763 lines
99 KiB
XML
763 lines
99 KiB
XML
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<mods:title id="D5DAC3AB1FD25876EFA66AAC46268175">Taxonomy and biostratigraphy of the elasmobranchs and bony fishes (Chondrichthyes and Osteichthyes) of the lower-to-middle Eocene (Ypresian to Bartonian) Claiborne Group in Alabama, USA, including an analysis of otoliths</mods:title>
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<mods:namePart id="319952346590E8DE26BB151BE770DCF1">Ebersole, Jun A.</mods:namePart>
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<mods:namePart id="4F2751347EF0BFEC6FA2D7B845CB1C86">Cicimurri, David J.</mods:namePart>
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<mods:namePart id="DCBFD5319BC6103A31E176E96AFE7A9E">Stringer, Gary L.</mods:namePart>
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<mods:title id="0FAD9D2D3E6324D1C883274A66C2674A">European Journal of Taxonomy</mods:title>
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<treatment id="18174D41FF0EFF20FD9099AF483C0D37" ID-DOI="http://doi.org/10.5281/zenodo.3664266" ID-GBIF-Taxon="162073147" ID-Zenodo-Dep="3664266" LSID="urn:lsid:plazi:treatment:18174D41FF0EFF20FD9099AF483C0D37" httpUri="http://treatment.plazi.org/id/18174D41FF0EFF20FD9099AF483C0D37" lastPageId="143" lastPageNumber="144" pageId="139" pageNumber="140">
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<subSubSection id="D8A4AFDCFF0EFF24FD9099AF4E0809ED" pageId="139" pageNumber="140" type="nomenclature">
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<paragraph id="9001FC57FF0EFF24FD9099AF4E960981" blockId="139.[598,989,1691,1754]" box="[598,989,1691,1718]" pageId="139" pageNumber="140">
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<heading id="CB494B3BFF0EFF24FD9099AF4E960981" box="[598,989,1691,1718]" centered="true" fontSize="11" level="2" pageId="139" pageNumber="140" reason="2">
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<taxonomicName id="57BE87D4FF0EFF24FD9099AF4E960981" authority="(Leidy, 1876)" baseAuthorityName="Leidy" baseAuthorityYear="1876" box="[598,989,1691,1718]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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<emphasis id="A2CA2045FF0EFF24FD9099AF4E7D0982" bold="true" box="[598,822,1691,1718]" italics="true" pageId="139" pageNumber="140">Leidybatis jugosus</emphasis>
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||
(
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<bibRefCitation id="F42F81A6FF0EFF24FC8099AF4E9E0981" author="Leidy J." box="[838,981,1691,1718]" pageId="139" pageNumber="140" pagination="86 - 87" refId="ref144835" refString="Leidy J. 1876. Remarks on fossils of the Ashley Phosphate Beds. Proceedings of the Academy of Natural Sciences of Philadelphia 28: 86 - 87." type="journal article" year="1876">Leidy, 1876</bibRefCitation>
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)
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="9001FC57FF0EFF24FD36998B4E0809ED" blockId="139.[598,989,1691,1754]" box="[752,835,1727,1754]" pageId="139" pageNumber="140">
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<figureCitation id="0885E0D2FF0EFF24FD36998B4E0809ED" box="[752,835,1727,1754]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="139" pageNumber="140">Fig. 52</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D8A4AFDCFF0EFF24FF7B98354E1708DC" pageId="139" pageNumber="140" type="reference_group">
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<paragraph id="9001FC57FF0EFF24FF7B98354FCF082C" blockId="139.[189,644,1793,1819]" box="[189,644,1793,1819]" pageId="139" pageNumber="140">
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<taxonomicName id="57BE87D4FF0EFF24FF7B98354FCB082C" authority="Leidy, 1876, p. 86" authorityName="Leidy" authorityPageNumber="86" authorityYear="1876" box="[189,640,1793,1819]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobates" higherTaxonomySource="CoL" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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<emphasis id="A2CA2045FF0EFF24FF7B98354CEA082C" box="[189,417,1793,1819]" italics="true" pageId="139" pageNumber="140">Myliobates jugosus</emphasis>
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<bibRefCitation id="F42F81A6FF0EFF24FE6E98354F7D082C" author="Leidy J." box="[424,566,1793,1819]" pageId="139" pageNumber="140" pagination="86 - 87" refId="ref144835" refString="Leidy J. 1876. Remarks on fossils of the Ashley Phosphate Beds. Proceedings of the Academy of Natural Sciences of Philadelphia 28: 86 - 87." type="journal article" year="1876">Leidy, 1876</bibRefCitation>
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, p. 86
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</taxonomicName>
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||
.
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</paragraph>
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||
<paragraph id="9001FC57FF0EFF24FF7B98764E0E086A" blockId="139.[189,1398,1858,2027]" box="[189,837,1858,1885]" pageId="139" pageNumber="140">
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<treatmentCitationGroup id="B0AEDB79FF0EFF24FF7B98764E0E086A" box="[189,837,1858,1885]" pageId="139" pageNumber="140">
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<taxonomicName id="57BE87D4FF0EFF24FF7B98764FC8086A" authority="- Leidy 1877: 240" authorityName="Leidy" authorityPageNumber="240" authorityYear="1877" box="[189,643,1858,1885]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobates" higherTaxonomySource="CoL" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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<emphasis id="A2CA2045FF0EFF24FF7B98764CEA086B" box="[189,417,1858,1884]" italics="true" pageId="139" pageNumber="140">Myliobates jugosus</emphasis>
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–
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<treatmentCitation id="111FDA46FF0EFF24FE7898764FC8086A" author="Leidy J." box="[446,643,1858,1885]" page="240" pageId="139" pageNumber="140" year="1877">
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||
<bibRefCitation id="F42F81A6FF0EFF24FE7898764FC8086A" author="Leidy J." box="[446,643,1858,1885]" pageId="139" pageNumber="140" pagination="209 - 261" refId="ref144864" refString="Leidy J. 1877. Description of vertebrate remains, chiefly from the Phosphate Beds of South Carolina. Journal of the Academy of Natural Sciences of Philadelphia 8 (2): 209 - 261." type="journal article" year="1877">Leidy 1877: 240</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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||
, pl. 31, figs 4–5.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="9001FC57FF0EFF24FF7B98524E4508B6" blockId="139.[189,1398,1858,2027]" box="[189,782,1894,1921]" pageId="139" pageNumber="140">
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||
<treatmentCitationGroup id="B0AEDB79FF0EFF24FF7B98524E4508B6" box="[189,782,1894,1921]" pageId="139" pageNumber="140">
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||
<taxonomicName id="57BE87D4FF0EFF24FF7B98524FF308B7" authority="- Leriche 1905: 182" authorityName="Leriche" authorityPageNumber="182" authorityYear="1905" box="[189,696,1894,1921]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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<emphasis id="A2CA2045FF0EFF24FF7B98524C7108B7" box="[189,314,1894,1920]" italics="true" pageId="139" pageNumber="140">Myliobatis</emphasis>
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cf.
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<emphasis id="A2CA2045FF0EFF24FEA198534C8A08B7" box="[359,449,1895,1920]" italics="true" pageId="139" pageNumber="140">jugosus</emphasis>
|
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–
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<treatmentCitation id="111FDA46FF0EFF24FE1898524FF308B7" author="Leriche M." box="[478,696,1894,1921]" page="182" pageId="139" pageNumber="140" year="1905">
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||
<bibRefCitation id="F42F81A6FF0EFF24FE1898524FF308B7" author="Leriche M." box="[478,696,1894,1921]" pageId="139" pageNumber="140" pagination="49 - 228" refId="ref144901" refString="Leriche M. 1905. Les poissons eocenes de la Belgique. Memoires du Musee royal d'Histoire naturelle de Belgique 3 (11): 49 - 228." type="journal article" year="1905">Leriche 1905: 182</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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, pl. 52.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="9001FC57FF0EFF24FF7B98BE4FC20894" blockId="139.[189,1398,1858,2027]" box="[189,649,1929,1956]" pageId="139" pageNumber="140">
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||
<treatmentCitationGroup id="B0AEDB79FF0EFF24FF7B98BE4FC20894" box="[189,649,1929,1956]" pageId="139" pageNumber="140">
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||
<taxonomicName id="57BE87D4FF0EFF24FF7B98BE4FCE0894" authority="- Leriche 1942: 26" authorityName="Leriche" authorityPageNumber="26" authorityYear="1942" box="[189,645,1929,1956]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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<emphasis id="A2CA2045FF0EFF24FF7B98BE4CD70894" box="[189,412,1930,1956]" italics="true" pageId="139" pageNumber="140">Myliobatis jugosus</emphasis>
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–
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<treatmentCitation id="111FDA46FF0EFF24FE7F98BD4FCE0894" author="Leriche M." box="[441,645,1929,1956]" page="26" pageId="139" pageNumber="140" year="1942">
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||
<bibRefCitation id="F42F81A6FF0EFF24FE7F98BD4FCE0894" author="Leriche M." box="[441,645,1929,1956]" pageId="139" pageNumber="140" pagination="1 - 110" refId="ref145078" refString="Leriche M. 1942. Contribution a l'etude des faunes ichthyologiques marines des terrains tertiaires de la Plaine cotiere atlantique et du centre des Etats-Unis. Les synchronismes des formations tertiaires des deux cotes de l'Atlantique. Memoires de la Societe geologique de France 45 (2 - 4): 1 - 110." type="journal article" year="1942">Leriche 1942: 26</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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||
.
