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<mods:title id="2029FB6194EA26C712179B9783EE652D">Phylogeny and systematics of the Trapeziidae Miers, 1886 (Crustacea: Brachyura), with the description of a new family</mods:title>
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<treatment id="162E87D3836E1F4EFA7DFA5CFE2BFE26" ID-DOI="http://doi.org/10.5281/zenodo.5675084" ID-GBIF-Taxon="120829194" ID-Zenodo-Dep="5675084" LSID="urn:lsid:plazi:treatment:162E87D3836E1F4EFA7DFA5CFE2BFE26" httpUri="http://treatment.plazi.org/id/162E87D3836E1F4EFA7DFA5CFE2BFE26" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<subSubSection id="D69D654E836E1F49FA7DFA5CFD42FA1E" box="[264,719,1492,1518]" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="9E3836C5836E1F49FA7DFA5CFD42FA1E" blockId="51.[264,719,1492,1518]" box="[264,719,1492,1518]" pageId="51" pageNumber="52">
<heading id="C57081A9836E1F49FA7DFA5CFD42FA1E" bold="true" box="[264,719,1492,1518]" fontSize="11" level="5" pageId="51" pageNumber="52" reason="0">
<taxonomicName id="59874D46836E1F49FA7DFA5CFD42FA1E" ID-CoL="57Y2G" authority="Herbst, 1799" authorityName="Herbst" authorityYear="1799" box="[264,719,1492,1518]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD7836E1F49FA7DFA5CFD42FA1E" bold="true" box="[264,719,1492,1518]" pageId="51" pageNumber="52">
<emphasis id="ACF3EAD7836E1F49FA7DFA5CFD98FA1E" bold="true" box="[264,533,1492,1518]" italics="true" pageId="51" pageNumber="52">Trapezia rufopunctata</emphasis>
(Herbst, 1799)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D69D654E836E1F49FA7DF992FEFBF940" pageId="51" pageNumber="52" type="reference_group">
<paragraph id="9E3836C5836E1F49FA7DF992FC13F9C0" blockId="51.[264,1323,1561,1712]" box="[264,926,1561,1585]" pageId="51" pageNumber="52">
<taxonomicName id="59874D46836E1F49FA7DF992FD20F9C0" authority="Herbst, 1799: 54" authorityName="Herbst" authorityPageNumber="54" authorityYear="1799" box="[264,685,1561,1585]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufopunctatus">
<emphasis id="ACF3EAD7836E1F49FA7DF992FE7CF9C1" box="[264,497,1562,1585]" italics="true" pageId="51" pageNumber="52">Cancer rufopunctatus</emphasis>
Herbst, 1799: 54
</taxonomicName>
, pl. 47, fig. 6 (colour).
