumber="479" masterPageNumber="461" pageNumber="471" updateTime="1699155222530" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="063ADE18252ACF5E93C84083D1DFCC15" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="19082D718C8206700C546482C35D2E9C">
<mods:title id="C3B7CA72B1B9F3C06184B14F15CC8DAB">On Orsunius II. Eight new species and additional records (Coleoptera: Staphylinidae: Paederinae: Medonina)</mods:title>
</mods:titleInfo>
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</mods:role>
<mods:namePart id="013B167E9BC3D4B97E1C23FC132B798A">Assing, V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="4A21DF722CD4E595562F825A1E927CE7">text</mods:typeOfResource>
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<mods:title id="A1CDCD3BA32B74BF9C5DE5737BAD4537">Linzer biologische Beiträge</mods:title>
</mods:titleInfo>
<mods:part id="4730511836A2948598DBDED6FC59AC67">
<mods:date id="A897733D1F5BC13E0A394AB1F1BF8FF9">2014</mods:date>
<mods:detail id="919F04783FEFBCDBCC4E10E92D0BE229" type="pubDate">
<mods:number id="95A26400A3783843D75C1B1577FC27AA">2014-07-31</mods:number>
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<mods:number id="EE8AD5F507E0645C45F62FDA9A7E28D0">46</mods:number>
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</mods:extent>
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<mods:classification id="3B3AFD37E1654392D3D338F1A41AD178">journal article</mods:classification>
<mods:identifier id="75AD352B6D25E23268FA8F06DA50931F" type="CLB-Dataset">55566</mods:identifier>
<mods:identifier id="B6C53EB73F86E238497D2A93F0C0AB87" type="DOI">10.5281/zenodo.5305057</mods:identifier>
<mods:identifier id="761C54AAA18B2F3F5201FC6AB8DA8C5D" type="GBIF-Dataset">c7ccba22-dcb8-4e60-b5fb-a8439522e453</mods:identifier>
<mods:identifier id="A4D22554E6EDCD80C75B379FDDE7DA0D" type="ISSN">0253-116X</mods:identifier>
<mods:identifier id="0751FDD924B2BD389791DD63CCE76B47" type="Zenodo-Dep">5305057</mods:identifier>
</mods:mods>
<treatment id="184887E9FFF5BD5E1F9CACE091B70776" ID-DOI="http://doi.org/10.5281/zenodo.6539887" ID-GBIF-Taxon="195300188" ID-Zenodo-Dep="6539887" LSID="urn:lsid:plazi:treatment:184887E9FFF5BD5E1F9CACE091B70776" httpUri="http://treatment.plazi.org/id/184887E9FFF5BD5E1F9CACE091B70776" lastPageId="11" lastPageNumber="472" pageId="10" pageNumber="471">
<subSubSection id="D8FB6574FFF5BD5F1F9CACE092F404CB" box="[178,721,1428,1451]" pageId="10" pageNumber="471" type="nomenclature">
<paragraph id="905E36FFFFF5BD5F1F9CACE092F404CB" blockId="10.[178,721,1428,1451]" box="[178,721,1428,1451]" pageId="10" pageNumber="471">
<heading id="CB168193FFF5BD5F1F9CACE092F404CB" bold="true" box="[178,721,1428,1451]" fontSize="9" level="2" pageId="10" pageNumber="471" reason="3">
<emphasis id="A295EAEDFFF5BD5F1F9CACE091D704CB" bold="true" box="[178,498,1428,1451]" pageId="10" pageNumber="471">
<taxonomicName id="57E14D7CFFF5BD5F1F9CACE091BF04CB" ID-CoL="6SXHW" authority="Assing, 2014" authorityName="Assing" authorityYear="2014" box="[178,410,1428,1451]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="471" phylum="Arthropoda" rank="species" species="granulosus" status="sp. nov.">
<emphasis id="A295EAEDFFF5BD5F1F9CACE091BF04CB" bold="true" box="[178,410,1428,1451]" italics="true" pageId="10" pageNumber="471">Orsunius granulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9A65796FFF5BD5F1E8CACE091D704CB" box="[418,498,1428,1451]" pageId="10" pageNumber="471" rank="species">nov.sp.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="08DA2A7AFFF5BD5F1D2EACE0925D04CB" box="[512,632,1428,1451]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="10" pageNumber="471">Figs 26-32</figureCitation>