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||
</treatmentCitationGroup>
|
||
</paragraph>
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||
<paragraph id="9001FC57FF0EFF24FF7B98994E1708DC" blockId="139.[189,1398,1858,2027]" pageId="139" pageNumber="140">
|
||
<treatmentCitationGroup id="B0AEDB79FF0EFF24FF7B98994E1708DC" pageId="139" pageNumber="140">
|
||
<taxonomicName id="57BE87D4FF0EFF24FF7B98994FD108F0" authority="- Cappetta 1986: 189" authorityName="Cappetta" authorityPageNumber="189" authorityYear="1986" box="[189,666,1965,1992]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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<emphasis id="A2CA2045FF0EFF24FF7B98994CD308F0" box="[189,408,1965,1991]" italics="true" pageId="139" pageNumber="140">Leidybatis jugosus</emphasis>
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–
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<treatmentCitation id="111FDA46FF0EFF24FE7598994FD108F0" author="Cappetta H." box="[435,666,1965,1991]" page="189" pageId="139" pageNumber="140" year="1986">
|
||
<bibRefCitation id="F42F81A6FF0EFF24FE7598994FD108F0" author="Cappetta H." box="[435,666,1965,1991]" pageId="139" pageNumber="140" pagination="185 - 207" refId="ref137813" refString="Cappetta H. 1986. Myliobatidae nouveaux (Neoselachii, Batomorphii) de l'Ypresien des Ouled Abdoun, Maroc. Geologica et Palaeontologica 20: 185 - 207." type="journal article" year="1986">Cappetta 1986: 189</bibRefCitation>
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</treatmentCitation>
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||
</taxonomicName>
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, pl. 4c. —
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<treatmentCitation id="111FDA46FF0EFF24FCD1989A484C08F0" author="Van den Eeckhaut G. & De Schutter P." box="[791,1287,1965,1992]" page="22" pageId="139" pageNumber="140" year="2009">
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||
<bibRefCitation id="F42F81A6FF0EFF24FCD1989A484C08F0" author="Van den Eeckhaut G. & De Schutter P." box="[791,1287,1965,1992]" pageId="139" pageNumber="140" pagination="1 - 57" refId="ref152865" refString="Van den Eeckhaut G. & De Schutter P. 2009. The elasmobranch fauna of the Lede Sand Formation at Oosterzele (Lutetian, middle Eocene of Belgium). Palaeofocus 1: 1 - 57." type="journal article" year="2009">Van den Eeckhaut & De Schutter 2009: 22</bibRefCitation>
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</treatmentCitation>
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, figs 5–7.
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<taxonomicName id="57BE87D4FF0EFF24FF7B98E54FA008DC" authority="- Clayton et al. 2013: 64" authorityName="Clayton" authorityPageNumber="64" authorityYear="2013" box="[189,747,2001,2027]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="139" pageNumber="140" phylum="Chordata" rank="species" species="jugosus">
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||
<emphasis id="A2CA2045FF0EFF24FF7B98E54C7C08DC" box="[189,311,2001,2027]" italics="true" pageId="139" pageNumber="140">Leidybatis</emphasis>
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||
cf.
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||
<emphasis id="A2CA2045FF0EFF24FEA298E64CF508DC" box="[356,446,2002,2027]" italics="true" pageId="139" pageNumber="140">jugosus</emphasis>
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||
–
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||
<treatmentCitation id="111FDA46FF0EFF24FE1D98E54FA008DC" author="Clayton A. A. & Ciampaglio, C. N. & Cicimurri, D. J." box="[475,747,2001,2027]" page="64" pageId="139" pageNumber="140" year="2013">
|
||
<bibRefCitation id="F42F81A6FF0EFF24FE1D98E54FA008DC" author="Clayton A. A. & Ciampaglio, C. N. & Cicimurri, D. J." box="[475,747,2001,2027]" pageId="139" pageNumber="140" pagination="60 - 73" refId="ref139906" refString="Clayton A. A., Ciampaglio, C. N. & Cicimurri, D. J. 2013. An inquiry into the stratigraphic occurrence of a Claibornian (Eocene) vertebrate fauna from Covington County, Alabama. Bulletin Alabama Museum of Natural History 31 (2): 60 - 73." type="journal article" year="2013">
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||
Clayton
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||
<emphasis id="A2CA2045FF0EFF24FDF998E64F3108DC" box="[575,634,2001,2027]" italics="true" pageId="139" pageNumber="140">et al.</emphasis>
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2013: 64
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</bibRefCitation>
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||
</treatmentCitation>
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||
</taxonomicName>
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, pl. 4a–c.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="D8A4AFDCFF09FF23FF7B9E3F4FCE0DAA" pageId="140" pageNumber="141" type="materials_examined">
|
||
<paragraph id="9001FC57FF09FF23FF7B9E3F4CE10E12" blockId="140.[189,426,267,293]" box="[189,426,267,293]" pageId="140" pageNumber="141">
|
||
<heading id="CB494B3BFF09FF23FF7B9E3F4CE10E12" bold="true" box="[189,426,267,293]" fontSize="11" level="3" pageId="140" pageNumber="141" reason="3">
|
||
<emphasis id="A2CA2045FF09FF23FF7B9E3F4CE10E12" bold="true" box="[189,426,267,293]" pageId="140" pageNumber="141">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF09FF23FF7B9E0F4FCE0DAA" blockId="140.[189,1399,314,669]" pageId="140" pageNumber="141">
|
||
<materialsCitation id="20D6F60AFF09FF23FF7B9E0F4FCA0DAA" ID-GBIF-Occurrence="2564277966" collectionCode="ALMNH, MMNS, MSC, SC, WSU" country="United States of America" location="Claiborne Group" pageId="140" pageNumber="141" specimenCode="NJSM 24033, WSU 18, WSU 30, CC 513, CC 522, WSU 5009" specimenCount="46" stateProvince="Alabama">
|
||
<collectingCountry id="E8A9BCC7FF09FF23FF7B9E0F4F270E62" box="[189,620,315,341]" name="United States of America" pageId="140" pageNumber="141">UNITED STATES OF AMERICA</collectingCountry>
|
||
–
|
||
<emphasis id="A2CA2045FF09FF23FD5D9E0E4E460E63" bold="true" box="[667,781,314,340]" pageId="140" pageNumber="141">
|
||
<collectingRegion id="527A32B5FF09FF23FD5D9E0E4E460E63" box="[667,781,314,340]" country="United States of America" name="Alabama" pageId="140" pageNumber="141">Alabama</collectingRegion>
|
||
</emphasis>
|
||
• 71 isolated teeth;
|
||
<location id="9561AA8CFF09FF23FBE29E0F49B30E62" LSID="urn:lsid:plazi:treatment:18174D41FF0EFF20FD9099AF483C0D37:9561AA8CFF09FF23FBE29E0F49B30E62" box="[1060,1272,315,341]" country="United States of America" name="Claiborne Group" pageId="140" pageNumber="141" stateProvince="Alabama">Claiborne Group</location>
|
||
;
|
||
<collectionCode id="F6AF6492FF09FF23FACD9E0F483C0E62" box="[1291,1399,315,341]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="140" pageNumber="141" type="Museum">ALMNH</collectionCode>
|
||
PV1992.28.21,
|
||
<collectionCode id="F6AF6492FF09FF23FEB89E6B4CA10E4E" box="[382,490,351,377]" country="USA" httpUri="http://grbio.org/cool/93n4-xhed" name="Alabama Museum of Natural History" pageId="140" pageNumber="141" type="Museum">ALMNH</collectionCode>
|
||
PV1992.28.6.3,
|
||
<collectionCode id="F6AF6492FF09FF23FD019E6B4E6A0E4E" box="[711,801,351,377]" pageId="140" pageNumber="141">MMNS</collectionCode>
|
||
VP-5643 (
|
||
<specimenCount id="86B837DEFF09FF23FC719E6B49130E4D" box="[951,1112,351,378]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FBB09E6B499B0E4E" box="[1142,1232,351,377]" pageId="140" pageNumber="141">MMNS</collectionCode>
|
||
VP-8189 (
|
||
<specimenCount id="86B837DEFF09FF23FAA19E6B4C770EA9" pageId="140" pageNumber="141" type="generic">6 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FE949EB04CE70EA9" box="[338,428,388,414]" pageId="140" pageNumber="141">MMNS</collectionCode>
|
||
VP-8224 (
|
||
<specimenCount id="86B837DEFF09FF23FDF79EB74F820EA9" box="[561,713,387,414]" pageId="140" pageNumber="141" type="generic">5 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FD199EB04E560EA9" box="[735,797,388,414]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
35005,
|
||
<collectionCode id="F6AF6492FF09FF23FC469EB04EF50EA9" box="[896,958,388,414]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37080,
|
||
<collectionCode id="F6AF6492FF09FF23FBE49EB0492B0EA9" box="[1058,1120,388,414]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37081,
|
||
<collectionCode id="F6AF6492FF09FF23FB029EB048490EA9" box="[1220,1282,388,414]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37082 (
|
||
<specimenCount id="86B837DEFF09FF23FAA19EB04C770EF4" pageId="140" pageNumber="141" type="generic">4 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FE929E9C4CD90EF5" box="[340,402,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37083,
|
||
<collectionCode id="F6AF6492FF09FF23FE3C9E9C4F730EF5" box="[506,568,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37084,
|
||
<collectionCode id="F6AF6492FF09FF23FD669E9C4F950EF5" box="[672,734,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37085,