</paragraph>
<paragraph id="9E3836C5836E1F49FA7DF9B2FD4DF9A0" blockId="51.[264,1323,1561,1712]" box="[264,704,1593,1617]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="BE9711EB836E1F49FA7DF9B2FD4DF9A0" box="[264,704,1593,1617]" pageId="51" pageNumber="52">
<taxonomicName id="59874D46836E1F49FA7DF9B2FD31F9A0" ID-CoL="6L4TZ" authority="Macleay, 1838: 67" authorityName="Macleay" authorityPageNumber="67" authorityYear="1838" box="[264,700,1593,1617]" class="Malacostraca" family="Trapeziidae" genus="Grapsillus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="ACF3EAD7836E1F49FA7DF9B2FE60F9A1" box="[264,493,1594,1617]" italics="true" pageId="51" pageNumber="52">Grapsillus maculatus</emphasis>
<treatmentCitation id="1F2610D4836E1F49FA86F9B1FD31F9A0" author="Macleay" box="[499,700,1593,1616]" page="67" pageId="51" pageNumber="52" year="1838">
<bibRefCitation id="FA164B34836E1F49FA86F9B1FD19F9A0" author="Macleay" box="[499,660,1593,1616]" pageId="51" pageNumber="52" refString="Macleay, W. S. (1838) On the Brachyurous Decapod Crustacea. brought from the Cape by Dr. Smith. In: Smith, A., Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa: ' together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe. Invertebratae. Smith, Elder &amp; Co., London, pp. 53 - 71, pls. 2, 3." type="journal article" year="1838">Macleay, 1838</bibRefCitation>
: 67
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E3836C5836E1F49FA7DF9D2FCAFF980" blockId="51.[264,1323,1561,1712]" box="[264,802,1625,1649]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="BE9711EB836E1F49FA7DF9D2FCAFF980" box="[264,802,1625,1649]" pageId="51" pageNumber="52">
<taxonomicName id="59874D46836E1F49FA7DF9D2FC93F980" ID-CoL="57XZB" authority="A. Milne Edwards, 1867: 281" authorityName="A. Milne Edwards" authorityPageNumber="281" authorityYear="1867" box="[264,798,1625,1649]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="acutifrons">
<emphasis id="ACF3EAD7836E1F49FA7DF9D2FE5AF981" box="[264,471,1626,1649]" italics="true" pageId="51" pageNumber="52">Trapezia acutifrons</emphasis>
A.
<treatmentCitation id="1F2610D4836E1F49FA8AF9D1FC93F980" author="Milne" box="[511,798,1625,1648]" page="281" pageId="51" pageNumber="52" year="1867">
<bibRefCitation id="FA164B34836E1F49FA8AF9D1FD64F980" author="Milne" box="[511,745,1625,1648]" pageId="51" pageNumber="52" refString="Milne Edwards, A. (1867) Descriptions de quelques especes nouvelles des Crustaces Brachyoures. Annales de la Societe Entomologique de France, ser. 4, 7 (4), 263 - 288." type="journal article" year="1867">Milne Edwards, 1867</bibRefCitation>
: 281
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9E3836C5836E1F49FA7DF9F2FEFBF940" blockId="51.[264,1323,1561,1712]" pageId="51" pageNumber="52">
<treatmentCitationGroup id="BE9711EB836E1F49FA7DF9F2FEFBF940" pageId="51" pageNumber="52">
<taxonomicName id="59874D46836E1F49FA7DF9F2FD36F960" ID-CoL="4QZ56" authority="Chen, 1933: 111" authorityName="Chen" authorityPageNumber="111" authorityYear="1933" box="[264,699,1657,1681]" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD7836E1F49FA7DF9F2FD88F961" box="[264,517,1658,1681]" italics="true" pageId="51" pageNumber="52">Quadrella rufopunctata</emphasis>
<treatmentCitation id="1F2610D4836E1F49F97EF9F1FD36F960" author="Chen" box="[523,699,1657,1680]" page="111" pageId="51" pageNumber="52" year="1933">
<bibRefCitation id="FA164B34836E1F49F97EF9F1FD04F960" author="Chen" box="[523,649,1657,1680]" pageId="51" pageNumber="52" refString="Chen, P. S. (1933) Zur Morphologie und Histologie der Respirations-organe von Grapsus grapsus L., nebst einer Liste Krabben der Sammlung Plate von Ceylon und Sudindien. Jenaische Zeitschrift fur Naturwissenschaft, 68, 31 - 116." type="journal article" year="1933">Chen, 1933</bibRefCitation>
: 111
</treatmentCitation>
</taxonomicName>
, fig. 55 (junior homonym of
<taxonomicName id="59874D46836E1F49F89BF9F2FEE3F940" authority="Herbst, 1799" authorityName="Herbst" authorityYear="1799" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufopunctatus">
<emphasis id="ACF3EAD7836E1F49F89BF9F2FB5BF961" box="[1006,1238,1658,1681]" italics="true" pageId="51" pageNumber="52">Cancer rufopunctatus</emphasis>
Herbst, 1799
</taxonomicName>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="D69D654E836E1F4EFA7DF954FE2BFE26" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="discussion">
<paragraph id="9E3836C5836E1F49FA7DF954FB92F89E" blockId="51.[264,1323,1756,1902]" pageId="51" pageNumber="52">
<emphasis id="ACF3EAD7836E1F49FA7DF954FEE3F905" box="[264,366,1756,1781]" italics="true" pageId="51" pageNumber="52">Remarks</emphasis>
. The taxonomy of this species is now quite stable.