,
<figureCitation id="08DA2A7AFFF5BD5F1DADACE092ED04CB" box="[643,712,1428,1451]" captionStart="Map 2" captionStartId="2.[178,223,1075,1095]" captionTargetBox="[178,1092,212,1064]" captionTargetId="figure-2@2.[178,1092,212,1064]" captionTargetPageId="2" captionText="Map 2: Distributions of Orsunius gladiator (white and black circles), O. latexcisus (black circle), O. flavovirgatus (black circle), O. heissi (black star), O. stimulans (black and grey triangles), O. granulosus (white and grey triangles), O. cuneatus (white diamond), and O. spathulatus (black square)." figureDoi="http://doi.org/10.5281/zenodo.5305061" httpUri="https://zenodo.org/record/5305061/files/figure.png" pageId="10" pageNumber="471">Map 2</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8FB6574FFF5BD5E1F9CACC891130062" lastPageId="11" lastPageNumber="472" pageId="10" pageNumber="471" type="materials_examined">
<paragraph id="905E36FFFFF5BD5E1F9CACC891130062" blockId="10.[178,1085,1468,1601]" lastBlockId="11.[178,1086,215,258]" lastPageId="11" lastPageNumber="472" pageId="10" pageNumber="471">
Type material:
<materialsCitation id="20893CA2FFF5BD5F1EB2ACC892F70774" ID-GBIF-Occurrence="3766395315" collectingDate="2008-06-13" collectingDateMax="2008-06-14" collectingDateMin="2008-06-13" collectionCode="ZMUC" collectorName="A. Solodovnikov &amp; J. Pedersen" country="Laos" county="Bolaven Plt" determinerName="V. Assing" elevation="1100" latitude="15.05285" location="Ban Hoyayteuy" longLatPrecision="1" longitude="106.28995" municipality="Muang Paxong" pageId="10" pageNumber="471" specimenCount="1" stateProvince="Champasak prov." typeStatus="holotype">
<typeStatus id="4F5A885DFFF5BD5F1EB2ACC891D704B0" box="[412,498,1468,1488]" pageId="10" pageNumber="471" type="holotype">Holotype</typeStatus>
: &quot;
<collectingCountry id="E8F6766FFFF5BD5F1D09ACC8927D04B0" box="[551,600,1468,1488]" name="Laos" pageId="10" pageNumber="471">Laos</collectingCountry>
:
<collectingRegion id="5225F81DFFF5BD5F1D4EACC8932104B0" box="[608,772,1468,1488]" country="Laos" name="Champasak" pageId="10" pageNumber="471">Champasak prov.</collectingRegion>
:
<collectingCounty id="793F4E73FFF5BD5F1C22ACC8935D04B0" box="[780,888,1468,1488]" pageId="10" pageNumber="471">Bolaven Plt</collectingCounty>
,
<collectingMunicipality id="703AAC85FFF5BD5F1CACACC8942A04B0" box="[898,1039,1468,1488]" pageId="10" pageNumber="471">Muang Paxong</collectingMunicipality>
,
<location id="953E6024FFF5BD5F1B36ACC8913D0486" LSID="urn:lsid:plazi:treatment:184887E9FFF5BD5E1F9CACE091B70776:953E6024FFF5BD5F1B36ACC8913D0486" country="Laos" county="Bolaven Plt" latitude="15.05285" longLatPrecision="1" longitude="106.28995" municipality="Muang Paxong" name="Ban Hoyayteuy" pageId="10" pageNumber="471" stateProvince="Champasak prov.">Ban Hoyayteuy</location>
,
<location id="953E6024FFF5BD5F1E0CACA691B20486" LSID="urn:lsid:plazi:treatment:184887E9FFF5BD5E1F9CACE091B70776:953E6024FFF5BD5F1E0CACA691B20486" box="[290,407,1490,1510]" country="Laos" county="Bolaven Plt" latitude="15.05285" longLatPrecision="1" longitude="106.28995" municipality="Muang Paxong" name="Mt. Phu Din" pageId="10" pageNumber="471" stateProvince="Champasak prov.">Mt. Phu Din</location>
,