|
||
<collectionCode id="F6AF6492FF09FF23FC809E9C4ECF0EF5" box="[838,900,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37086,
|
||
<collectionCode id="F6AF6492FF09FF23FC2A9E9C49610EF5" box="[1004,1066,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37135,
|
||
<collectionCode id="F6AF6492FF09FF23FB549E9C499B0EF5" box="[1170,1232,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37135,
|
||
<collectionCode id="F6AF6492FF09FF23FAFE9E9C483D0EF5" box="[1336,1398,424,450]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37249 (
|
||
<specimenCount id="86B837DEFF09FF23FED19EF84CE60ED0" box="[279,429,460,487]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FE049EF84F4B0ED1" box="[450,512,460,486]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37256,
|
||
<collectionCode id="F6AF6492FF09FF23FDA69EF84FD50ED1" box="[608,670,460,486]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37257 (
|
||
<specimenCount id="86B837DEFF09FF23FCC79EF84EDD0ED0" box="[769,918,460,487]" pageId="140" pageNumber="141" type="generic">3 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FC6D9EF84EA20ED1" box="[939,1001,460,486]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37282,
|
||
<collectionCode id="F6AF6492FF09FF23FB8F9EF849CC0ED1" box="[1097,1159,460,486]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37291,
|
||
<collectionCode id="F6AF6492FF09FF23FB219EF8486E0ED1" box="[1255,1317,460,486]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37321 (
|
||
<specimenCount id="86B837DEFF09FF23FF019EC54C100D3B" box="[199,347,497,524]" pageId="140" pageNumber="141" type="generic">3 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FEAB9EC54CE00D3C" box="[365,427,497,523]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
37681,
|
||
<collectionCode id="F6AF6492FF09FF23FDCE9EC54F0D0D3C" box="[520,582,497,523]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
38829 (
|
||
<specimenCount id="86B837DEFF09FF23FD609EC54E710D3B" box="[678,826,497,524]" pageId="140" pageNumber="141" type="generic">3 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FC8A9EC54EC10D3C" box="[844,906,497,523]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
38876 (
|
||
<specimenCount id="86B837DEFF09FF23FC2F9EC549350D3B" box="[1001,1150,497,524]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FB569EC549850D3C" box="[1168,1230,497,523]" pageId="140" pageNumber="141">MSC</collectionCode>
|
||
38948,
|
||
<specimenCode id="C018542CFF09FF23FAED9EC54C420D18" collectionCode="NJSM" country="USA" httpUri="http://grbio.org/cool/fe4d-5yka" name="New Jersey State Museum" pageId="140" pageNumber="141">
|
||
<collectionCode id="F6AF6492FF09FF23FAED9EC5483D0D3C" box="[1323,1398,497,523]" pageId="140" pageNumber="141">NJSM</collectionCode>
|
||
24033
|
||
</specimenCode>
|
||
,
|
||
<collectionCode id="F6AF6492FF09FF23FEDE9D214C700D18" box="[280,315,533,559]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.16,
|
||
<collectionCode id="F6AF6492FF09FF23FE149D214CBF0D18" box="[466,500,533,559]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.17 (
|
||
<specimenCount id="86B837DEFF09FF23FD489D214E6C0D07" box="[654,807,533,560]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FCF89D214E2B0D18" box="[830,864,533,559]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.197 (
|
||
<specimenCount id="86B837DEFF09FF23FBCF9D2149E90D07" box="[1033,1186,533,560]" pageId="140" pageNumber="141" type="generic">3 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FB7F9D2149970D18" box="[1209,1244,533,559]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.198,
|
||
<collectionCode id="F6AF6492FF09FF23FF7B9D0E4DAB0D63" box="[189,224,570,596]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.199,
|
||
<collectionCode id="F6AF6492FF09FF23FE4B9D0E4CFB0D63" box="[397,432,570,596]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.200,
|
||
<collectionCode id="F6AF6492FF09FF23FD989D0E4FCA0D63" box="[606,641,570,596]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.201 (
|
||
<specimenCount id="86B837DEFF09FF23FCF69D0E4E9A0D62" box="[816,977,570,597]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FC369D0E49580D63" box="[1008,1043,570,596]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.202,
|
||
<collectionCode id="F6AF6492FF09FF23FB069D0E49A90D63" box="[1216,1250,570,596]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.203 (
|
||
<specimenCount id="86B837DEFF09FF23FF009D6A4C2B0D4E" box="[198,352,606,633]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FEB19D6A4CD10D4F" box="[375,410,606,632]" pageId="140" pageNumber="141">SC</collectionCode>
|
||
2012.47.204,
|
||
<specimenCode id="C018542CFF09FF23FD869D6A4FE00D4F" box="[576,683,606,632]" collectionCode="WSU" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">
|
||
<collectionCode id="F6AF6492FF09FF23FD869D6A4FCA0D4F" box="[576,641,606,632]" pageId="140" pageNumber="141">WSU</collectionCode>
|
||
18
|
||
</specimenCode>
|
||
,
|
||
<specimenCode id="C018542CFF09FF23FD7A9D6A4E630D4F" box="[700,808,606,632]" collectionCode="WSU" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">
|
||
<collectionCode id="F6AF6492FF09FF23FD7A9D6A4FB60D4F" box="[700,765,606,632]" pageId="140" pageNumber="141">WSU</collectionCode>
|
||
30
|
||
</specimenCode>
|
||
,
|
||
<collectionCode id="F6AF6492FF09FF23FCFE9D6A4E320D4F" box="[824,889,606,632]" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">WSU</collectionCode>
|
||
<specimenCode id="C018542CFF09FF23FC429D6A4EA80D4E" box="[900,995,606,633]" collectionCode="WSU" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">CC 513</specimenCode>
|
||
(
|
||
<specimenCount id="86B837DEFF09FF23FC319D6A49DA0D4E" box="[1015,1169,606,633]" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<collectionCode id="F6AF6492FF09FF23FB6E9D6A49A20D4F" box="[1192,1257,606,632]" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">WSU</collectionCode>
|
||
<specimenCode id="C018542CFF09FF23FB329D6A48180D4E" box="[1268,1363,606,633]" collectionCode="WSU" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">CC 522</specimenCode>
|
||
(
|
||
<specimenCount id="86B837DEFF09FF23FAA19D6A4C770DAA" pageId="140" pageNumber="141" type="generic">2 specimens</specimenCount>
|
||
),
|
||
<specimenCode id="C018542CFF09FF23FE899DB74C980DAA" box="[335,467,642,669]" collectionCode="WSU" country="USA" httpUri="http://grbio.org/cool/1raa-g3is" name="Weber State University, Bird and Mammal Collection" pageId="140" pageNumber="141">
|
||
<collectionCode id="F6AF6492FF09FF23FE899DB74CDB0DAA" box="[335,400,643,669]" pageId="140" pageNumber="141">WSU</collectionCode>
|
||
5009
|
||
</specimenCode>
|
||
(
|
||
<specimenCount id="86B837DEFF09FF23FE259DB74F320DAA" box="[483,633,642,669]" pageId="140" pageNumber="141" type="generic">3 specimens</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8A4AFDCFF09FF23FF7B9DF1491F0B97" pageId="140" pageNumber="141" type="description">
|
||
<paragraph id="9001FC57FF09FF23FF7B9DF14C050DE8" blockId="140.[189,334,709,735]" box="[189,334,709,735]" pageId="140" pageNumber="141">
|
||
<heading id="CB494B3BFF09FF23FF7B9DF14C050DE8" bold="true" box="[189,334,709,735]" fontSize="11" level="3" pageId="140" pageNumber="141" reason="3">
|
||
<emphasis id="A2CA2045FF09FF23FF7B9DF14C050DE8" bold="true" box="[189,334,709,735]" pageId="140" pageNumber="141">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF09FF23FF7B9DC1491F0B97" blockId="140.[189,1399,757,1184]" pageId="140" pageNumber="141">
|
||
Median teeth very wide and narrow. All median teeth with distinctive swelling on the crown; swelling generally located medially but can be closer to the lateral margin (resulting in an asymmetrical labial/ lingual outline). Swelling is in contrast to remainder of crown, which is dorsoventrally very thin. Occlusal surface has thick enameloid covering that is heavily tuberculated on unworn parts of tooth; surface is punctate where enameloid is missing, due to
|
||
<emphasis id="A2CA2045FF09FF23FC849CBC4EDA0C96" box="[834,913,904,929]" italics="true" pageId="140" pageNumber="141">in vivo</emphasis>
|
||
use or taphonomic processes (
|
||
<emphasis id="A2CA2045FF09FF23FB3F9CBC48030C96" box="[1273,1352,904,929]" italics="true" pageId="140" pageNumber="141">in vivo</emphasis>
|
||
use is indicated by flat wear facets that on the highly convex portion of the crown of some teeth). Labial crown face overhangs the root; lingual face with thick and rounded transverse ridge at the crown base. Labial and lingual crown faces possess coarse parallel vertical folds that anastomose and become finer apically. Distal ends of crown angular. Root at least as high as thinnest part of crown; is subdivided into thin lamellae by evenly spaced nutritive grooves. Lamellae do not extend lingually past the crown. Lateral teeth are six-sided, nearly symmetrical. Lateral teeth with same tuberculation and wrinkling as observed on median teeth, but root subdivided into only two or three lamellae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D8A4AFDCFF09FF20FF7B9BFD483C0D37" lastPageId="143" lastPageNumber="144" pageId="140" pageNumber="141" type="discussion">