<bibRefCitation id="FA164B34836E1F49F8BFF954FB44F906" author="Ng" box="[970,1225,1756,1782]" pageId="51" pageNumber="52" refString="Ng, P. K. L. &amp; Ahyong, S. T. (2000) Brachyuran type specimens (Crustacea: Decapoda) in the Macleay collection, University of Sydney, Australia. Raffles Bulletin of Zoology (Singapore), 49 (1), 83 - 100." type="journal article" year="2000">Ng &amp; Ahyong (2000)</bibRefCitation>
showed that the
<typeStatus id="413C8867836E1F49FA1DF88CFE59F8EE" box="[360,468,1796,1822]" pageId="51" pageNumber="52" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="59874D46836E1F49FA88F88CFC22F8EE" authority="Macleay, 1838" authorityName="Macleay" authorityYear="1838" box="[509,943,1796,1822]" class="Malacostraca" family="Trapeziidae" genus="Grapsillus" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="ACF3EAD7836E1F49FA88F88CFD77F8ED" box="[509,762,1796,1821]" italics="true" pageId="51" pageNumber="52">Grapsillus maculatus</emphasis>
<bibRefCitation id="FA164B34836E1F49F877F88CFC22F8EE" author="Macleay" box="[770,943,1796,1822]" pageId="51" pageNumber="52" refString="Macleay, W. S. (1838) On the Brachyurous Decapod Crustacea. brought from the Cape by Dr. Smith. In: Smith, A., Illustrations of the Zoology of South Africa; consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836; fitted out by ' The Cape of Good Hope Association for Exploring Central Africa: ' together with a summary of African Zoology, and an inquiry into the geographical ranges of species in that quarter of the globe. Invertebratae. Smith, Elder &amp; Co., London, pp. 53 - 71, pls. 2, 3." type="journal article" year="1838">Macleay, 1838</bibRefCitation>
</taxonomicName>
was still extant (in the Macleay Museum, University of Sydney) and that it fits the concept of
<taxonomicName id="59874D46836E1F49F891F8A4FB14F8B5" box="[996,1177,1836,1861]" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD7836E1F49F891F8A4FB14F8B5" box="[996,1177,1836,1861]" italics="true" pageId="51" pageNumber="52">T. rufopunctata</emphasis>
</taxonomicName>
as currently understood (see
<bibRefCitation id="FA164B34836E1F49FABDF8DCFD7DF89E" author="Galil" box="[456,752,1876,1902]" pageId="51" pageNumber="52" refString="Galil, B. &amp; Lewinsohn, C. (1985) On the taxonomic status of Trapezia rufopunctata (Herbst) and Trapezia flavopunctata Eydoux &amp; Souleyet (Decapoda, Brachyura). Crustaceana, 48 (2), 209 - 217." type="journal article" year="1985">Galil &amp; Lewinsohn 1985</bibRefCitation>
: 209;
<bibRefCitation id="FA164B34836E1F49F842F8DCFC5AF89E" author="Castro" box="[823,983,1876,1902]" pageId="51" pageNumber="52" refString="Castro, P. (1997 a) Trapeziid crabs (Brachyura: Xanthoidea: Trapeziidae) of New Caledonia, eastern Australia, and the Coral Sea. In: Richer de Forges, B. (Ed.), Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, Benthos). Etudes et Theses, 3, 59 - 107." type="journal article" year="1997" yearSuffix="a">Castro 1997a</bibRefCitation>
: 127).