<date id="E45F103FFFF5BD5F1E8CACA692010486" box="[418,548,1490,1510]" pageId="10" pageNumber="471" value="2008-06-13" valueMax="2008-06-14" valueMin="2008-06-13">
<collectingDate id="F41BE9D7FFF5BD5F1E8CACA692010486" box="[418,548,1490,1510]" pageId="10" pageNumber="471" value="2008-06-13" valueMax="2008-06-14" valueMin="2008-06-13">13-14.vi.2008</collectingDate>
</date>
;
<quantity id="57199B1AFFF5BD5F1D00ACA692530486" box="[558,630,1490,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="471" unit="m" value="1100.0">
<elevation id="1BCCD1CCFFF5BD5F1D00ACA692530486" box="[558,630,1490,1510]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="10" pageNumber="471" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<geoCoordinate id="F5D55038FFF5BD5F1DAEACA692D60486" box="[640,755,1490,1510]" degrees="15" direction="north" minutes="03.171" orientation="latitude" pageId="10" pageNumber="471" precision="1" value="15.05285">15°03.171'N</geoCoordinate>
<geoCoordinate id="F5D55038FFF5BD5F1DD7ACA693500486" box="[761,885,1490,1510]" degrees="106" direction="east" minutes="17.397" orientation="longitude" pageId="10" pageNumber="471" precision="1" value="106.28995">106°17.397'E</geoCoordinate>
/
<collectorName id="3D145329FFF5BD5F1CA6ACA694000486" box="[904,1061,1490,1510]" pageId="10" pageNumber="471">A. Solodovnikov</collectorName>
&amp;
<collectorName id="3D145329FFF5BD5F1F9CAC9D913E049D" box="[178,283,1513,1533]" pageId="10" pageNumber="471">J. Pedersen</collectorName>
leg., Disturbed primary rainforest, sifted leaf litter; ZMUC collection / Holotypus
<originalDetermination id="B3FF8E03FFF5BD5F1F9CAF7491820774" box="[178,423,1536,1556]" pageId="10" pageNumber="471">
<taxonomicName id="57E14D7CFFF5BD5F1F9CAF7491560774" authority="Assing, 2014" authorityName="Assing" authorityYear="2014" box="[178,371,1536,1556]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="471" phylum="Arthropoda" rank="species" species="granulosus" status="sp. nov.">
<emphasis id="A295EAEDFFF5BD5F1F9CAF7491560774" box="[178,371,1536,1556]" italics="true" pageId="10" pageNumber="471">Orsunius granulosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B9A65796FFF5BD5F1E55AF7491820774" box="[379,423,1536,1556]" pageId="10" pageNumber="471" rank="species">sp.n.</taxonomicNameLabel>
</originalDetermination>
det.
<determinerName id="6582DA3BFFF5BD5F1EF7AF7492120774" box="[473,567,1536,1556]" pageId="10" pageNumber="471">V. Assing</determinerName>
2012&quot; (
<collectionCode id="F6F0AE3AFFF5BD5F1DABAF7492E90774" box="[645,716,1536,1556]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="10" pageNumber="471">ZMUC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="20893CA2FFF5BD5F1DF0AF7491B1074B" ID-GBIF-Occurrence="3766395312" collectingDate="2008-06-13" collectingDateMax="2008-06-14" collectingDateMin="2008-06-13" collectionCode="ZMUC" collectorName="A. Solodovnikov &amp; J. Pedersen" country="Laos" county="Bolaven Plt" elevation="1100" latitude="15.05285" location="Ban Hoyayteuy" longLatPrecision="1" longitude="106.28995" municipality="Muang Paxong" pageId="10" pageNumber="471" specimenCount="1" stateProvince="Champasak" typeStatus="paratype">
<typeStatus id="4F5A885DFFF5BD5F1DF0AF7493130774" box="[734,822,1536,1556]" pageId="10" pageNumber="471" type="paratype">Paratypes</typeStatus>
: 3, 3: same data as holotype (
<collectionCode id="F6F0AE3AFFF5BD5F1E20AF639176074B" box="[270,339,1559,1579]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="10" pageNumber="471">ZMUC</collectionCode>