|
||
<paragraph id="9001FC57FF09FF23FF7B9BFD4C640BD4" blockId="140.[189,303,1225,1251]" box="[189,303,1225,1251]" pageId="140" pageNumber="141">
|
||
<heading id="CB494B3BFF09FF23FF7B9BFD4C640BD4" bold="true" box="[189,303,1225,1251]" fontSize="11" level="3" pageId="140" pageNumber="141" reason="3">
|
||
<emphasis id="A2CA2045FF09FF23FF7B9BFD4C640BD4" bold="true" box="[189,303,1225,1251]" pageId="140" pageNumber="141">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF09FF23FF7B9BCC4F1D08DC" blockId="140.[189,1399,1272,2027]" pageId="140" pageNumber="141">
|
||
Ascribing the teeth described above to a particular species is complicated by the fact that fossil
|
||
<taxonomicName id="57BE87D4FF09FF23FF7B9A284C120A01" authorityName="Bonaparte" authorityYear="1838" box="[189,345,1308,1334]" class="Elasmobranchii" family="Myliobatidae" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="family">Myliobatidae</taxonomicName>
|
||
are often based on a single median tooth, and our knowledge of how the tooth was incorporated into the upper or lower dentition is far from complete. Whole and even partial upper and lower dentitions are more informative, but as
|
||
<bibRefCitation id="F42F81A6FF09FF23FD0A9A5149310A48" author="Hovestadt D. C. & Hovestadt-Euler M." box="[716,1146,1381,1407]" pageId="140" pageNumber="141" pagination="1 - 66" refId="ref143545" refString="Hovestadt D. C. & Hovestadt-Euler M. 2013. Generic assessment and reallocation of Cenozoic Myliobatinae based on new information of tooth, tooth plate and caudal spine morphology of extant taxa. Palaeontos 24: 1 - 66." type="journal article" year="2013">Hovestadt & Hovestadt-Euler (2013)</bibRefCitation>
|
||
have recently shown, there can be a great deal of intraspecific variation within the dentitions of extant species. Several fossil species based on isolated median teeth, including
|
||
<taxonomicName id="57BE87D4FF09FF23FCD49A9A49DE0AFF" authority="Salinas, 1901" authorityName="Salinas" authorityYear="1901" box="[786,1173,1454,1480]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="siculus">
|
||
<emphasis id="A2CA2045FF09FF23FCD49A9A4EA20AFF" box="[786,1001,1454,1480]" italics="true" pageId="140" pageNumber="141">Myliobatis siculus</emphasis>
|
||
Salinas, 1901
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="57BE87D4FF09FF23FB659A9B4C170ADA" authority="Fowler, 1911" authorityName="Fowler" authorityYear="1911" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="pachyrhizodus">
|
||
<emphasis id="A2CA2045FF09FF23FB659A9B483C0AFF" box="[1187,1399,1454,1480]" italics="true" pageId="140" pageNumber="141">M. pachyrhizodus</emphasis>
|
||
<bibRefCitation id="F42F81A6FF09FF23FF7B9AE74C170ADA" author="Fowler H. W." box="[189,348,1491,1517]" pageId="140" pageNumber="141" pagination="1 - 182" refId="ref141918" refString="Fowler H. W. 1911. A description of the fossil fish remains of the Cretaceous, Eocene and Miocene formations of New-Jersey. Geological Survey of New Jersey Bulletin 4: 1 - 182. https: // doi. org / 10.5962 / bhl. title. 63936" type="journal article" year="1911">Fowler, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="57BE87D4FF09FF23FEAA9AE04F930ADA" authority="Woodward, 1889" authorityName="Woodward" authorityYear="1889" box="[364,728,1491,1517]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="tumidens">
|
||
<emphasis id="A2CA2045FF09FF23FEAA9AE04CB40ADA" box="[364,511,1491,1517]" italics="true" pageId="140" pageNumber="141">M. tumidens</emphasis>
|
||
<bibRefCitation id="F42F81A6FF09FF23FDCC9AE74F930ADA" author="Woodward A. S." box="[522,728,1491,1517]" pageId="140" pageNumber="141" refId="ref154310" refString="Woodward A. S. 1889. Catalogue of the Fossil Fishes in the British Museum. Part. I. British Museum (Natural History), London. https: // doi. org / 10.5962 / bhl. title. 61854" type="book" year="1889">Woodward, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="57BE87D4FF09FF23FCDA9AE049700ADA" authority="Leidy, 1876" authorityName="Leidy" authorityYear="1876" box="[796,1083,1491,1517]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF09FF23FCDA9AE04EEB0ADA" box="[796,928,1492,1517]" italics="true" pageId="140" pageNumber="141">M. jugosus</emphasis>
|
||
<bibRefCitation id="F42F81A6FF09FF23FC6C9AE749700ADA" author="Leidy J." box="[938,1083,1491,1517]" pageId="140" pageNumber="141" pagination="86 - 87" refId="ref144835" refString="Leidy J. 1876. Remarks on fossils of the Ashley Phosphate Beds. Proceedings of the Academy of Natural Sciences of Philadelphia 28: 86 - 87." type="journal article" year="1876">Leidy, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
, are very convex, but this phenomenon can largely be attributed to the teeth being from the upper dentition (i.e., see description for
|
||
<taxonomicName id="57BE87D4FF09FF23FF2E99284C350901" authorityName="Garman" authorityYear="1908" box="[232,382,1564,1590]" class="Elasmobranchii" family="Myliobatidae" genus="Aetomylaeus" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FF2E99284C350901" box="[232,382,1564,1590]" italics="true" pageId="140" pageNumber="141">Aetomylaeus</emphasis>
|
||
</taxonomicName>
|
||
above).
|
||
<bibRefCitation id="F42F81A6FF09FF23FE23992F4FDF0901" author="Leidy J." box="[485,660,1563,1590]" pageId="140" pageNumber="141" pagination="86 - 87" refId="ref144835" refString="Leidy J. 1876. Remarks on fossils of the Ashley Phosphate Beds. Proceedings of the Academy of Natural Sciences of Philadelphia 28: 86 - 87." type="journal article" year="1876">Leidy’s (1876)</bibRefCitation>
|
||
taxon, the
|
||
<taxonomicName id="57BE87D4FF09FF23FCDC99284E3F0902" authorityName="Leidy" authorityYear="1876" box="[794,884,1564,1589]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF09FF23FCDC99284E3F0902" box="[794,884,1564,1589]" italics="true" pageId="140" pageNumber="141">jugosus</emphasis>
|
||
</taxonomicName>
|
||
morphology, has unusually convex median teeth like the ones from the
|
||
<collectingRegion id="527A32B5FF09FF23FDCD99744F17096D" box="[523,604,1600,1626]" country="Portugal" name="Lisboa" pageId="140" pageNumber="141">Lisbon</collectingRegion>
|
||
Formation we described above.
|
||
<bibRefCitation id="F42F81A6FF09FF23FC27997449EB096C" author="Cappetta H." box="[993,1184,1600,1627]" pageId="140" pageNumber="141" pagination="185 - 207" refId="ref137813" refString="Cappetta H. 1986. Myliobatidae nouveaux (Neoselachii, Batomorphii) de l'Ypresien des Ouled Abdoun, Maroc. Geologica et Palaeontologica 20: 185 - 207." type="journal article" year="1986">Cappetta (1986)</bibRefCitation>
|
||
erected the genus
|
||
<taxonomicName id="57BE87D4FF09FF23FF7B99504C7C0949" authorityName="Cappetta" authorityYear="1986" box="[189,311,1636,1662]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FF7B99504C7C0949" box="[189,311,1636,1662]" italics="true" pageId="140" pageNumber="141">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
to include the
|
||
<taxonomicName id="57BE87D4FF09FF23FE3399514F040949" authorityName="Leidy" authorityYear="1876" box="[501,591,1637,1662]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF09FF23FE3399514F040949" box="[501,591,1637,1662]" italics="true" pageId="140" pageNumber="141">jugosus</emphasis>
|
||
</taxonomicName>
|
||
morphology, as well as his new species,
|
||
<taxonomicName id="57BE87D4FF09FF23FB88995149B80949" box="[1102,1267,1636,1662]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="granulosus">
|
||
<emphasis id="A2CA2045FF09FF23FB88995149B80949" box="[1102,1267,1636,1662]" italics="true" pageId="140" pageNumber="141">L. granulosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="F42F81A6FF09FF23FAC099504C400994" author="Mendiola C." pageId="140" pageNumber="141" pagination="1 - 42" refId="ref146654" refString="Mendiola C. 1999. Myliobatoideos nuevos (Neoselachii, Batomorphii) del Thanetiense de Oed Zem (Cuenca de los Ouled Abdoun, Marruecos). Revista de la Societat Paleontologica d'Elx 6: 1 - 42." type="journal article" year="1999">Mendiola (1999)</bibRefCitation>
|
||
named several species of
|
||
<taxonomicName id="57BE87D4FF09FF23FD8499BD4FF70994" authorityName="Cappetta" authorityYear="1986" box="[578,700,1673,1699]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FD8499BD4FF70994" box="[578,700,1673,1699]" italics="true" pageId="140" pageNumber="141">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
, but
|
||
<bibRefCitation id="F42F81A6FF09FF23FD3F99BD4EFC0994" author="Cappetta H." box="[761,951,1673,1699]" pageId="140" pageNumber="141" refId="ref137948" refString="Cappetta H. 2012. Chondrichthyes (Mesozoic and Cenozoic Elasmobranchii, Teeth). Handbook of Palaeoichthyology. Verlag Friedrich Pfeil, Munchen." type="book" year="2012">Cappetta (2012)</bibRefCitation>
|
||
considered those species as a
|
||
<emphasis id="A2CA2045FF09FF23FAD899BE4C4B09F0" italics="true" pageId="140" pageNumber="141">nomina dubia</emphasis>
|
||
because they lacked stratigraphic context.
