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<mods:title id="0627020675C898DF01208DEC06B32D21">On the morphology of the astragalus and calcaneus of the amphicyonids (Carnivora, Mammalia) from the Paleogene of Europe: implications for the ecology of the European bear-dogs</mods:title>
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<mods:namePart id="4C9BB1E10469034852363193950AA5B1">Fournier, Morgane</mods:namePart>
<mods:affiliation id="53EBADB8B72D5FDE71046685B62F8A6D">CR 2 P (CNRS, MNHN, Sorbonne Université), Département Origines et Évolution, Muséum national dHistoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="A49DDB7BB5AA72F806BA615EDB3D69F8">Ladevèze, Sandrine</mods:namePart>
<mods:affiliation id="B86DFAFAFABC175446EE489F8C4649E7">CR 2 P (CNRS, MNHN, Sorbonne Université), Département Origines et Évolution, Muséum national dHistoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="BB1F9C356EB1D08D4EF6D7D17BF7709A">Verger, Kévin Le</mods:namePart>
<mods:affiliation id="48012F9EC63970E5F0CFBB83F6F92AE2">CR 2 P (CNRS, MNHN, Sorbonne Université), Département Origines et Évolution, Muséum national dHistoire naturelle, case postale 38, 57 rue Cuvier, F- 75231 Paris cedex 05 (France)</mods:affiliation>
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<mods:namePart id="03F42622F5C39C02240AC66E30C3A4E7">Fischer, Valentin</mods:namePart>
<mods:affiliation id="4DF7865CA7596A40F3EF37FE97F718BC">Evolution &amp; Diversity Dynamics Lab, Université de Liège (ULiège), Quartier Agora, Bâtiment B 18, Allée du six août, 14, 4000 Liège (Belgium)</mods:affiliation>
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<mods:namePart id="AE41A0FA617031848C488198401EE149">Speijer, Robert P.</mods:namePart>
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<mods:namePart id="1BE21C3CC85217D96CB665FCEA5D452A">Solé, Floréal</mods:namePart>
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<mods:title id="AFC07A2DDD4695DAF0F6409102908EE9">Geodiversitas</mods:title>
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<subSubSection id="D7A59BADFFA8FFCAFBABFF3EC2C5994B" pageId="13" pageNumber="316" type="nomenclature">
<paragraph id="9F00C826FFA8FFCAFBABFF3EC2C5994B" blockId="13.[1054,1214,217,276]" pageId="13" pageNumber="316">
<heading id="C4487F4AFFA8FFCAFBABFF3EC2DD98AB" box="[1054,1214,217,244]" pageId="13" pageNumber="316">
<taxonomicName id="58BFB3A5FFA8FFCAFBABFF3EC2DD98AB" box="[1054,1214,217,244]" class="Mammalia" family="Amphicyonidae" genus="Morphotype b" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="316" phylum="Chordata" rank="genus">Morphotype B</taxonomicName>
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(
<figureCitation id="0784D4A3FFA8FFCAFB8BFF1EC21F994C" box="[1086,1148,249,275]" captionStart="FIG" captionStartId="14.[132,143,1581,1598]" captionTargetBox="[271,1316,215,1539]" captionTargetId="figure@14.[270,842,782,1352]" captionTargetPageId="14" captionText="FIG. 4. — Calcanei of undetermined amphicyonids from the Quercy (France; Priabonian-Chattian, MP18-MP30): A-E, Morphotype A, MNHN.F.Qu9845, left calcaneus (reversed views): A, dorsal view; B, lateral view; C, medial view; D, distal view; E, proximal view; F-J, Morphotype B, KUL.PLV1542_28, left calcaneus (reversed views); F, dorsal view; G, lateral view; H, medial view; I, distal view; J, proximal view; K-O, Morphotype C, KUL.PLV1542_29, left calcaneus (reversed views): K, dorsal view; L, lateral view;M, medial view; N, distal view; O, proximal view. Abbreviations:cf, cuboid facet; ef, ectal facet; lpt, lateral process of tuber; mpt, medial process of tuber; pp, peroneal process; plt, plantar tubercle; ptg, tendinous plantar groove; sg, sagittal groove; sf, sustentacular facet; st, sustentaculum tali; tc, tuber calcanei. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.3952813" httpUri="https://zenodo.org/record/3952813/files/figure.png" pageId="13" pageNumber="316">Fig. 4</figureCitation>
F-J)
</paragraph>
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<subSubSection id="D7A59BADFFA8FFCAFC98FEDEC2B99DD0" pageId="13" pageNumber="316" type="description">
<paragraph id="9F00C826FFA8FFCAFC98FEDEC5CB990C" blockId="13.[813,1456,313,882]" box="[813,936,313,339]" pageId="13" pageNumber="316">
<emphasis id="ADCB1434FFA8FFCAFC98FEDEC5CB990C" box="[813,936,313,339]" italics="true" pageId="13" pageNumber="316">Dorsal view</emphasis>
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FEBEC2749B2D" blockId="13.[813,1456,313,882]" pageId="13" pageNumber="316">
The sagittal groove is rather visible. The