, cAss)
</materialsCitation>
;
<materialsCitation id="20893CA2FFF5BD5F1EB0AF63941F0721" ID-GBIF-Occurrence="3766395305" collectingDate="2010-04-11" collectingDateMax="2010-04-24" collectingDateMin="2010-04-11" collectionCode="NHMB, cAss" collectorName="L. Dembicky" country="Vietnam-N" elevation="400" latitude="17.689444" location="Vietnam-Laos border area" longLatPrecision="21" longitude="105.7625" municipality="Quang Binh prov." pageId="10" pageNumber="471" specimenCount="4">
<specimenCount id="86E7FD76FFF5BD5F1EB0AF63918C074B" box="[414,425,1559,1579]" pageId="10" pageNumber="471">4</specimenCount>
, 3: &quot;
<collectingCountry id="E8F6766FFFF5BD5F1D0FAF6392A9074B" box="[545,652,1559,1579]" pageId="10" pageNumber="471">Vietnam-N</collectingCountry>
,
<collectingMunicipality id="703AAC85FFF5BD5F1DBBAF63931B074B" box="[661,830,1559,1579]" pageId="10" pageNumber="471">Quang Binh prov.</collectingMunicipality>
,
<locationDeviation id="81EC2FDDFFF5BD5F1C69AF639383074B" bearing="0.0" box="[839,934,1559,1579]" distance="1000.0" pageId="10" pageNumber="471">
<quantity id="57199B1AFFF5BD5F1C69AF639353074B" box="[839,886,1559,1579]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="471" unit="km" value="1.0">1 km</quantity>
N of
</locationDeviation>
<collectingMunicipality id="703AAC85FFF5BD5F1C83AF6393D6074B" box="[941,1011,1559,1579]" pageId="10" pageNumber="471">Cha Lo</collectingMunicipality>
,
<quantity id="57199B1AFFF5BD5F1CD3AF63941F074B" box="[1021,1082,1559,1579]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="471" unit="m" value="400.0">
<elevation id="1BCCD1CCFFF5BD5F1CD3AF63941F074B" box="[1021,1082,1559,1579]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="471" unit="m" value="400.0">400 m</elevation>
</quantity>
,
<location id="953E6024FFF5BD5F1F9CAF5991B00721" LSID="urn:lsid:plazi:treatment:184887E9FFF5BD5E1F9CACE091B70776:953E6024FFF5BD5F1F9CAF5991B00721" box="[178,405,1581,1601]" country="Vietnam-N" latitude="17.689444" longLatPrecision="21" longitude="105.7625" municipality="Quang Binh prov." name="Vietnam-Laos border area" pageId="10" pageNumber="471">Vietnam-Laos border area</location>
,
<geoCoordinate id="F5D55038FFF5BD5F1EB1AF5992270721" box="[415,514,1581,1601]" degrees="17" direction="north" minutes="41" orientation="latitude" pageId="10" pageNumber="471" precision="15" seconds="22" value="17.689444">17°41'22''N</geoCoordinate>
,
<geoCoordinate id="F5D55038FFF5BD5F1D22AF5992520721" box="[524,631,1581,1601]" degrees="105" direction="east" minutes="45" orientation="longitude" pageId="10" pageNumber="471" precision="15" seconds="45" value="105.7625">105°45'45''E</geoCoordinate>
,
<collectorName id="3D145329FFF5BD5F1DAFAF5992D60721" box="[641,755,1581,1601]" pageId="10" pageNumber="471">L. Dembický</collectorName>
leg.,
<date id="E45F103FFFF5BD5F1C0BAF5993850721" box="[805,928,1581,1601]" pageId="10" pageNumber="471" value="2010-04-11" valueMax="2010-04-24" valueMin="2010-04-11">
<collectingDate id="F41BE9D7FFF5BD5F1C0BAF5993850721" box="[805,928,1581,1601]" pageId="10" pageNumber="471" value="2010-04-11" valueMax="2010-04-24" valueMin="2010-04-11">11.-24.iv.2010</collectingDate>
</date>
&quot; (
<collectionCode id="F6F0AE3AFFF5BD5F1C9AAF5993DD0721" box="[948,1016,1581,1601]" country="0" httpUri="http://grbio.org/cool/1d5a-4efs" name="Natural History Museum Bucharest" pageId="10" pageNumber="471">NHMB</collectionCode>