|
||
<bibRefCitation id="F42F81A6FF09FF23FCCB9999498209F0" author="Hovestadt D. C. & Hovestadt-Euler M." box="[781,1225,1709,1735]" pageId="140" pageNumber="141" pagination="1 - 66" refId="ref143545" refString="Hovestadt D. C. & Hovestadt-Euler M. 2013. Generic assessment and reallocation of Cenozoic Myliobatinae based on new information of tooth, tooth plate and caudal spine morphology of extant taxa. Palaeontos 24: 1 - 66." type="journal article" year="2013">Hovestadt & Hovestadt-Euler (2013)</bibRefCitation>
|
||
indicated that the various tooth morphologies attributed to
|
||
<taxonomicName id="57BE87D4FF09FF23FD1F99E64E1809DB" authorityName="Cappetta" authorityYear="1986" box="[729,851,1746,1772]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FD1F99E64E1809DB" box="[729,851,1746,1772]" italics="true" pageId="140" pageNumber="141">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
fall within the range they observed in extant species of
|
||
<taxonomicName id="57BE87D4FF09FF23FEFD99C24C9A0827" authorityName="Garman" authorityYear="1908" box="[315,465,1782,1808]" class="Elasmobranchii" family="Myliobatidae" genus="Aetomylaeus" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FEFD99C24C9A0827" box="[315,465,1782,1808]" italics="true" pageId="140" pageNumber="141">Aetomylaeus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="57BE87D4FF09FF23FDCA99C24FC20827" authorityName="Cuvier" authorityYear="1816" box="[524,649,1782,1808]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FDCA99C24FC20827" box="[524,649,1782,1808]" italics="true" pageId="140" pageNumber="141">Myliobatis</emphasis>
|
||
</taxonomicName>
|
||
, and they considered
|
||
<taxonomicName id="57BE87D4FF09FF23FC4899C349430827" authorityName="Leidy" authorityYear="1876" box="[910,1032,1783,1808]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF09FF23FC4899C349430827" box="[910,1032,1783,1808]" italics="true" pageId="140" pageNumber="141">L. jugosus</emphasis>
|
||
</taxonomicName>
|
||
as a
|
||
<emphasis id="A2CA2045FF09FF23FB8099C349BE0827" box="[1094,1269,1782,1808]" italics="true" pageId="140" pageNumber="141">nomen dubium</emphasis>
|
||
. However,
|
||
<taxonomicName id="57BE87D4FF09FF23FF7B982F4C7C0802" authorityName="Cappetta" authorityYear="1986" box="[189,311,1819,1845]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FF7B982F4C7C0802" box="[189,311,1819,1845]" italics="true" pageId="140" pageNumber="141">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
possess labial and lingual ornamentation of parallel vertical folds, which contrasts with the pitting and granulation seen on these areas of
|
||
<taxonomicName id="57BE87D4FF09FF23FD14980B4E23086E" authorityName="Garman" authorityYear="1908" box="[722,872,1855,1881]" class="Elasmobranchii" family="Myliobatidae" genus="Aetomylaeus" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FD14980B4E23086E" box="[722,872,1855,1881]" italics="true" pageId="140" pageNumber="141">Aetomylaeus</emphasis>
|
||
</taxonomicName>
|
||
teeth (
|
||
<bibRefCitation id="F42F81A6FF09FF23FC71980B4909086E" author="Leidy J." box="[951,1090,1855,1881]" pageId="140" pageNumber="141" pagination="209 - 261" refId="ref144864" refString="Leidy J. 1877. Description of vertebrate remains, chiefly from the Phosphate Beds of South Carolina. Journal of the Academy of Natural Sciences of Philadelphia 8 (2): 209 - 261." type="journal article" year="1877">Leidy 1877</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F42F81A6FF09FF23FB8B980B49B1086E" author="Cappetta H." box="[1101,1274,1855,1881]" pageId="140" pageNumber="141" pagination="185 - 207" refId="ref137813" refString="Cappetta H. 1986. Myliobatidae nouveaux (Neoselachii, Batomorphii) de l'Ypresien des Ouled Abdoun, Maroc. Geologica et Palaeontologica 20: 185 - 207." type="journal article" year="1986">Cappetta 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F42F81A6FF09FF23FAC0980B4DB00849" author="Mendiola C." pageId="140" pageNumber="141" pagination="1 - 42" refId="ref146654" refString="Mendiola C. 1999. Myliobatoideos nuevos (Neoselachii, Batomorphii) del Thanetiense de Oed Zem (Cuenca de los Ouled Abdoun, Marruecos). Revista de la Societat Paleontologica d'Elx 6: 1 - 42." type="journal article" year="1999">Mendiola 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F42F81A6FF09FF23FECF98504CF20849" author="Cappetta H." box="[265,441,1892,1918]" pageId="140" pageNumber="141" refId="ref137948" refString="Cappetta H. 2012. Chondrichthyes (Mesozoic and Cenozoic Elasmobranchii, Teeth). Handbook of Palaeoichthyology. Verlag Friedrich Pfeil, Munchen." type="book" year="2012">Cappetta 2012</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="F42F81A6FF09FF23FE0198574E3B0849" author="Hovestadt D. C. & Hovestadt-Euler M." box="[455,880,1891,1918]" pageId="140" pageNumber="141" pagination="1 - 66" refId="ref143545" refString="Hovestadt D. C. & Hovestadt-Euler M. 2013. Generic assessment and reallocation of Cenozoic Myliobatinae based on new information of tooth, tooth plate and caudal spine morphology of extant taxa. Palaeontos 24: 1 - 66." type="journal article" year="2013">Hovestadt & Hovestadt-Euler 2013</bibRefCitation>
|
||
). In addition, the labial root face is vertical on
|
||
<taxonomicName id="57BE87D4FF09FF23FF2598BC4C160895" authorityName="Cappetta" authorityYear="1986" box="[227,349,1928,1954]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FF2598BC4C160895" box="[227,349,1928,1954]" italics="true" pageId="140" pageNumber="141">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
teeth and root lobes do not extend lingually past the crown base. This contrasts with the roots of
|
||
<taxonomicName id="57BE87D4FF09FF23FED898984CD008F1" authorityName="Cuvier" authorityYear="1816" box="[286,411,1964,1990]" class="Elasmobranchii" family="Myliobatidae" genus="Myliobatis" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FED898984CD008F1" box="[286,411,1964,1990]" italics="true" pageId="140" pageNumber="141">Myliobatis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="57BE87D4FF09FF23FE1398984F2008F1" authorityName="Garman" authorityYear="1908" box="[469,619,1964,1990]" class="Elasmobranchii" family="Myliobatidae" genus="Aetomylaeus" kingdom="Animalia" order="Myliobatiformes" pageId="140" pageNumber="141" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF09FF23FE1398984F2008F1" box="[469,619,1964,1990]" italics="true" pageId="140" pageNumber="141">Aetomylaeus</emphasis>
|
||
</taxonomicName>
|
||
, which have oblique labial faces and the lobes very conspicuously extend past the lingual crown face.
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF08FF22FF7B9E3F49660EC9" blockId="141.[189,1399,267,510]" pageId="141" pageNumber="142">
|
||
The root characteristics of
|
||
<taxonomicName id="57BE87D4FF08FF22FE289E3F4F230E12" authorityName="Cappetta" authorityYear="1986" box="[494,616,267,293]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FE289E3F4F230E12" box="[494,616,267,293]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
are similar to the condition seen on
|
||
<taxonomicName id="57BE87D4FF08FF22FC399E3F49C80E12" box="[1023,1155,267,293]" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FC399E3F49C80E12" box="[1023,1155,267,293]" italics="true" pageId="141" pageNumber="142">Rhinoptera</emphasis>
|
||
</taxonomicName>
|
||
teeth, and specimens illustrated by
|
||
<bibRefCitation id="F42F81A6FF08FF22FEA29E044F6B0E7E" author="Cappetta H." box="[356,544,303,330]" pageId="141" pageNumber="142" pagination="185 - 207" refId="ref137813" refString="Cappetta H. 1986. Myliobatidae nouveaux (Neoselachii, Batomorphii) de l'Ypresien des Ouled Abdoun, Maroc. Geologica et Palaeontologica 20: 185 - 207." type="journal article" year="1986">Cappetta (1986</bibRefCitation>
|
||
: fig. 3, 1–8) indicate that
|
||
<taxonomicName id="57BE87D4FF08FF22FC9A9E04494B0E7E" box="[860,1024,303,329]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="species" species="granulosus">
|
||
<emphasis id="A2CA2045FF08FF22FC9A9E04494B0E7E" box="[860,1024,303,329]" italics="true" pageId="141" pageNumber="142">L. granulosus</emphasis>
|
||
</taxonomicName>
|
||
had lateral teeth of decreasing width like
|
||
<taxonomicName id="57BE87D4FF08FF22FEFD9E604CF40E59" box="[315,447,340,366]" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FEFD9E604CF40E59" box="[315,447,340,366]" italics="true" pageId="141" pageNumber="142">Rhinoptera</emphasis>
|
||
</taxonomicName>
|
||
. In support of this is a
|
||
<taxonomicName id="57BE87D4FF08FF22FD179E604E000E59" authorityName="Cappetta" authorityYear="1986" box="[721,843,340,366]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD179E604E000E59" box="[721,843,340,366]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
lateral tooth morphology that is comparable to those within more proximal lateral rows of a
|
||
<taxonomicName id="57BE87D4FF08FF22FD1C9E4C4E150EA5" box="[730,862,376,402]" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD1C9E4C4E150EA5" box="[730,862,376,402]" italics="true" pageId="141" pageNumber="142">Rhinoptera</emphasis>
|
||
</taxonomicName>
|
||
dentition. SC2012.47.202 (
|
||
<figureCitation id="0885E0D2FF08FF22FB619E4C49B60EA5" box="[1191,1277,375,402]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="141" pageNumber="142">Fig. 52</figureCitation>
|
||
SS–TT) is twice as wide as long and is similar to
|
||
<taxonomicName id="57BE87D4FF08FF22FD409EA84E410E81" box="[646,778,412,438]" class="Elasmobranchii" family="Myliobatidae" genus="Rhinoptera" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD409EA84E410E81" box="[646,778,412,438]" italics="true" pageId="141" pageNumber="142">Rhinoptera</emphasis>
|
||
</taxonomicName>
|
||
lateral teeth in our Claiborne sample. We assign this tooth to
|
||
<taxonomicName id="57BE87D4FF08FF22FEE29EF44CD50EED" authorityName="Cappetta" authorityYear="1986" box="[292,414,448,474]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FEE29EF44CD50EED" box="[292,414,448,474]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
based on the distinctly ornamented occlusal surface and lingual and labial crown ornamentation that is identical to that on the median teeth described above.