</paragraph>
<paragraph id="9E3836C583691F4EFA4DFE94FE2BFE26" blockId="52.[264,1324,284,470]" pageId="52" pageNumber="53">
The dried male and female
<typeStatus id="413C886783691F4EF9F6FE94FD67FEC6" box="[643,746,284,310]" pageId="52" pageNumber="53" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="59874D4683691F4EF861FE94FAA1FEC6" authority="A. Milne Edwards, 1867" authorityName="A. Milne Edwards" authorityYear="1867" box="[788,1324,284,310]" class="Malacostraca" family="Trapeziidae" genus="Trapezia" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="acutifrons">
<emphasis id="ACF3EAD783691F4EF861FE94FC76FEC5" box="[788,1019,284,309]" italics="true" pageId="52" pageNumber="53">Trapezia acutifrons</emphasis>
A.
<bibRefCitation id="FA164B3483691F4EFF5FFE94FAA1FEC6" author="Milne" box="[1066,1324,284,310]" pageId="52" pageNumber="53" refString="Milne Edwards, A. (1867) Descriptions de quelques especes nouvelles des Crustaces Brachyoures. Annales de la Societe Entomologique de France, ser. 4, 7 (4), 263 - 288." type="journal article" year="1867">Milne Edwards, 1867</bibRefCitation>
</taxonomicName>
(MNHN­B 2913) from the Hawaiian Islands are clearly identical to
<taxonomicName id="59874D4683691F4EFF50FECCFB54FEAD" box="[1061,1241,324,349]" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783691F4EFF50FECCFB54FEAD" box="[1061,1241,324,349]" italics="true" pageId="52" pageNumber="53">T. rufopunctata</emphasis>
</taxonomicName>
. From its description, but not the figure (
<bibRefCitation id="FA164B3483691F4EF87EFEE4FBA9FE76" author="Chen" box="[779,1060,364,390]" pageId="52" pageNumber="53" refString="Chen, P. S. (1933) Zur Morphologie und Histologie der Respirations-organe von Grapsus grapsus L., nebst einer Liste Krabben der Sammlung Plate von Ceylon und Sudindien. Jenaische Zeitschrift fur Naturwissenschaft, 68, 31 - 116." type="journal article" year="1933">Chen 1933: 111, fig. 55</bibRefCitation>
),
<taxonomicName id="59874D4683691F4EFF4DFEE4FEB8FE5D" class="Malacostraca" family="Trapeziidae" genus="Quadrella" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783691F4EFF4DFEE4FEB8FE5D" italics="true" pageId="52" pageNumber="53">Quadrella rufopunctata</emphasis>
</taxonomicName>
(
<typeStatus id="413C886783691F4EFA30FE1CFEF4FE5E" box="[325,377,404,430]" pageId="52" pageNumber="53">type</typeStatus>
locality
<collectingCountry id="E690765583691F4EFAABFE1CFDDFFE5E" box="[478,594,404,430]" name="Sri Lanka" pageId="52" pageNumber="53">Sri Lanka</collectingCountry>
) (not
<taxonomicName id="59874D4683691F4EF9E3FE1CFBA7FE5E" authority="Herbst, 1799" authorityName="Herbst" authorityYear="1799" box="[662,1066,404,430]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783691F4EF9E3FE1CFC06FE5D" box="[662,907,404,429]" italics="true" pageId="52" pageNumber="53">Cancer rufopunctata</emphasis>
Herbst, 1799
</taxonomicName>
) is almost certainly
<taxonomicName id="59874D4683691F4EFE60FE1CFE12FE25" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="rufopunctata">
<emphasis id="ACF3EAD783691F4EFE60FE1CFE12FE25" italics="true" pageId="52" pageNumber="53">T. rufopunctata</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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