,
<collectionCode id="F6F0AE3AFFF5BD5F1B2DAF5994160721" box="[1027,1075,1581,1601]" pageId="10" pageNumber="471">cAss</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="20893CA2FFF4BD5E1F9CA9A391130062" ID-GBIF-Occurrence="3766395320" pageId="11" pageNumber="472">
2: &quot;
<collectingCountry id="E8F6766FFFF4BD5E1FD4A9A39154018B" box="[250,369,215,235]" name="Thailand" pageId="11" pageNumber="472">THAILAND</collectingCountry>
- river near Ban Pong Din,
<quantity id="57199B1AFFF4BD5E1DA9A9A392E1018B" box="[647,708,215,235]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="472" unit="km" value="32.0">32 km</quantity>
ENE
<collectingRegion id="5225F81DFFF4BD5E1C2DA9A39351018B" box="[771,884,215,235]" country="Thailand" name="Chiang Mai" pageId="11" pageNumber="472">Chiang Mai</collectingRegion>
,
<date id="E45F103FFFF4BD5E1CAFA9A393CF018B" box="[897,1002,215,235]" pageId="11" pageNumber="472" value="2004-11-27">27.XI.2004</date>
, leg. W. Rossi&quot; (cAss).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D8FB6574FFF4BD5E1F9CA86491AF0023" pageId="11" pageNumber="472" type="etymology">
<paragraph id="905E36FFFFF4BD5E1F9CA86491AF0023" blockId="11.[178,1085,272,323]" pageId="11" pageNumber="472">Etymology: The specific epithet (Latin, adjective) refers to the granulose punctation of the pronotum.</paragraph>
</subSubSection>
<subSubSection id="D8FB6574FFF4BD5E1F9CA826913905E9" pageId="11" pageNumber="472" type="description">
<paragraph id="905E36FFFFF4BD5E1F9CA826919700A0" blockId="11.[178,1085,338,448]" pageId="11" pageNumber="472">
Description: Body length
<quantity id="57199B1AFFF4BD5E1D33A82692B20008" box="[541,663,338,360]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.2" metricValueMin="3.7" pageId="11" pageNumber="472" unit="mm" value="3.95" valueMax="4.2" valueMin="3.7">3.7-4.2 mm</quantity>
; length of forebody
<quantity id="57199B1AFFF4BD5E1C44A82693C60008" box="[874,995,338,360]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.1" metricValueMin="1.9" pageId="11" pageNumber="472" unit="mm" value="2.0" valueMax="2.1" valueMin="1.9">1.9-2.1 mm</quantity>
. Habitus as in
<figureCitation id="08DA2A7AFFF4BD5E1FC7A804911100E6" box="[233,308,368,390]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 26</figureCitation>
. Coloration: body reddish to dark-brown, with the elytra sometimes slightly paler and the elytral postero-lateral angles weakly infuscate; legs yellowish to yellowishbrown; antennae reddish.
</paragraph>
<paragraph id="905E36FFFFF4BD5E1F9CA8A4922B03D2" blockId="11.[178,1086,464,690]" pageId="11" pageNumber="472">
Head (
<figureCitation id="08DA2A7AFFF4BD5E1FD6A8A491610086" box="[248,324,464,486]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 27</figureCitation>
) of transversely rectangular shape, approximately 1.15 times as broad as long; posterior margin distinctly concave in the middle; punctation distinct and dense; interstices narrower than diameter of punctures and glossy, without microsculpture; sides with long dark-brown setae. Eyes large, on average approximately as long as postocular portion in dorsal view, often slightly shorter or longer. Antennae slender,
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long; preapical antennomeres approximately as wide as long. Anterior margin of labrum with broadly V-shaped excision in the middle; left mandible with three, right mandible with four pronounced molar teeth.