<emphasis id="ADCB1434FFA8FFCAFB7FFEBEC336992C" box="[1226,1365,345,371]" italics="true" pageId="13" pageNumber="316">tuber calcanei</emphasis>
is proximodistally quite elongated. The ectal facet is more inclined relative to the proximodistal axis than in other morphotypes, as well as in
<taxonomicName id="58BFB3A5FFA8FFCAFC03FE5FC21F998D" box="[950,1148,440,466]" class="Mammalia" family="Amphicyonidae" genus="Cynodictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="316" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="ADCB1434FFA8FFCAFC03FE5FC21F998D" box="[950,1148,440,466]" italics="true" pageId="13" pageNumber="316">Cynodictis lacustris</emphasis>
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. It is strongly convex in the middle and very slightly concave in its proximal part. The edges of this facet are rather well pronounced, except the distal edge, which merges with the body of the calcaneus and is therefore difficult to delimit. The sustentacular facet is rounded and extends slightly on the proximal edge of the
<emphasis id="ADCB1434FFA8FFCAFC98FD9FC58B9ACD" box="[813,1000,632,658]" italics="true" pageId="13" pageNumber="316">sustentaculum tali</emphasis>
. It is placed more proximal on the body of the calcaneus than in the other two calcaneal morphotypes. The peroneal process and its proximal and distal edges are quite well defined. The cuboid facet is inclined with respect to the mediolateral axis and its lateral edge rises over the body of the calcaneus. The plantar tubercle is distally very long and clearly visible in dorsal view. It is longer than that of
<taxonomicName id="58BFB3A5FFA8FFCAFCFFFCB0C26C9B2E" box="[842,1039,855,881]" class="Mammalia" family="Amphicyonidae" genus="Cynodictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="13" pageNumber="316" phylum="Chordata" rank="species" species="lacustris">
<emphasis id="ADCB1434FFA8FFCAFCFFFCB0C26C9B2E" box="[842,1039,855,881]" italics="true" pageId="13" pageNumber="316">Cynodictis lacustris</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FC70C5CE9BEF" blockId="13.[813,1455,919,1041]" box="[813,941,919,945]" pageId="13" pageNumber="316">
<emphasis id="ADCB1434FFA8FFCAFC98FC70C5CE9BEF" box="[813,941,919,945]" italics="true" pageId="13" pageNumber="316">Lateral view</emphasis>
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FC50C3C99C4E" blockId="13.[813,1455,919,1041]" pageId="13" pageNumber="316">
The dorsal and ventral edges of the
<emphasis id="ADCB1434FFA8FFCAFB08FC50C32C9B8E" box="[1213,1359,951,977]" italics="true" pageId="13" pageNumber="316">tuber calcanei</emphasis>
are very slightly concave. The cavity, proximal to the ectal facet, is deep and positioned higher on the tuber than in Morphotype A.
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FBD1C5CD9C0F" blockId="13.[813,1333,1078,1137]" box="[813,942,1078,1104]" pageId="13" pageNumber="316">
<emphasis id="ADCB1434FFA8FFCAFC98FBD1C5CD9C0F" box="[813,942,1078,1104]" italics="true" pageId="13" pageNumber="316">Medial view</emphasis>
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FBB1C3569C2F" blockId="13.[813,1333,1078,1137]" box="[813,1333,1110,1137]" pageId="13" pageNumber="316">The tendinous plantar groove is quite excavated.</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FB71C5C09CEF" blockId="13.[813,1455,1174,1296]" box="[813,931,1174,1200]" pageId="13" pageNumber="316">
<emphasis id="ADCB1434FFA8FFCAFC98FB71C5C09CEF" box="[813,931,1174,1200]" italics="true" pageId="13" pageNumber="316">Distal view</emphasis>
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FB51C2189D50" blockId="13.[813,1455,1174,1296]" pageId="13" pageNumber="316">The cuboid facet is smaller than in Morphotype A. The plantar tubercle is ventrally quite broad. The edge separating these two structures is not very clear.</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FAD2C5A19D10" blockId="13.[813,1455,1333,1423]" box="[813,962,1333,1359]" pageId="13" pageNumber="316">
<emphasis id="ADCB1434FFA8FFCAFC98FAD2C5A19D10" box="[813,962,1333,1359]" italics="true" pageId="13" pageNumber="316">Proximal view</emphasis>
</paragraph>
<paragraph id="9F00C826FFA8FFCAFC98FAB2C2B99DD0" blockId="13.[813,1455,1333,1423]" pageId="13" pageNumber="316">
The proximal part of the
<emphasis id="ADCB1434FFA8FFCAFB98FAB2C2D49D30" box="[1069,1207,1365,1391]" italics="true" pageId="13" pageNumber="316">tuber calcanei</emphasis>
has a rather oval surface that is slightly dorsoventrally elongated.
</paragraph>
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