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF08FF22FF7B9D134EE70CF9" blockId="141.[189,1399,550,974]" pageId="141" pageNumber="142">
|
||
We herein support the validity of
|
||
<taxonomicName id="57BE87D4FF08FF22FD8B9D124F8C0D77" authorityName="Cappetta" authorityYear="1986" box="[589,711,550,576]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD8B9D124F8C0D77" box="[589,711,550,576]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
based on our samples of
|
||
<taxonomicName id="57BE87D4FF08FF22FC3F9D1249DE0D77" authorityName="Bonaparte" authorityYear="1835" box="[1017,1173,550,576]" class="Elasmobranchii" family="Myliobatidae" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="subFamily" subFamily="Myliobatinae">Myliobatinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="57BE87D4FF08FF22FB169D12483D0D77" authorityName="Jordan & Evermann" authorityYear="1896" box="[1232,1398,550,576]" class="Elasmobranchii" family="Rhinopteridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="subFamily" subFamily="Rhinopterinae">Rhinopterinae</taxonomicName>
|
||
teeth from the Tallahatta and
|
||
<collectingRegion id="527A32B5FF08FF22FDE49D7E4F380D53" box="[546,627,586,612]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formations. As mentioned above, the labial and lingual vertical wrinkling on
|
||
<taxonomicName id="57BE87D4FF08FF22FE9D9D5A4C9E0DBF" authorityName="Cappetta" authorityYear="1986" box="[347,469,622,648]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FE9D9D5A4C9E0DBF" box="[347,469,622,648]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
teeth contrasts with the ornamentation we observed on
|
||
<taxonomicName id="57BE87D4FF08FF22FBA69D5A49BD0DBF" authorityName="Garman" authorityYear="1908" box="[1120,1270,622,648]" class="Elasmobranchii" family="Myliobatidae" genus="Aetomylaeus" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FBA69D5A49BD0DBF" box="[1120,1270,622,648]" italics="true" pageId="141" pageNumber="142">Aetomylaeus</emphasis>
|
||
</taxonomicName>
|
||
teeth from both formations. If one considered the
|
||
<taxonomicName id="57BE87D4FF08FF22FD4F9DA64E480D9B" authorityName="Cappetta" authorityYear="1986" box="[649,771,658,684]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD4F9DA64E480D9B" box="[649,771,658,684]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
morphology aberrant, as do Hovestadt & Hovestadt- Euler (2013), the labial and lingual ornamentation of teeth should still serve as a generic identifier. As an example, we examined a partial
|
||
<taxonomicName id="57BE87D4FF08FF22FD9F9DEF4FA40DC2" authorityName="Garman" authorityYear="1908" box="[601,751,731,757]" class="Elasmobranchii" family="Myliobatidae" genus="Aetomylaeus" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD9F9DEF4FA40DC2" box="[601,751,731,757]" italics="true" pageId="141" pageNumber="142">Aetomylaeus</emphasis>
|
||
</taxonomicName>
|
||
median tooth (MSC 37079,
|
||
<figureCitation id="0885E0D2FF08FF22FBFB9DEF49C40DC1" box="[1085,1167,731,758]" captionStart="Fig" captionStartId="126.[189,232,1645,1671]" captionTargetBox="[225,1363,263,1613]" captionTargetId="figure@126.[215,1374,248,1633]" captionTargetPageId="126" captionText="Fig. 46. Aetomylaeus sp., teeth. A–E. MSC 38836, lower median tooth, “upper” Lisbon Formation. A. Oral view. B. Basal view. C. Lingual view. D. Labial view. E. Profile view. F–J. MSC 199.2, median tooth, basal Gosport Sand. F. Oral view. G. Basal view. H. Lingual view. I. Labial view. J. Profile view. K–O. MSC 199.1, median tooth, basal Gosport Sand. K. Oral view. L. Basal view. M. Lingual view. N. Labial view. O. Profile view. P–T. MSC 38872, lower median tooth, Gosport Sand. P. Oral view. Q. Basal view. R. Lingual view. S. Labial view. T. Profile view. U–Y. MSC 37910, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. U. Oral view. V. Basal view. W. Mesial view. X. Distal view. Y. Labial view. Z–DD. MSC 38851, lateral tooth, basal Gosport Sand. Z. Oral view. AA. Basal view. BB. Mesial view. CC. Distal view. DD. Labial view. EE–II. MSC 37079, upper median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. EE. Oral view. FF. Basal view. GG. Lingual view. HH. Labial view. II. Profile view. Labial at the top in oral or basal views of the median teeth. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660349" httpUri="https://zenodo.org/record/3660349/files/figure.png" pageId="141" pageNumber="142">Fig. 46</figureCitation>
|
||
EE–II) that exhibits a convex medial portion but thinner lateral side, similar to
|
||
<taxonomicName id="57BE87D4FF08FF22FCB99DCB4EB20C2E" authorityName="Cappetta" authorityYear="1986" box="[895,1017,767,793]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FCB99DCB4EB20C2E" box="[895,1017,767,793]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
. The vertical wrinkling seen on
|
||
<taxonomicName id="57BE87D4FF08FF22FF7B9C174FC30C09" authority="(Tallahatta Formation)" baseAuthorityName="Tallahatta Formation" box="[189,648,803,830]" class="Elasmobranchii" family="Myliobatidae" genus="Pseudaetobatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FF7B9C174C3C0C0A" box="[189,375,803,829]" italics="true" pageId="141" pageNumber="142">Pseudaetobatus</emphasis>
|
||
(Tallahatta Formation)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="57BE87D4FF08FF22FD049C174E7C0C0A" authorityName="de Blainville" authorityYear="1816" box="[706,823,803,829]" class="Elasmobranchii" family="Myliobatidae" genus="Aetobatus" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FD049C174E7C0C0A" box="[706,823,803,829]" italics="true" pageId="141" pageNumber="142">Aetobatus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<collectingRegion id="527A32B5FF08FF22FC819C174ED10C0A" box="[839,922,803,829]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
Formation) is much weaker than that of
|
||
<taxonomicName id="57BE87D4FF08FF22FF7B9C734C7C0C56" authorityName="Cappetta" authorityYear="1986" box="[189,311,839,865]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FF7B9C734C7C0C56" box="[189,311,839,865]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
, and the root lobes of the former two taxa conspicuously extend lingually past the crown. Lastly, a conspicuous feature of
|
||
<taxonomicName id="57BE87D4FF08FF22FDF99C5F4FF20CB2" authorityName="Cappetta" authorityYear="1986" box="[575,697,875,901]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FDF99C5F4FF20CB2" box="[575,697,875,901]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
teeth is the thick enameloid covering on the crown, which is tuberculated on the occlusal surface. This contrasts with the smooth enameloid occurring on all other
|
||
<taxonomicName id="57BE87D4FF08FF22FF7B9C874C120CFA" authorityName="Bonaparte" authorityYear="1835" box="[189,345,947,973]" class="Elasmobranchii" family="Myliobatidae" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="subFamily" subFamily="Myliobatinae">Myliobatinae</taxonomicName>
|
||
and
|
||
<taxonomicName id="57BE87D4FF08FF22FE549C874F730CFA" authorityName="Jordan & Evermann" authorityYear="1896" box="[402,568,947,973]" class="Elasmobranchii" family="Rhinopteridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="subFamily" subFamily="Rhinopterinae">Rhinopterinae</taxonomicName>
|
||
teeth within Claibornian strata.
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF08FF20FF7B9CC14E890E7E" blockId="141.[189,1399,1013,1329]" lastBlockId="143.[189,1398,267,329]" lastPageId="143" lastPageNumber="144" pageId="141" pageNumber="142">
|
||
Within the species of
|
||
<taxonomicName id="57BE87D4FF08FF22FE039CC14F740B38" authorityName="Cappetta" authorityYear="1986" box="[453,575,1013,1039]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF08FF22FE039CC14F740B38" box="[453,575,1013,1039]" italics="true" pageId="141" pageNumber="142">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
, the crowns of
|
||
<taxonomicName id="57BE87D4FF08FF22FD3B9CC24EEA0B38" box="[765,929,1013,1039]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="species" species="granulosus">
|
||
<emphasis id="A2CA2045FF08FF22FD3B9CC24EEA0B38" box="[765,929,1013,1039]" italics="true" pageId="141" pageNumber="142">L. granulosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="57BE87D4FF08FF22FC749CC2497B0B38" box="[946,1072,1014,1039]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="species" species="rusticus">
|
||
<emphasis id="A2CA2045FF08FF22FC749CC2497B0B38" box="[946,1072,1014,1039]" italics="true" pageId="141" pageNumber="142">L. rusticus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="57BE87D4FF08FF22FBB39CC249B50B38" box="[1141,1278,1014,1039]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="species" species="zemensis">
|
||
<emphasis id="A2CA2045FF08FF22FBB39CC249B50B38" box="[1141,1278,1014,1039]" italics="true" pageId="141" pageNumber="142">L. zemensis</emphasis>
|
||
</taxonomicName>
|
||
appear to be more uniformly convex than
|
||
<taxonomicName id="57BE87D4FF08FF22FDFA9B2E4FFD0B04" authorityName="Leidy" authorityYear="1876" box="[572,694,1050,1075]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF08FF22FDFA9B2E4FFD0B04" box="[572,694,1050,1075]" italics="true" pageId="141" pageNumber="142">L. jugosus</emphasis>
|
||
</taxonomicName>
|
||
. MSC 35005 (
|
||
<figureCitation id="0885E0D2FF08FF22FCA09B2D4EF00B03" box="[870,955,1049,1076]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="141" pageNumber="142">Fig. 52</figureCitation>
|
||
A–E) is virtually identical to the
|
||
<typeStatus id="4F0542F5FF08FF22FA829B2E483C0B03" box="[1348,1399,1050,1076]" pageId="141" pageNumber="142">type</typeStatus>
|
||
specimen illustrated by
|
||
<bibRefCitation id="F42F81A6FF08FF22FE119B0A4F260B6F" author="Leidy J." box="[471,621,1086,1112]" pageId="141" pageNumber="142" pagination="209 - 261" refId="ref144864" refString="Leidy J. 1877. Description of vertebrate remains, chiefly from the Phosphate Beds of South Carolina. Journal of the Academy of Natural Sciences of Philadelphia 8 (2): 209 - 261." type="journal article" year="1877">Leidy (1877</bibRefCitation>
|
||
: pl. 31, figs 4–5) and
|
||
<bibRefCitation id="F42F81A6FF08FF22FCB69B0A48540B6F" author="Hovestadt D. C. & Hovestadt-Euler M." box="[880,1311,1086,1112]" pageId="141" pageNumber="142" pagination="1 - 66" refId="ref143545" refString="Hovestadt D. C. & Hovestadt-Euler M. 2013. Generic assessment and reallocation of Cenozoic Myliobatinae based on new information of tooth, tooth plate and caudal spine morphology of extant taxa. Palaeontos 24: 1 - 66." type="journal article" year="2013">Hovestadt & Hovestadt-Euler (2013</bibRefCitation>
|
||
: pl. 49, fig. 19), and it compares closely with a specimen from the middle Eocene of western Africa shown by
|
||
<bibRefCitation id="F42F81A6FF08FF22FF7B9BB24C320B97" author="Cappetta H." box="[189,377,1158,1184]" pageId="141" pageNumber="142" pagination="185 - 207" refId="ref137813" refString="Cappetta H. 1986. Myliobatidae nouveaux (Neoselachii, Batomorphii) de l'Ypresien des Ouled Abdoun, Maroc. Geologica et Palaeontologica 20: 185 - 207." type="journal article" year="1986">Cappetta (1986</bibRefCitation>
|
||
: pl. 2, fig. 4).