</paragraph>
<paragraph id="905E36FFFFF4BD5E1F9CABB794250250" blockId="11.[178,1085,706,816]" pageId="11" pageNumber="472">
Pronotum (
<figureCitation id="08DA2A7AFFF4BD5E1E0BABB6915503B8" box="[293,368,706,728]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 27</figureCitation>
) approximately 1.2 times as wide as long and 1.05 times as broad as head; posterior angles rounded, weakly marked; lateral and anterior margins with long dark setae; punctation very dense and distinctly, finely granulose, rendering the pronotum matt; with or without rudiment of an impunctate median band in posterior half.
</paragraph>
<paragraph id="905E36FFFFF4BD5E1F9CAA4B92F302CC" blockId="11.[178,1086,831,941]" pageId="11" pageNumber="472">
Elytra (
<figureCitation id="08DA2A7AFFF4BD5E1E2FAA4B91680235" box="[257,333,831,853]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 27</figureCitation>
) 1.10-1.15 times as long as pronotum; punctation very dense, fine, and somewhat asperate. Hind wings fully developed. Protarsomeres I-IV sexually dimorphic, more distinctly dilated in males than in females. Metatarsomere I longer than II and slightly shorter than the combined length of II and III.
</paragraph>
<paragraph id="905E36FFFFF4BD5E1F9CAAC893A2028F" blockId="11.[178,1085,956,1007]" pageId="11" pageNumber="472">Abdomen narrower than elytra; punctation extremely fine and dense, barely noticeable in the microsculpture; posterior margin of tergite VII with palisade fringe.</paragraph>
<paragraph id="905E36FFFFF4BD5E1FE9AA8B913905E9" blockId="11.[178,1085,1023,1161]" pageId="11" pageNumber="472">
: sternite VII (
<figureCitation id="08DA2A7AFFF4BD5E1E71AA8B918E0575" box="[351,427,1023,1045]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 28</figureCitation>
) with weakly concave posterior margin; sternite VIII (
<figureCitation id="08DA2A7AFFF4BD5E1CC7AA8B94100575" box="[1001,1077,1023,1045]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 29</figureCitation>
) oblong and with posterior excision of trapezoid shape (
<figureCitation id="08DA2A7AFFF4BD5E1DDAAD6893660552" box="[756,835,1052,1074]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Fig. 30</figureCitation>
); aedeagus (
<figureCitation id="08DA2A7AFFF4BD5E1CEAAD6894130552" box="[964,1078,1052,1074]" captionStart="Figs 26-35" captionStartId="12.[178,218,1516,1536]" captionTargetBox="[187,1087,216,1505]" captionTargetId="figure-2@12.[178,1091,212,1505]" captionTargetPageId="12" captionText="Figs 26-35: Orsunius granulosus nov.sp. (26-32), O. latexcisus nov.sp. (33) and O. flavovirgatus nov.sp. (34-35): (26) habitus; (27) forebody; (28) male sternite VII; (29, 33, 34) male sternite VIII; (30, 35) posterior portion of male sternite VIII; (31) aedeagus in lateral view; (32) aedeagus in ventral view. Scale bars: 26-27: 1.0 mm; 28-29, 31-34: 0.2 mm; 30, 35: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5305069" httpUri="https://zenodo.org/record/5305069/files/figure.png" pageId="11" pageNumber="472">Figs 31-32</figureCitation>
) small, approximately
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long; internal sac with a pair of strongly sclerotized short and curved apical structures and with a pair of longer, less strongly sclerotized basal structures.