|
||
<bibRefCitation id="F42F81A6FF08FF22FDEB9BB24F800B97" author="Leidy J." box="[557,715,1158,1184]" pageId="141" pageNumber="142" pagination="209 - 261" refId="ref144864" refString="Leidy J. 1877. Description of vertebrate remains, chiefly from the Phosphate Beds of South Carolina. Journal of the Academy of Natural Sciences of Philadelphia 8 (2): 209 - 261." type="journal article" year="1877">Leidy (1877)</bibRefCitation>
|
||
made no mention of the tuberculated ornamentation we observed on our Claiborne specimens, and he attributed the “dull” surface and thinness of the lateral parts of the crown to
|
||
<emphasis id="A2CA2045FF08FF22FE759BFB4F4A0BDF" box="[435,513,1231,1256]" italics="true" pageId="141" pageNumber="142">in vivo</emphasis>
|
||
usage. The apparent lack of ornament on Leidy’s illustrated specimen could be the result of wear or poor representation by the illustrator, but our Claiborne specimens show that the thinness of the sides of the crown reflects the natural morphology of the tooth. Four specimens within our sample indicate a degree of heterodonty within the median row of
|
||
<taxonomicName id="57BE87D4FF0BFF21FC2E9E3F49290E12" authorityName="Cappetta" authorityYear="1986" box="[1000,1122,267,293]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="142" pageNumber="143" phylum="Chordata" rank="genus">
|
||
<emphasis id="A2CA2045FF0BFF21FC2E9E3F49290E12" box="[1000,1122,267,293]" italics="true" pageId="142" pageNumber="143">Leidybatis</emphasis>
|
||
</taxonomicName>
|
||
dentitions. MSC 37082 (
|
||
<figureCitation id="0885E0D2FF0BFF21FF009E1A4C500E7E" box="[198,283,302,329]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="142" pageNumber="143">Fig. 52</figureCitation>
|
||
T–X) has a broad but very low convexity that is skewed to one side of the tooth. MSC 37086 (
|
||
<figureCitation id="0885E0D2FF0BFF21FF009E664C510E5A" box="[198,282,338,365]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="142" pageNumber="143">Fig. 52</figureCitation>
|
||
K–O) has a more convex swelling that is also offset to one side of the crown, and the shoulders are more elongated than on MSC 37082. MSC 37083 (
|
||
<figureCitation id="0885E0D2FF0BFF21FC919E414EE50EA7" box="[855,942,373,400]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="142" pageNumber="143">Fig. 52</figureCitation>
|
||
NN–RR) is a smaller tooth with high, medially located swelling and short shoulders, whereas MSC 35005 (a much larger tooth,
|
||
<figureCitation id="0885E0D2FF0BFF21FB259EAC487D0E84" box="[1251,1334,408,435]" captionStart="Fig" captionStartId="141.[189,232,1399,1425]" captionTargetBox="[195,1392,543,2022]" captionTargetId="figure@142.[189,1398,537,2049]" captionTargetPageId="142" captionText="Fig. 52 (opposite page). Leidybatis jugosus (Leidy, 1876), teeth. A–E. MSC 35005, median tooth, Tallahatta/Lisbon formation contact zone. A. Profile view. B. Labial view. C. Lingual view. D. Oral view. E. Basal view. F–J. MSC 37080, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. F. Labial view. G. Profile view. H. Lingual view. I. Oral view. J. Basal view. K–O. MSC 37086, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. K. Profile view. L. Labial view. M. Lingual view. N. Oral view. O. Basal view. P–S. MSC 37291, lateral tooth, basal Lisbon Formation. P. Profile view. Q. Lingual view. R. Oral view. S. Basal view. T–X. MSC 37082, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. T. Profile view. U. Labial view. V. Lingual view. W. Oral view. X. Basal view. Y–CC. MSC 37081, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. Y. Lingual view. Z. Profile view. AA. Labial view. BB. Oral view. CC. Basal view. DD–HH. SC2012.47.198, median tooth, basal Lisbon Formation. DD. Profile view. EE. Labial view. FF. Lingual view. GG. Oral view. HH. Basal view. II– MM. MSC 37135, lateral tooth, Tallahatta/Lisbon formation contact zone, courtesy of James Lowery. II. Labial view. JJ. Profile view. KK. Lingual view. LL. Oral view. MM. Basal view. NN–RR. MSC 37083, median tooth, Tallahatta/Lisbon formation contact zone, courtesy of Bruce Relihan. NN. Profile view. OO. Labial view. PP. Lingual view. QQ. Oral view. RR. Basal view. SS–TT. SC2012.47.202, lateral tooth, basal Lisbon Formation. SS. Oral view. TT. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =1 cm; F–J, P–S, Y–CC, II–MM, SS–TT=5 mm." figureDoi="http://doi.org/10.5281/zenodo.3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="142" pageNumber="143">Fig. 52</figureCitation>
|
||
A–E) has a very high medial swelling and elongated shoulders. Unfortunately, we lack dentitions and cannot accurately distinguish upper from lower teeth, but we believe the
|
||
<collectingRegion id="527A32B5FF0BFF21FC7E9EEA49420ECF" box="[952,1033,478,504]" country="Portugal" name="Lisboa" pageId="142" pageNumber="143">Lisbon</collectingRegion>
|
||
sample represents intraspecific variation within a single species. There is currently no evidence to suggest that the two tooth fragments from the Tallahatta Formation are not conspecific with
|
||
<taxonomicName id="57BE87D4FF0AFF20FC859E1B4EF70E7F" baseAuthorityName="Leidy" baseAuthorityYear="1876" box="[835,956,303,328]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="143" pageNumber="144" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF0AFF20FC859E1B4EF70E7F" box="[835,956,303,328]" italics="true" pageId="143" pageNumber="144">L. jugosus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<caption id="C4C1ACDFFF08FF22FF7B9A43483C08DF" ID-DOI="http://doi.org/10.5281/zenodo.3660361" ID-Zenodo-Dep="3660361" httpUri="https://zenodo.org/record/3660361/files/figure.png" pageId="141" pageNumber="142" targetBox="[195,1392,543,2022]" targetPageId="142">
|
||
<paragraph id="9001FC57FF08FF22FF7B9A43483C08DF" blockId="141.[189,1399,1399,2025]" pageId="141" pageNumber="142">
|
||
<emphasis id="A2CA2045FF08FF22FF7B9A434C5D0AA6" bold="true" box="[189,278,1399,1425]" pageId="141" pageNumber="142">Fig. 52</emphasis>
|
||
(opposite page).
|
||
<taxonomicName id="57BE87D4FF08FF22FE369A434EC80AA5" authority="(Leidy, 1876)" authorityName="Leidy" authorityYear="1876" baseAuthorityName="Leidy" baseAuthorityYear="1876" box="[496,899,1399,1426]" class="Elasmobranchii" family="Myliobatidae" genus="Leidybatis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Myliobatiformes" pageId="141" pageNumber="142" phylum="Chordata" rank="species" species="jugosus">
|
||
<emphasis id="A2CA2045FF08FF22FE369A434F9B0AA6" box="[496,720,1399,1425]" italics="true" pageId="141" pageNumber="142">Leidybatis jugosus</emphasis>
|
||
(
|
||
<bibRefCitation id="F42F81A6FF08FF22FD229A434E320AA5" author="Leidy J." box="[740,889,1399,1426]" pageId="141" pageNumber="142" pagination="86 - 87" refId="ref144835" refString="Leidy J. 1876. Remarks on fossils of the Ashley Phosphate Beds. Proceedings of the Academy of Natural Sciences of Philadelphia 28: 86 - 87." type="journal article" year="1876">Leidy, 1876</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, teeth.
|
||
<emphasis id="A2CA2045FF08FF22FC199A43495D0AA6" bold="true" box="[991,1046,1399,1425]" pageId="141" pageNumber="142">A–E</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FBEC9A4349230AA6" box="[1066,1128,1399,1425]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
35005, median tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FEFC9AAE4CC10A83" box="[314,394,1434,1460]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone.
|
||
<emphasis id="A2CA2045FF08FF22FD619AAE4FF70A83" bold="true" box="[679,700,1434,1460]" pageId="141" pageNumber="142">A</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FCA79AAF4E3E0A82" bold="true" box="[865,885,1435,1461]" pageId="141" pageNumber="142">B</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FBD09AAE49600A83" bold="true" box="[1046,1067,1434,1460]" pageId="141" pageNumber="142">C</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FB1A9AAF49BA0A82" bold="true" box="[1244,1265,1435,1461]" pageId="141" pageNumber="142">D</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FF7B9A8A4D9A0AEF" bold="true" box="[189,209,1470,1496]" pageId="141" pageNumber="142">E</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FEB29A8A4CE80AEF" bold="true" box="[372,419,1470,1496]" pageId="141" pageNumber="142">F–J</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FE739A8A4CB80AEF" box="[437,499,1470,1496]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37080, lateral tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FCB59A8A4E880AEF" box="[883,963,1470,1496]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone, courtesy of Bruce Relihan.
|
||
<emphasis id="A2CA2045FF08FF22FEB49AD54CCF0ACC" bold="true" box="[370,388,1505,1531]" pageId="141" pageNumber="142">F</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FDE19AD54F750ACC" bold="true" box="[551,574,1505,1531]" pageId="141" pageNumber="142">G</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FD229AD54FB00ACC" bold="true" box="[740,763,1505,1531]" pageId="141" pageNumber="142">H</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FC699AD54EF10ACC" bold="true" box="[943,954,1505,1531]" pageId="141" pageNumber="142">I</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FB819AD5491D0ACC" bold="true" box="[1095,1110,1505,1531]" pageId="141" pageNumber="142">J</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FB299AD548600ACC" bold="true" box="[1263,1323,1505,1531]" pageId="141" pageNumber="142">K–O</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FAFE9AD5483D0ACC" box="[1336,1398,1505,1531]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37086, median tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FDF899304FC50929" box="[574,654,1540,1566]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone, courtesy of James Lowery.