</paragraph>
</subSubSection>
<subSubSection id="D8FB6574FFF4BD5E1F9CADED91FC04F9" pageId="11" pageNumber="472" type="discussion">
<paragraph id="905E36FFFFF4BD5E1F9CADED91FC04F9" blockId="11.[178,1085,1177,1433]" pageId="11" pageNumber="472">
Comparative notes: This and the two following species are attributed to
<taxonomicName id="57E14D7CFFF4BD5E1F9CADC3912B05AC" authorityName="ASSING" authorityYear="2011" box="[178,270,1207,1228]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="472" phylum="Arthropoda" rank="genus">
<emphasis id="A295EAEDFFF4BD5E1F9CADC3912B05AC" box="[178,270,1207,1228]" italics="true" pageId="11" pageNumber="472">Orsunius</emphasis>
</taxonomicName>
with some hesitation. They share the habitus, the head shape, and other external characters with the species previously assigned to the genus, but are distinguished from them by the presence of four (rather than three) molar teeth on the right mandible, the broadly V-shaped median incision of the anterior margin of the labrum, the conspicuous granulose punctation of the pronotum, and by a rather different morphology of the aedeagus. From the closely related
<taxonomicName id="57E14D7CFFF4BD5E1D1EAC3D928B043F" authorityName="Assing" authorityYear="2014" box="[560,686,1353,1375]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="472" phylum="Arthropoda" rank="species" species="cuneatus">
<emphasis id="A295EAEDFFF4BD5E1D1EAC3D928B043F" box="[560,686,1353,1375]" italics="true" pageId="11" pageNumber="472">O. cuneatus</emphasis>
</taxonomicName>
and
<taxonomicName id="57E14D7CFFF4BD5E1DC7AC3D9363043F" authorityName="Assing" authorityYear="2014" box="[745,838,1353,1375]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="472" phylum="Arthropoda" rank="species" species="heissi">
<emphasis id="A295EAEDFFF4BD5E1DC7AC3D9363043F" box="[745,838,1353,1375]" italics="true" pageId="11" pageNumber="472">O. heissi</emphasis>
</taxonomicName>
,
<taxonomicName id="57E14D7CFFF4BD5E1C77AC3D93C9043F" authorityName="Assing" authorityYear="2014" box="[857,1004,1353,1375]" class="Insecta" family="Staphylinidae" genus="Orsunius" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="472" phylum="Arthropoda" rank="species" species="granulosus">
<emphasis id="A295EAEDFFF4BD5E1C77AC3D93C9043F" box="[857,1004,1353,1375]" italics="true" pageId="11" pageNumber="472">O. granulosus</emphasis>
</taxonomicName>
is best distinguished by the trapezoid posterior excision of the male sternite VII and by the morphology of the aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="D8FB6574FFF4BD5E1F9CACDC91B70776" pageId="11" pageNumber="472" type="distribution">
<paragraph id="905E36FFFFF4BD5E1F9CACDC91B70776" blockId="11.[178,1085,1448,1558]" pageId="11" pageNumber="472">
Distribution and natural history: The species is known from three localities in southern
<collectingCountry id="E8F6766FFFF4BD5E1EEBACB291DC04BB" box="[453,505,1478,1499]" name="Laos" pageId="11" pageNumber="472">Laos</collectingCountry>
, North
<collectingCountry id="E8F6766FFFF4BD5E1D6BACB2928704BC" box="[581,674,1478,1500]" name="Thailand" pageId="11" pageNumber="472">Thailand</collectingCountry>
, and North
<collectingCountry id="E8F6766FFFF4BD5E1C36ACB2934A04BC" box="[792,879,1478,1500]" name="Vietnam" pageId="11" pageNumber="472">Vietnam</collectingCountry>
(
<figureCitation id="08DA2A7AFFF4BD5E1C52ACB293E704BC" box="[892,962,1478,1500]" captionStart="Map 2" captionStartId="2.[178,223,1075,1095]" captionTargetBox="[178,1092,212,1064]" captionTargetId="figure-2@2.[178,1092,212,1064]" captionTargetPageId="2" captionText="Map 2: Distributions of Orsunius gladiator (white and black circles), O. latexcisus (black circle), O. flavovirgatus (black circle), O. heissi (black star), O. stimulans (black and grey triangles), O. granulosus (white and grey triangles), O. cuneatus (white diamond), and O. spathulatus (black square)." figureDoi="http://doi.org/10.5281/zenodo.5305061" httpUri="https://zenodo.org/record/5305061/files/figure.png" pageId="11" pageNumber="472">Map 2</figureCitation>
). The partly teneral specimens from
<collectingCountry id="E8F6766FFFF4BD5E1E84AC9091FE0499" box="[426,475,1508,1529]" name="Laos" pageId="11" pageNumber="472">Laos</collectingCountry>
were sifted from leaf litter in a degraded primary forest at an altitude of
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.
</paragraph>
</subSubSection>
</treatment>
</document>
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