|
||
<emphasis id="A2CA2045FF08FF22FAC7993048530929" bold="true" box="[1281,1304,1540,1566]" pageId="141" pageNumber="142">K</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FEC2991C4C530975" bold="true" box="[260,280,1576,1602]" pageId="141" pageNumber="142">L</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FE02991C4CAB0975" bold="true" box="[452,480,1576,1602]" pageId="141" pageNumber="142">M</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FD5A991C4FFA0975" bold="true" box="[668,689,1576,1602]" pageId="141" pageNumber="142">N</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FC8099134E160976" bold="true" box="[838,861,1575,1601]" pageId="141" pageNumber="142">O</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FC399913497B0976" bold="true" box="[1023,1072,1575,1601]" pageId="141" pageNumber="142">P–S</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FB86991349350976" box="[1088,1150,1575,1601]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37291, lateral tooth, basal
|
||
<collectingRegion id="527A32B5FF08FF22FEC2997E4C1E0953" box="[260,341,1610,1636]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
Formation.
|
||
<emphasis id="A2CA2045FF08FF22FE2A997F4CB50952" bold="true" box="[492,510,1611,1637]" pageId="141" pageNumber="142">P</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FD74997E4F820953" bold="true" box="[690,713,1610,1636]" pageId="141" pageNumber="142">Q</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FC4F997F4ED50952" bold="true" box="[905,926,1611,1637]" pageId="141" pageNumber="142">R</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FBFE997E49030953" bold="true" box="[1080,1096,1610,1636]" pageId="141" pageNumber="142">S</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FB29997F486D0952" bold="true" box="[1263,1318,1611,1637]" pageId="141" pageNumber="142">T–X</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FAFE997F483D0952" box="[1336,1398,1611,1637]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37082, median tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FDF9995A4FC409BF" box="[575,655,1646,1672]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone, courtesy of Bruce Relihan.
|
||
<emphasis id="A2CA2045FF08FF22FAC2995A485309BF" bold="true" box="[1284,1304,1646,1672]" pageId="141" pageNumber="142">T</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FEC599A54C53099C" bold="true" box="[259,280,1681,1707]" pageId="141" pageNumber="142">U</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FE7899A54C98099C" bold="true" box="[446,467,1681,1707]" pageId="141" pageNumber="142">V</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FD4C99A54FEC099C" bold="true" box="[650,679,1681,1707]" pageId="141" pageNumber="142">W</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FCF199A54E07099C" bold="true" box="[823,844,1681,1707]" pageId="141" pageNumber="142">X</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FC2F99A5497C099C" bold="true" box="[1001,1079,1681,1707]" pageId="141" pageNumber="142">Y–CC</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FB8399A549C8099C" box="[1093,1155,1681,1707]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37081, lateral tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FEFC99804CC109F9" box="[314,394,1716,1742]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone, courtesy of Bruce Relihan.
|
||
<emphasis id="A2CA2045FF08FF22FC349980494C09F9" bold="true" box="[1010,1031,1716,1742]" pageId="141" pageNumber="142">Y</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FB069980499F09F9" bold="true" box="[1216,1236,1716,1742]" pageId="141" pageNumber="142">Z</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FF7B99E34DAC09C6" bold="true" box="[189,231,1751,1777]" pageId="141" pageNumber="142">AA</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FE4D99EC4CF909C5" bold="true" box="[395,434,1752,1778]" pageId="141" pageNumber="142">BB</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FDF999E34F2209C6" bold="true" box="[575,617,1751,1777]" pageId="141" pageNumber="142">CC</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FCC599EC4E2109C5" bold="true" box="[771,874,1752,1778]" pageId="141" pageNumber="142">DD–HH</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FCBE99E34ED009C6" box="[888,923,1751,1777]" pageId="141" pageNumber="142">SC</collectionCode>
|
||
2012.47.198, median tooth, basal
|
||
<collectingRegion id="527A32B5FF08FF22FAE399E3483D09C6" box="[1317,1398,1751,1777]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
Formation.
|
||
<emphasis id="A2CA2045FF08FF22FE8099CF4C3B0822" bold="true" box="[326,368,1787,1813]" pageId="141" pageNumber="142">DD</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FDDA99CF4F080822" bold="true" box="[540,579,1787,1813]" pageId="141" pageNumber="142">EE</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FD2D99CF4E440822" bold="true" box="[747,783,1787,1813]" pageId="141" pageNumber="142">FF</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FC0199CE4EBE0823" bold="true" box="[967,1013,1786,1812]" pageId="141" pageNumber="142">GG</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FB4399CF49F80822" bold="true" box="[1157,1203,1787,1813]" pageId="141" pageNumber="142">HH</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FA9799CF4DBF080F" bold="true" pageId="141" pageNumber="142">II– MM</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FEC2982A4C09080F" box="[260,322,1822,1848]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37135, lateral tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FD78982A4E45080F" box="[702,782,1822,1848]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone, courtesy of James Lowery.
|
||
<emphasis id="A2CA2045FF08FF22FF7B98754D9F086C" bold="true" box="[189,212,1857,1883]" pageId="141" pageNumber="142">II</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FEB898754CD0086C" bold="true" box="[382,411,1857,1883]" pageId="141" pageNumber="142">JJ</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FD8C98754F33086C" bold="true" box="[586,632,1857,1883]" pageId="141" pageNumber="142">KK</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FCF498754E12086C" bold="true" box="[818,857,1857,1883]" pageId="141" pageNumber="142">LL</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FC2B9875496F086C" bold="true" box="[1005,1060,1857,1883]" pageId="141" pageNumber="142">MM</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FB0398754863086C" bold="true" box="[1221,1320,1857,1883]" pageId="141" pageNumber="142">NN–RR</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FAFE9875483D086C" box="[1336,1398,1857,1883]" pageId="141" pageNumber="142">MSC</collectionCode>
|
||
37083, median tooth, Tallahatta/
|
||
<collectingRegion id="527A32B5FF08FF22FDFF98504FC20849" box="[569,649,1892,1918]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
formation contact zone, courtesy of Bruce Relihan.
|
||
<emphasis id="A2CA2045FF08FF22FB29985048520849" bold="true" box="[1263,1305,1892,1918]" pageId="141" pageNumber="142">NN</emphasis>
|
||
. Profile view.
|
||
<emphasis id="A2CA2045FF08FF22FEC398B34C780896" bold="true" box="[261,307,1927,1953]" pageId="141" pageNumber="142">OO</emphasis>
|
||
. Labial view.
|
||
<emphasis id="A2CA2045FF08FF22FE1998BC4F480895" bold="true" box="[479,515,1928,1954]" pageId="141" pageNumber="142">PP</emphasis>
|
||
. Lingual view.
|
||
<emphasis id="A2CA2045FF08FF22FD0698B34FA50896" bold="true" box="[704,750,1927,1953]" pageId="141" pageNumber="142">QQ</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FC4298BC4EE50895" bold="true" box="[900,942,1928,1954]" pageId="141" pageNumber="142">RR</emphasis>
|
||
. Basal view.
|
||
<emphasis id="A2CA2045FF08FF22FB9798B349EC0896" bold="true" box="[1105,1191,1927,1953]" pageId="141" pageNumber="142">SS–TT</emphasis>
|
||
.
|
||
<collectionCode id="F6AF6492FF08FF22FB7F98B349970896" box="[1209,1244,1927,1953]" pageId="141" pageNumber="142">SC</collectionCode>
|
||
2012.47.202, lateral tooth, basal
|
||
<collectingRegion id="527A32B5FF08FF22FE65989E4CBF08F3" box="[419,500,1962,1988]" country="Portugal" name="Lisboa" pageId="141" pageNumber="142">Lisbon</collectingRegion>
|
||
Formation.
|
||
<emphasis id="A2CA2045FF08FF22FD4F989E4FE108F3" bold="true" box="[649,682,1962,1988]" pageId="141" pageNumber="142">SS</emphasis>
|
||
. Oral view.
|
||
<emphasis id="A2CA2045FF08FF22FC87989F4E2308F2" bold="true" box="[833,872,1963,1989]" pageId="141" pageNumber="142">TT</emphasis>
|
||
. Basal view. Labial at top in oral and basal views. Scale bars: A–E, K–O, T–X, DD–HH, NN–RR =
|
||
<quantity id="574651B2FF08FF22FC8F98FA4ECD08DF" box="[841,902,1998,2024]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="141" pageNumber="142" unit="cm" value="1.0">1 cm</quantity>
|
||
; F–J, P–S, Y–CC, II–MM, SS–TT=
|
||
<quantity id="574651B2FF08FF22FAEA98FA483908DF" box="[1324,1394,1998,2025]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="141" pageNumber="142" unit="mm" value="5.0">5 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9001FC57FF0AFF20FF7B9E444E5F0EBD" blockId="143.[189,788,368,394]" box="[189,788,368,394]" pageId="143" pageNumber="144">
|
||
<heading id="CB494B3BFF0AFF20FF7B9E444E5F0EBD" bold="true" box="[189,788,368,394]" fontSize="11" level="3" pageId="143" pageNumber="144" reason="3">
|
||
<emphasis id="A2CA2045FF0AFF20FF7B9E444E5F0EBD" bold="true" box="[189,788,368,394]" pageId="143" pageNumber="144">
|
||
Stratigraphic and geographic range in
|
||
<collectingRegion id="527A32B5FF0AFF20FD649E444E5F0EBD" box="[674,788,368,394]" country="United States of America" name="Alabama" pageId="143" pageNumber="144">Alabama</collectingRegion>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="9001FC57FF0AFF20FF7B9EAB483C0D37" blockId="143.[189,1399,414,512]" pageId="143" pageNumber="144">
|
||
Two tooth fragments were collected from the lower Tallahatta Formation at site ADl-1. The remaining sample is from the contact of the Tallahatta and
|
||
<collectingRegion id="527A32B5FF0AFF20FD3A9EF64E060EEB" box="[764,845,450,476]" country="Portugal" name="Lisboa" pageId="143" pageNumber="144">Lisbon</collectingRegion>
|
||
formations at sites ACh-14 and ACov-11, and the basal
|
||
<collectingRegion id="527A32B5FF0AFF20FEEC9ED14C300EC8" box="[298,379,485,511]" country="Portugal" name="Lisboa" pageId="143" pageNumber="144">Lisbon</collectingRegion>
|
||
Formation at site ACov-11. Upper Ypresian to middle Lutetian, zones NP14 and NP15.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |