treatments-xml/data/13/F4/81/13F48102D2695F0F9E55656C158A365C.xml

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<mods:title id="A57950BEEDBCC774090638A7C71B1635">A review of Copelatus diving beetles from the Solomon Islands, reporting the discovery of six new species (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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<mods:namePart id="D6C050E2BBD3B9AEBBF39E196D95738C">Hajek, Jiri</mods:namePart>
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<mods:affiliation id="BEEC9D5FE3B11F6C98ED897049E7FCA7">Department of Entomology, National Museum, Cirkusova 1740, CZ- 193 00, Prague 9 - Horni Pocernice, Czech Republic</mods:affiliation>
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<mods:namePart id="CE9D47610AB84AAAB4DA46856A284D69">Shaverdo, Helena</mods:namePart>
<mods:affiliation id="FE0E2EC856933B52384F4170BBD53EBA">Naturhistorisches Museum Wien, Burgring 7, 1010, Vienna, Austria</mods:affiliation>
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<mods:namePart id="F7F0FCC458DAC29757E93F494DEB7E59">Hendrich, Lars</mods:namePart>
<mods:affiliation id="B4172DCCD921AA3D7F7BAF6605EF8510">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany</mods:affiliation>
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<mods:namePart id="86F657E71F886A5BFEABAAFC9CF48B34">Balke, Michael</mods:namePart>
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<mods:affiliation id="2F170ADE660DF98BEC686E4EEEDA7B1D">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247, Munich, Germany &amp; GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
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<mods:date id="307B2728A39CAAF96448B69A2CCEAF30">2021</mods:date>
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<treatment id="13F48102D2695F0F9E55656C158A365C" ID-GBIF-Taxon="180331294" LSID="urn:lsid:zoobank.org:act:507C2AEA-9E85-4957-A8BC-F3442EC83D7A" httpUri="http://treatment.plazi.org/id/13F48102D2695F0F9E55656C158A365C" lastPageNumber="81" pageId="0" pageNumber="81" scope_family="Dytiscidae" scope_order="Coleoptera" scope_subFamily="Copelatinae">
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<paragraph id="F71700FDAEF8FB87DC64C0B538E3D729" pageId="0" pageNumber="81">
<taxonomicName id="97C865E27A91D8A3A1965FFF89D394F4" ID-CoL="8M23S" LSID="http://zoobank.org/507C2AEA-9E85-4957-A8BC-F3442EC83D7A" authority="Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke, 2021" authorityName="Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus laevipennis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="laevipennis" status="sp. nov.">Copelatus laevipennis</taxonomicName>
<taxonomicNameLabel id="E18F1D27624243C07F22BD98582AEB36" pageId="0" pageNumber="81">sp. nov.</taxonomicNameLabel>
<figureCitation id="6350A4C91610B08EB197D65EF2B8F527" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Habitus of Copelatus 3 C. kietensis sp. nov. (holotype; TL: 6.3 mm) 4 C. laevipennis sp. nov. (holotype; TL: 6.9 mm)." figureDoi="10.3897/zookeys.1023.61478.figures3-4" httpUri="https://binary.pensoft.net/fig/519049" pageId="0" pageNumber="81">Figures 4</figureCitation>
<figureCitation id="09F1FAB6DCF0B4D366CFAFAC1CB2DBEA" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">, 18</figureCitation>
<figureCitation id="21FDEF930A13B5374B9EC4F100A7858D" captionStart="Figures 23–25" captionStartId="F15" captionText="Figures 23 - 25. Variability of Copelatus male genitalia: median lobe in lateral view, left side 23 Copelatus laevipennis sp. nov. (paratype) 24, 25 C. variistriatus sp. nov. (paratypes). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figures23-25" httpUri="https://binary.pensoft.net/fig/519062" pageId="0" pageNumber="81">, 23</figureCitation>
</paragraph>
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<typeStatus id="473A6A071320EF36BE067B55D91D36A5">Type</typeStatus>
locality.
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<paragraph id="4CADB895CAFDEA2A31746CC21EFFDDFC" pageId="0" pageNumber="81">
<materialsCitation id="B5261A39BF776425A76D8795A93C0384" ID-GBIF-Occurrence="3063630346" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Solomon Islands" latitude="-9.628166" location="Mbaole Village" longLatPrecision="13" longitude="160.1115" specimenCount="1" typeStatus="holotype">
<collectingCountry id="C99615066DE3E31E5E028C3F84990380" name="Solomon Islands">Solomon Islands</collectingCountry>
, 
<taxonomicName id="DFBE34BC55A620D621A73832A19F46B4" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
, 
<quantity id="798F000D2AB081B254DF9C61552DD938" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="km" value="0.5">0.5 km</quantity>
N of 
<location id="A242A72866D1AE45519DB00F16C98AF9" LSID="urn:lsid:plazi:treatment:13F48102D2695F0F9E55656C158A365C:A242A72866D1AE45519DB00F16C98AF9" country="Solomon Islands" latitude="-9.628166" longLatPrecision="13" longitude="160.1115" name="Mbaole Village">Mbaole Village</location>
, 
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, 
<geoCoordinate id="6EE91ED94C5B9DF54DBC2585B5ED9E5A" degrees="160" direction="east" minutes="06.69" orientation="longitude" precision="9" value="160.1115">160°06.69'E</geoCoordinate>
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.
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<paragraph id="06E24C479B5D375D9F655E83967D39D6" pageId="0" pageNumber="81">
<typeStatus id="DEB5E6E541107E27094964B790FE45DC">Type</typeStatus>
material.
</paragraph>
<paragraph id="FB96397CBB1EE21EAC568A53E076D403" pageId="0" pageNumber="81">
<emphasis id="763C08E82ED267E4B13E1CB1819EAC3F" bold="true" pageId="0" pageNumber="81">
<emphasis id="99E4D345B3C695893B5C536BF629982D" italics="true" pageId="0" pageNumber="81">
<typeStatus id="5B7D292DCE02951A1EE816A79628EF7B">Holotype</typeStatus>
</emphasis>
:
</emphasis>
<materialsCitation id="607EC4213AFA17CD2F1785F353AAA25B" ID-GBIF-Occurrence="3063630324" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="ZSMG" collectorName="K. Mailautoka &amp; M. Balke" country="Solomon Islands" determinerName="Balke" elevation="853" location="Mbaole" longitude="9.628166" specimenCount="2" typeStatus="holotype">
<specimenCount id="AFC6AAE67290025B9266F9FD55B8A3E2">♂</specimenCount>
, labelled: &quot; 
<collectingCountry id="6381D882FAB950E789E549BC4BB2EE6D" name="Solomon Islands">Salomonen</collectingCountry>
: C- 
<taxonomicName id="6C4136D5DF069F109311C2E2AAEA5776" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
, / 
<quantity id="ACFCCA844A59D5538D55F79C61675697" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="km" value="0.5">0.5 km</quantity>
N 
<location id="D58439957059719A5EE20E5DAA4381AA" LSID="urn:lsid:plazi:treatment:13F48102D2695F0F9E55656C158A365C:D58439957059719A5EE20E5DAA4381AA" country="Solomon Islands" longitude="9.628166" name="Mbaole">Mbaole</location>
, 
<quantity id="80CAFEBB0F6DCC4C55EC52781787887F" metricMagnitude="2" metricUnit="m" metricValue="8.531352" unit="ft" value="2799.0">
<elevation id="687ABB0658124C7B259BA51558E01454" metricMagnitude="2" metricUnit="m" metricValue="8.531352" unit="ft" value="2799.0">2799 feet</elevation>
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/ S 
<geoCoordinate id="2859BCAA0343AF1EC7BAE71AA8E00005" degrees="09" direction="east" minutes="37.69" orientation="longitude" precision="9" value="9.628166">09°37.69 E</geoCoordinate>
<geoCoordinate id="6F12A6B9AFBE8BA39101B4AFCC5A89AE" degrees="160" direction="east" minutes="06.69" orientation="longitude" precision="9" value="160.1115">160°06.69E</geoCoordinate>
/ 2007 
<collectorName id="0D56C73EA47DAB409F869A5054E2BBC2">K. Mailautoka</collectorName>
leg. [p] // DNA / 
<collectorName id="020E265EDF62D7BD4A3AF0B3E6A945A2">M. Balke</collectorName>
/ 2907 [green label, p] // 
<typeStatus id="B589B6BBD01BE628661DED4368AB5C5E">HOLOTYPE</typeStatus>
<specimenCount id="6582AD73EB521B84FB19BAF235027B7D">♂</specimenCount>
/ 
<taxonomicName id="6F4A666A50938EF0CB9C310E7514AEE4" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="COPELATUS" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">
<emphasis id="3DC9E38FC80C768E81AA6477BC48253D" italics="true" pageId="0" pageNumber="81">COPELATUS</emphasis>
</taxonomicName>
/ 
<taxonomicName id="323DCC90A8E64CC2E84F3D60EAD7CBF0" authorityName="Hájek &amp; Shaverdo &amp; Hendrich &amp; Balke" authorityYear="2021" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus laevipennis" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="species" species="laevipennis">
<emphasis id="7A9A998AA5024FAF31953CC9CF9186B2" italics="true" pageId="0" pageNumber="81">Copelatus laevipennis</emphasis>
</taxonomicName>
sp. nov. / 
<normalizedToken id="3981A09B37E3345FAE3C252384F024A3" originalValue="Hájek">Hajek</normalizedToken>
, Shaverdo, Hendrich &amp; 
<determinerName id="F6300B721833D2E282D9A91DDB687BF5">Balke</determinerName>
det. 2020 [red label, p]&quot; (
<collectionCode id="BEA76F86B497A93F0081D737F153D51D">ZSMG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="2960C102D93129F3FB4A57DA716EC07C" pageId="0" pageNumber="81">
<materialsCitation id="321B3F50E07418032B436085CD6B798B" ID-GBIF-Occurrence="3063630301" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NMPC" collectorName="M. Balke" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="D9C11AFC664A2DE2123FABC6ED59C216" bold="true" pageId="0" pageNumber="81">
<emphasis id="55578F88B9D8E2DA0E32602102D74AF0" italics="true" pageId="0" pageNumber="81">
<typeStatus id="F789521A06F444FB2B1F157DD1B66D4F">Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<specimenCount id="D7C2323F26203B840DABE5C6910C8E98" type="male">1 ♂</specimenCount>
, same locality data as 
<typeStatus id="B23A46A1AE44721561AE084FDAEB3C9A">holotype</typeStatus>
, with additional label: &quot;DNA / 
<collectorName id="7AEE1439E397C55AAFF849EAA77326A1">M. Balke</collectorName>
/ 3334 [green label, p]&quot; (
<collectionCode id="35091535214AAA2095EF50293C3E9DF9" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="D567834F3359DA2714C9DADB5A7E147C" ID-GBIF-Occurrence="3063630314" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NMPC, ZSMG" collectorName="K. Mailautoka" country="Solomon Islands" elevation="853" latitude="-9.628166" location="Mbaole" longLatPrecision="13" longitude="160.11467" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="2A4EB311D0C9F19DA4057575EF7750E4" type="female">2 ♀♀</specimenCount>
, labelled: &quot; 
<collectingCountry id="9AF179E167B3B3FE392C28CB7D1EB76E" name="Solomon Islands">Solomon Islands</collectingCountry>
: C- 
<taxonomicName id="F2F0031AAA96925C25FB463F727A887D" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
, / 
<quantity id="779B21A670ACDF0F5FDD4CFC5146BE30" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="km" value="0.5">0.5 km</quantity>
N 
<location id="241E8166136A6E3B4C27012B5BFD5674" LSID="urn:lsid:plazi:treatment:13F48102D2695F0F9E55656C158A365C:241E8166136A6E3B4C27012B5BFD5674" country="Solomon Islands" latitude="-9.628166" longLatPrecision="13" longitude="160.11467" name="Mbaole">Mbaole</location>
, 
<quantity id="21F468ABA63678D4501E95E1CB89207F" metricMagnitude="2" metricUnit="m" metricValue="8.53" unit="m" value="853.0">
<elevation id="C644754689DD2C96A047CD980AC9C335" metricMagnitude="2" metricUnit="m" metricValue="8.53" unit="m" value="853.0">853 m</elevation>
</quantity>
, 2007, / 
<geoCoordinate id="EB553112BA7AC4FACF46594FE8564D93" degrees="09" direction="south" minutes="37.69" orientation="latitude" precision="9" value="-9.628166">09°37.69S</geoCoordinate>
, 
<geoCoordinate id="D4619E9EE45C3DCFD9202337C6839468" degrees="160" direction="east" minutes="06.88" orientation="longitude" precision="9" value="160.11467">160°06.88E</geoCoordinate>
, 
<collectorName id="F76A096035CE687EE69ACED7AF7094E2">K. Mailautoka</collectorName>
[p]&quot; (
<collectionCode id="83496A12642B3216E3773E9F28E234B2" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
, 
<collectionCode id="B3596B779EB55E7B721A0527D715F9E7">ZSMG</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="13A702B51C2941B97057133AF92F7B40" ID-GBIF-Occurrence="3063630318" collectingDate="1956-06-27" collectionCode="NHMUK, NMPC, ZSMG" collectorName="E. S. Brown" country="Solomon Islands" location="Suta" municipality="Is." specimenCount="17" specimenCount-female="7" specimenCount-male="10" typeStatus="paratype">
<specimenCount id="96DEE444D8A745E890EBB5567B58EC73" type="male">10 ♂♂</specimenCount>
, 
<specimenCount id="02AC7729FE987C4B6118C529DA7B284B" type="female">7 ♀♀</specimenCount>
, labelled: &quot;SOLOMON IS.: / 
<taxonomicName id="13C62AC0DD35BFC87DA47E6E4B6F5F38" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
<collectingMunicipality id="6CD2A54FF18EEF8C6634EF61144D727A">Is.</collectingMunicipality>
/ 
<location id="72C4BF06EE0FEFEEEA691E614B9554A3" LSID="urn:lsid:plazi:treatment:13F48102D2695F0F9E55656C158A365C:72C4BF06EE0FEFEEEA691E614B9554A3" country="Solomon Islands" municipality="Is." name="Suta">Suta</location>
/ 
<date id="3A314B09DB35EEE6A9BCCD3CD6670867" value="1956-06-27">
<collectingDate id="C2A887B150D133C21432A52E8CBD15BD" value="1956-06-27">27.vi.1956</collectingDate>
</date>
[p] // 
<collectorName id="373FB2EB56384DF07A2CE7BB37A7DF6F">E.S.Brown</collectorName>
/ B.M.1957-201 [p]&quot; (
<collectionCode id="FB972C4B5DD0F44FABD915E7FDB5DB02" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
, 
<collectionCode id="958569A77CBD66DBC7BF3ECDF60E648A" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" type="">NMPC</collectionCode>
, 
<collectionCode id="A020D3FFD236F55A173E9DC015EDE537">ZSMG</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="B9933FD8A0E902C401DA350F473E7EEA" ID-GBIF-Occurrence="3063630317" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMUK" collectorName="E. S. Brown" country="Solomon Islands" location="Suta" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="157D2B0EACCC09115AFBAB6B8E1EE354" type="male">1 ♂</specimenCount>
, labelled: &quot;5327 [on side, hw] / SOLOMON IS. [red underlined] / 
<taxonomicName id="12B77CC3E1EE77D5DA5A24200C9C7A8C" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
[p] / 
<location id="2DE5169C4D9651DE95498AA75FFED44F" LSID="urn:lsid:plazi:treatment:13F48102D2695F0F9E55656C158A365C:2DE5169C4D9651DE95498AA75FFED44F" country="Solomon Islands" name="Suta">Suta</location>
/ 27.vi. [hw] 195 [p] 6 [hw] / 
<collectorName id="FF84DDA37634FBF4688A1CDFCF199E52">E.S.Brown</collectorName>
[p] / Sangava / R [on reverse, hw] // E.S.BROWN coll / C.I.E. 1957-24 [hw] // 
<collectorName id="C6005170FA30C2C19C4A9F66E70BDDA0">E.S.Brown</collectorName>
/ B.M.1957-201 [p]&quot; (
<collectionCode id="002005EC373E751BC68A73F8CF931FE0" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="F663C50BDECB36ED7CF6A69EE4237478" ID-GBIF-Occurrence="3063630327" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectionCode="NHMUK" country="Solomon Islands" determinerName="J. Balfour-Browne" location="GUADALCANAL" specimenCount="2" specimenCount-male="1" stateProvince="Guadalcanal" typeStatus="paratype">
<specimenCount id="A3B7F16C0386CA44661A8B5FE32C3BDF" type="male">1 ♂</specimenCount>
, labelled: &quot; 
<specimenCount id="2BAD3AC403730517DF8A45050F4FB0BB">♂</specimenCount>
[p] // Type [round label with red frame, p] // SOLOMON IS: / 
<collectingRegion id="67072D4B3681A70B6F9FD6310333783B" country="Solomon Islands" name="Guadalcanal">GUADALCANAL</collectingRegion>
/ Tenaru R. headwaters. / 1820' 5.viii.53. / In hole in tree trunk. [hw] // Brit.Mus. / 1987-14 [p] // aC 2 [hw] // 
<taxonomicName id="0B4791F35491E6D5F643D35E9AD02EC9" class="Insecta" family="Dytiscidae" genus="Copelatus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Copelatus" order="Coleoptera" pageId="0" pageNumber="81" phylum="Arthropoda" rank="genus">Copelatus</taxonomicName>
/ torosus Type! [hw] / 
<determinerName id="36A92DDC405F6A1FD5FCF69E4428AEF0">J. Balfour-Browne</determinerName>
det., 195 [p] 3 [hw]&quot; (
<collectionCode id="F1233529F5E68E2299479432B32FA243" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="3E691C0D7AFA097EDFA2095C3D5266E0" ID-GBIF-Occurrence="3063630341" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" country="Solomon Islands" location="All" specimenCount="1" stateProvince="Guadalcanal" typeStatus="paratype">
<location id="2D71BE5833E7683940537DE3DD10DA22" LSID="urn:lsid:plazi:treatment:13F48102D2695F0F9E55656C158A365C:2D71BE5833E7683940537DE3DD10DA22" country="Solomon Islands" name="All" stateProvince="Guadalcanal">All</location>
<typeStatus id="2B46567687182D5AB26C1B969996527E">paratypes</typeStatus>
with the respective printed red label
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="776138C0AD49C4D2CA5966563759D0DC" pageId="0" pageNumber="81" type="description">
<paragraph id="14E92228DE6122147CB0885721DB4B65" pageId="0" pageNumber="81">
Description of male 
<typeStatus id="8F3581D7BA38B088FED7554AA8F69C79">holotype</typeStatus>
.
</paragraph>
<paragraph id="28114BE60A1D7D2C55B813B006238A69" pageId="0" pageNumber="81">
<emphasis id="A5EA04EE4FC943582991661CF5C86B2E" bold="true" pageId="0" pageNumber="81">
<emphasis id="FE5C04BB562CFFD7E439B56D365F4E22" italics="true" pageId="0" pageNumber="81">Habitus</emphasis>
:
</emphasis>
Elongate, oblong-oval, broadest shortly before mid-length of elytra; body distinctly convex in lateral view. Body outline continuous, without discontinuity between pronotum and elytra. Dorsal surface shiny (Fig. 
<figureCitation id="62FDFF6A8DD9B3EBDFCA9D56F7FC21CF" captionStart="Figures 3, 4" captionStartId="F2" captionText="Figures 3, 4. Habitus of Copelatus 3 C. kietensis sp. nov. (holotype; TL: 6.3 mm) 4 C. laevipennis sp. nov. (holotype; TL: 6.9 mm)." figureDoi="10.3897/zookeys.1023.61478.figures3-4" httpUri="https://binary.pensoft.net/fig/519049" pageId="0" pageNumber="81">4</figureCitation>
).
</paragraph>
<paragraph id="C497BDB0EF049FEE79D9CF43E794DAB7" pageId="0" pageNumber="81">
<emphasis id="0E4099DB002F2E8DABFE8BE6BCB1C0EE" bold="true" pageId="0" pageNumber="81">
<emphasis id="51459A099926FC991BDB1964AC0D995B" italics="true" pageId="0" pageNumber="81">Colouration</emphasis>
:
</emphasis>
Body colour pitchy brown; sides of pronotum (except for lateral margin) and appendages paler, ferruginous.
</paragraph>
<paragraph id="094F5F32041428EB8F2A1C4C7AF6B5DD" pageId="0" pageNumber="81">
<emphasis id="5C79C7C6D4A8314BF0B794C0C9B064DB" bold="true" pageId="0" pageNumber="81">
<emphasis id="3F4A73F10B99A26A1B4844800D140DDE" italics="true" pageId="0" pageNumber="81">Head</emphasis>
:
</emphasis>
Moderately broad, ca. 0.65 
<normalizedToken id="F665CBFAA778AE1BAFE5941F54E7581E" originalValue="×">x</normalizedToken>
width of pronotum, trapezoidal. Anterior margin of clypeus indistinctly concave. Antenna with antennomeres long and slender. Reticulation consisting of moderately deeply impressed isodiametric meshes. Punctation double, consisting of coarse setigerous punctures, and very small punctures spread sparsely on surface; row of coarse punctures present around inner margin of eyes, few punctures present at frontal level of eyes, and several punctures anterolaterally to eyes in fronto-clypeal depressions.
</paragraph>
<paragraph id="B0BD50C2E2FECA4C6FC76DB79A36864C" pageId="0" pageNumber="81">
<emphasis id="D6D9EECCEE4C6DBCD4482DC018D7CE72" bold="true" pageId="0" pageNumber="81">
<emphasis id="A077B0050EE73AA119B63242ABCA7449" italics="true" pageId="0" pageNumber="81">Pronotum</emphasis>
:
</emphasis>
Transverse (width/length ratio = 2.78), broadest between posterior angles, lateral margins moderately curved. Sides with lateral beading very thin and indistinct. Reticulation similar to that of head. Punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin, laterally in longitudinal depression close to sides, several punctures present also in basolateral depressions along basal margin. Disc of pronotum with shallow medial longitudinal scratch.
</paragraph>
<paragraph id="B53BC7ED1C9B5250EE3E2A5320C24ECD" pageId="0" pageNumber="81">
<emphasis id="E8DDA04A0779D03D72BD208C830721C5" bold="true" pageId="0" pageNumber="81">
<emphasis id="1B5273FBA6D1A29041DFDE387F1EFF4A" italics="true" pageId="0" pageNumber="81">Elytra</emphasis>
:
</emphasis>
Base of elytra as broad as pronotal base; lateral margins of elytra subparallel-sided in basal half, distinctly narrowing in apical half. Elytral striae absent. Reticulation similar to that of head and pronotum, meshes somewhat elongated longitudinally. Punctation consisting of coarse setigerous punctures and very fine sparse punctures. Coarse punctures arranged in three distinct longitudinal puncture lines: two discal and one lateral; another row of punctures present along lateral margin of elytra, and few coarse punctures present also in interspace between discal and lateral puncture lines.
</paragraph>
<paragraph id="6B62D8BB5F5A6391379993021F5EE976" pageId="0" pageNumber="81">
<emphasis id="E5BB7FCB48BF23F5903B3FD78BF44C04" bold="true" pageId="0" pageNumber="81">
<emphasis id="66ABACDD81163F69064ACED9A2E162CF" italics="true" pageId="0" pageNumber="81">Legs</emphasis>
:
</emphasis>
Protibia modified, angled near base, distinctly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1-3 distinctly broadened, with adhesive setae on their ventral side.
</paragraph>
<paragraph id="38E2C1C6EB0765F66EC1559A6EE68777" pageId="0" pageNumber="81">
<emphasis id="180CC4B5A99BFE1DF397BA00BE6FAD93" bold="true" pageId="0" pageNumber="81">
<emphasis id="A21DF60B4B07E4BC747ED57F5DEE0A3E" italics="true" pageId="0" pageNumber="81">Ventral side</emphasis>
:
</emphasis>
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation almost effaced. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (&quot;metasternal wings&quot;) tongue-shaped, slender. Metacoxal lines nearly complete, absent only very close to metaventrite. Metacoxal plates covered laterally with long, deep longitudinal strioles; reticulation consisting of extremely elongated, longitudinal polygonal meshes. Metacoxal processes rounded and incised at posterior margin. Abdominal ventrites I and II with longitudinal strioles; ventrites III and IV with oblique strioles laterally. Tuft of setae present antero-medially on ventrites III-V; ventrite VI with setigerous punctures laterally on either side. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I and II, oblique on ventrite III and transverse on ventrites IV-VI. Punctation consisting of fine, sparsely distributed punctures.
</paragraph>
<paragraph id="EE856219B9D9591BC6533D91C34200F2" pageId="0" pageNumber="81">
<emphasis id="F6CFDA3CBA6C94BA81AB0B9D4A03F4DD" bold="true" pageId="0" pageNumber="81">
<emphasis id="B841E842F08527BFD16C59AD7AEA8B48" italics="true" pageId="0" pageNumber="81">Genitalia</emphasis>
:
</emphasis>
Median lobe of aedeagus (Fig. 
<figureCitation id="7A891E01276CB40F3A76ACA03A9F40A5" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">18</figureCitation>
) sickle-shaped, with evident dorsal and ventral sclerites; dorsal sclerite without surface sculpture and divided into two parts in apical half: left part shorter than right one, both slightly curved, with small crests, notches and truncate apexes (Fig. 
<figureCitation id="739B9A84DD040A83A5B12FCF26A2EF64" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">18A, B</figureCitation>
); ventral sclerite divided into two parts apically: left part more strongly sclerotised, broader, shorter, with broadly pointed apex, right part longer, partly sclerotised (medially membranous), with elongate, thin apex in shape of very weak hook (Fig. 
<figureCitation id="1533BD43DE9AFEBD0E137932D9270CA9" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">18C</figureCitation>
).
</paragraph>
<paragraph id="ACAFC7441C747F83146D8895470CEC6C" pageId="0" pageNumber="81">
Lateral lobes (parameres) of narrow triangular form, with broader subdistal part due to curved setigerous dorsal margin; setae numerous, dense, and strong distally, and distinctly less numerous, weaker, and sparser basally (Fig. 
<figureCitation id="A2369918B766A2B96314D75FED4253C7" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">18D</figureCitation>
).
</paragraph>
<paragraph id="905CD41BB1DF5D523BB999FB0D58FD4D" pageId="0" pageNumber="81">
<emphasis id="A1AE64619C4390CCC1FCC6EBCB0278C9" bold="true" pageId="0" pageNumber="81">Female.</emphasis>
Identical to male in habitus. Protibia simple, not angled basally and only slightly broadened distally; pro- and mesotarsomeres not broadened, without adhesive setae.
</paragraph>
</subSubSection>
<subSubSection id="1DE1E0350A8BF1249597130A96C1FCC4" pageId="0" pageNumber="81" type="variability">
<paragraph id="D49939229F6707431722D597F4045594" pageId="0" pageNumber="81">Variability.</paragraph>
<paragraph id="1FE9F1DAFF763D5142D4B1FA537C1AFA" pageId="0" pageNumber="81">
All specimens of the type series agree well with the 
<typeStatus id="FB4096D7C16DCB0736B9A568244EDC7C">holotype</typeStatus>
. There is only slight variability in body colouration: some specimens have head with pale frontal part and dark vertex, and the sides of elytra are indistinctly paler than elytral disc. Few longitudinal strioles present in depression close to posterior angles of pronotum in one female. Small differences were detected also in the shape of the male median lobe: a notch on right part of dorsal sclerite can be almost absent in some specimens (cf. Figs 
<figureCitation id="8D146B850D3B737F64CAF8666C64BA6E" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">18B</figureCitation>
, 
<figureCitation id="D62CE43564F115230AA4E9CC3A03B1B4" captionStart="Figures 23–25" captionStartId="F15" captionText="Figures 23 - 25. Variability of Copelatus male genitalia: median lobe in lateral view, left side 23 Copelatus laevipennis sp. nov. (paratype) 24, 25 C. variistriatus sp. nov. (paratypes). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figures23-25" httpUri="https://binary.pensoft.net/fig/519062" pageId="0" pageNumber="81">23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="BB936F9D2D63074A7E906EC584A28A58" pageId="0" pageNumber="81" type="description">
<paragraph id="78878A46F95F1300C8105C54A7C6C419" pageId="0" pageNumber="81">Measurements.</paragraph>
<paragraph id="0224E6D1A3759F0FA70F14B91961E17D" pageId="0" pageNumber="81">
TL: 
<quantity id="95FF444BC229C958952F26AB9913B24F" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.2" metricValueMin="6.3" unit="mm" value="6.75" valueMax="7.2" valueMin="6.3">6.3-7.2 mm</quantity>
(mean value: 
<quantity id="CBBD143BF1E05F568409DFA4A5FF9E52" metricMagnitude="-3" metricUnit="m" metricValue="6.9" metricValueMax="7.1" metricValueMin="6.7" unit="mm" value="6.9" valueMax="7.1" valueMin="6.7">
6.9 
<normalizedToken id="9017399B6DC88235E75235F206B5842E" originalValue="±">+/-</normalizedToken>
0.2 mm
</quantity>
); 
<typeStatus id="E09305E522D344E02E28E3415A6B286B">holotype</typeStatus>
: 
<quantity id="CA72354A5AE6DF1A541808E9F6E1F7FC" metricMagnitude="-3" metricUnit="m" metricValue="6.9" unit="mm" value="6.9">6.9 mm</quantity>
. TL-h: 
<quantity id="0693F4265144F0E845E075B0A6119FE3" metricMagnitude="-3" metricUnit="m" metricValue="6.05" metricValueMax="6.4" metricValueMin="5.7" unit="mm" value="6.05" valueMax="6.4" valueMin="5.7">5.7-6.4 mm</quantity>
(mean value: 
<quantity id="D3BFA4DD713E15FE93E4BF1922783CCC" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.4" metricValueMin="6.0" unit="mm" value="6.2" valueMax="6.4" valueMin="6.0">
6.2 
<normalizedToken id="40C9EEA9D87361AC7CE59679BE5B9090" originalValue="±">+/-</normalizedToken>
0.2 mm
</quantity>
); 
<typeStatus id="8AEEBEFCCBBC24495EAA7A05DE8A4624">holotype</typeStatus>
: 
<quantity id="3C0C9172FD6C4C6263D84DDE4E4BB0C6" metricMagnitude="-3" metricUnit="m" metricValue="6.3" unit="mm" value="6.3">6.3 mm</quantity>
. MW: 
<quantity id="EBE8AACC80FCF56177F21AA530F9CDA8" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" unit="mm" value="3.4" valueMax="3.6" valueMin="3.2">3.2-3.6 mm</quantity>
(mean value: 
<quantity id="E664F362EE12DC38396DE8367394B641" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.5" metricValueMin="3.3" unit="mm" value="3.4" valueMax="3.5" valueMin="3.3">
3.4 
<normalizedToken id="159A5C6BDC4ABA46944C1D11FB859036" originalValue="±">+/-</normalizedToken>
0.1 mm
</quantity>
); 
<typeStatus id="B6D624A1462392B581A842F49450D013">holotype</typeStatus>
: 
<quantity id="7A7B0B9AD7D5F0963165F6785B5FD281" metricMagnitude="-3" metricUnit="m" metricValue="3.4" unit="mm" value="3.4">3.4 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="FCA2BC10C419A5E33E1B2DB610423703" pageId="0" pageNumber="81" type="differential diagnosis">
<paragraph id="56AC2A02A04DD5DA28CBC8F28A0FF2B1" pageId="0" pageNumber="81">Differential diagnosis.</paragraph>
<paragraph id="C71242F53578B32C8BB2765FDB750EF4" pageId="0" pageNumber="81">
Based on the absence of elytral striae, the new species can be included into the 
<taxonomicName id="3104DA4A98FF89C1B861586514C2408E" lsidName="C. hydroporoides" pageId="0" pageNumber="81" rank="species" species="hydroporoides">
<emphasis id="FD37E564550764324363CBDBE770890D" italics="true" pageId="0" pageNumber="81">C. hydroporoides</emphasis>
</taxonomicName>
species group (or 
<emphasis id="A4F32126F12B69F6F031190C089098E2" italics="true" pageId="0" pageNumber="81">haemorrhoidalis</emphasis>
, in earlier studies). This species group contains currently 55 species distributed predominantly in Afrotropical and Neotropical regions (
<bibRefCitation id="88B75A2B606DF22F59CE01E24400C103" author="Nilsson, AN" journalOrPublisher="Australian Entomologist" pageId="0" pageNumber="81" publicationUrl="http://www.waterbeetles.eu" refId="B16" refString="Nilsson, AN, Hajek, J, 2020. A World Catalogue of the Family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1.I.2020. Distributed as a PDF file via Internet. http://www.waterbeetles.eu" title="A World Catalogue of the Family Dytiscidae, or the diving beetles (Coleoptera, Adephaga). Version 1. I. 2020. Distributed as a PDF file via Internet." url="http://www.waterbeetles.eu" year="2020">
Nilsson and 
<normalizedToken id="D713877FA8E30FDE14D155ACD0EDE3E6" originalValue="Hájek">Hajek</normalizedToken>
2020
</bibRefCitation>
). Only two species of this group were recently described from the Oriental (
<collectingRegion id="DEFE6AFD3E90D89223A33EBD9BCAFAEF" country="Indonesia" name="Maluku Utara">Maluku</collectingRegion>
, 
<collectingCountry id="0856297A141F59723E90BC56C5DC1F50" name="Indonesia">Indonesia</collectingCountry>
) and Australian regions (
<collectingCountry id="C82DED8035145926816EFBCD93775D6D" name="Papua New Guinea">Papua New Guinea</collectingCountry>
) by 
<bibRefCitation id="C5A270FD71D15711719C65A1E9E46D29" author="Hajek, J" journalOrPublisher="Acta Entomologica Musei Nationalis Pragae" pageId="0" pageNumber="81" pagination="437 - 443" refId="B12" refString="Hajek, J, Hendrich, L, Hawlitschek, O, Balke, M, 2010. Copelatus sibelaemontis sp. nov. (Coleoptera: Dytiscidae) from the Moluccas with generic assignment based on morphology and DNA sequence data. Acta Entomologica Musei Nationalis Pragae 50: 437 - 443" title="Copelatus sibelaemontis sp. nov. (Coleoptera: Dytiscidae) from the Moluccas with generic assignment based on morphology and DNA sequence data." volume="50" year="2010">
<normalizedToken id="9FFBAAD8E0B5B286A357066988FAC6CD" originalValue="Hájek">Hajek</normalizedToken>
et al. (2010)
</bibRefCitation>
and 
<bibRefCitation id="35F8771838B5B44CE9B312E460D8C66B" author="Megna, YS" journalOrPublisher="Australian Entomologist" pageId="0" pageNumber="81" pagination="49 - 55" refId="B15" refString="Megna, YS, Atthakor, W, Manaono, M, Hendrich, L, Balke, M, 2017. A new species of Copelatus Erichson, 1832 (Coleoptera: Dytiscidae: Copelatinae) from the Adelbert Range of Papua New Guinea. Australian Entomologist 44: 49 - 55" title="A new species of Copelatus Erichson, 1832 (Coleoptera: Dytiscidae: Copelatinae) from the Adelbert Range of Papua New Guinea." volume="44" year="2017">Megna et al. (2017)</bibRefCitation>
, respectively.
</paragraph>
<paragraph id="E3FC5B8F1E42A098A28C624367684BE7" pageId="0" pageNumber="81">
Based on shape of the median lobe and paramere, 
<taxonomicName id="F376F8F744EDCA2ACD4DF1292CE34DF9" lsidName="C. laevipennis" pageId="0" pageNumber="81" rank="species" species="laevipennis">
<emphasis id="ECAE6BC59F3B0FD065BEC351DA0F85A0" italics="true" pageId="0" pageNumber="81">C. laevipennis</emphasis>
</taxonomicName>
sp. nov. is without any doubt closely related to 
<taxonomicName id="5EB6AF3753FEFC632DA1A891802236B6" lsidName="C. variistriatus" pageId="0" pageNumber="81" rank="species" species="variistriatus">
<emphasis id="58678D012792DB8D7EA1B69269FF36E7" italics="true" pageId="0" pageNumber="81">C. variistriatus</emphasis>
</taxonomicName>
sp. nov., which it resembles also in size, body shape, and colouration, and 
<taxonomicName id="E200BC631E9DCA3C9734395040129A4D" lsidName="C. urceolus" pageId="0" pageNumber="81" rank="species" species="urceolus">
<emphasis id="79D40FA6272F74B4131012F54E3D3E5B" italics="true" pageId="0" pageNumber="81">C. urceolus</emphasis>
</taxonomicName>
sp. nov. The 
<typeStatus id="4DC7E65BB548348CD0DA8BAA80072420">type</typeStatus>
localities of 
<taxonomicName id="2DBEDFDA5F716CC24D0639E9DD234CDF" lsidName="C. laevipennis" pageId="0" pageNumber="81" rank="species" species="laevipennis">
<emphasis id="324F6C697EA29411BBDD66DCF1F61138" italics="true" pageId="0" pageNumber="81">C. laevipennis</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName id="365946F532B1AAE6AFD0EBCEA86930B3" lsidName="C. variistriatus" pageId="0" pageNumber="81" rank="species" species="variistriatus">
<emphasis id="7892E2EEE19494050B24EE6A73766030" italics="true" pageId="0" pageNumber="81">C. variistriatus</emphasis>
</taxonomicName>
sp. nov. are only ca. 
<quantity id="6E1F61F7FF70B3A6423EED9F6E8AB13D" metricMagnitude="4" metricUnit="m" metricValue="2.0" unit="km" value="20.0">20 km</quantity>
apart, although they differ significantly in altitude. Due to great variability of elytral striation in 
<taxonomicName id="A36DD158A5A3BFD92E573A4E993AE839" lsidName="C. variistriatus" pageId="0" pageNumber="81" rank="species" species="variistriatus">
<emphasis id="1B9FE6DA40058DA34C61872A1F8DAC22" italics="true" pageId="0" pageNumber="81">C. variistriatus</emphasis>
</taxonomicName>
sp. nov. described below, we hesitated first to describe another species, which may represent only a non-striated population of one species. Finally, we have decided to describe both species, because of: 1) 
<taxonomicName id="BFF942392F227667C6B7767C8BD30CFD" lsidName="C. laevipennis" pageId="0" pageNumber="81" rank="species" species="laevipennis">
<emphasis id="E6BCD9DAF8910AC9B57EC0BB45F2BFD7" italics="true" pageId="0" pageNumber="81">C. laevipennis</emphasis>
</taxonomicName>
sp. nov. has more parallel habitus than 
<taxonomicName id="6B93DEAB911FF94D98BF501C78F66BA6" lsidName="C. variistriatus" pageId="0" pageNumber="81" rank="species" species="variistriatus">
<emphasis id="A65B92AC53B1CD3B88E78A79E6A08EB9" italics="true" pageId="0" pageNumber="81">C. variistriatus</emphasis>
</taxonomicName>
sp. nov.; 2) there are no intermediate specimens, i.e., there are no specimens of 
<taxonomicName id="8C96F8E6A320D898093C8B8D2692C4EC" lsidName="C. variistriatus" pageId="0" pageNumber="81" rank="species" species="variistriatus">
<emphasis id="8E17312AC7FB2ABAA89CFE28B83EF88E" italics="true" pageId="0" pageNumber="81">C. variistriatus</emphasis>
</taxonomicName>
sp. nov. without elytral striae, and no specimens of 
<taxonomicName id="835BE89385C7C3FCF64DF4AED15E9395" lsidName="C. laevipennis" pageId="0" pageNumber="81" rank="species" species="laevipennis">
<emphasis id="7F9CBF5487697E221A40C79B077CF234" italics="true" pageId="0" pageNumber="81">C. laevipennis</emphasis>
</taxonomicName>
sp. nov. with traces of striae; 3) minor differences in the shape of the median lobe can be detected between both species, mainly in shape of the apexes of parts of the dorsal and ventral sclerites (cf. Figs 
<figureCitation id="B2A01F5BD5962D81FC31A962EF799358" captionStart="Figure 18" captionStartId="F10" captionText="Figure 18. Male genitalia of Copelatus laevipennis sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure18" httpUri="https://binary.pensoft.net/fig/519057" pageId="0" pageNumber="81">18</figureCitation>
, 
<figureCitation id="1E63F016BF507A2E7CB40ED04F83576E" captionStart="Figure 22" captionStartId="F14" captionText="Figure 22. Male genitalia of Copelatus variistriatus sp. nov. (holotype) A median lobe in lateral view, right side B median lobe in lateral view, left side C median lobe in ventral view D left paramere in external view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figure22" httpUri="https://binary.pensoft.net/fig/519061" pageId="0" pageNumber="81">22</figureCitation>
- 
<figureCitation id="E9A5F6A0FFC974CF06E35E6AFAAD41BE" captionStart="Figures 23–25" captionStartId="F15" captionText="Figures 23 - 25. Variability of Copelatus male genitalia: median lobe in lateral view, left side 23 Copelatus laevipennis sp. nov. (paratype) 24, 25 C. variistriatus sp. nov. (paratypes). Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1023.61478.figures23-25" httpUri="https://binary.pensoft.net/fig/519062" pageId="0" pageNumber="81">25</figureCitation>
); 4) there is significant genetic divergence between both species: uncorrected genetic distances between the sequenced CO1 haplotypes of 
<taxonomicName id="3DCFC6A8A701C5336B959E726B218932" lsidName="C. laevipennis" pageId="0" pageNumber="81" rank="species" species="laevipennis">
<emphasis id="516B5F6ABB09D1721E1B4EAF21176020" italics="true" pageId="0" pageNumber="81">C. laevipennis</emphasis>
</taxonomicName>
sp. nov. and 
<taxonomicName id="00C4C5EA0953A7FADB6C18CFBF926E3D" lsidName="C. variistriatus" pageId="0" pageNumber="81" rank="species" species="variistriatus">
<emphasis id="6694D055401FCAA02DF68230848FBECA" italics="true" pageId="0" pageNumber="81">C. variistriatus</emphasis>
</taxonomicName>
sp. nov. are ca. 7% (pers. obs.).
</paragraph>
</subSubSection>
<subSubSection id="F0539AC43541998499EE4ED4709C5BC8" pageId="0" pageNumber="81" type="etymology">
<paragraph id="6EF59E1CD49AD4195CA7DD37BF8086F4" pageId="0" pageNumber="81">Etymology.</paragraph>
<paragraph id="72D0A9867A5820227EE97DEB2C375DFA" pageId="0" pageNumber="81">
The species name is composed from Latin adjective 
<emphasis id="6D0B6D506B1F98F32CADA9D26FAB8CDC" italics="true" pageId="0" pageNumber="81">laevis</emphasis>
(- 
<emphasis id="CB3F68FCE3BBBA6E5C5942ECA311961E" italics="true" pageId="0" pageNumber="81">is</emphasis>
, - 
<emphasis id="BBA7629325923D715733FBC9678F638A" italics="true" pageId="0" pageNumber="81">e</emphasis>
, = smooth) and noun 
<emphasis id="E63D2FF51BAFF20C215236B385F4B144" italics="true" pageId="0" pageNumber="81">penna</emphasis>
(- 
<emphasis id="67EC9C8535EF73C86F554195BA305B12" italics="true" pageId="0" pageNumber="81">ae</emphasis>
, feminine, = wing), referring to smooth elytra without striae.
</paragraph>
</subSubSection>
<subSubSection id="115874AC861C05A39083C6B67065D91D" pageId="0" pageNumber="81" type="distribution">
<paragraph id="E6E3A80420E23C93341165B353BDE6A3" pageId="0" pageNumber="81">Distribution.</paragraph>
<paragraph id="23E0023742F560B338B3AB71B15C6250" pageId="0" pageNumber="81">
The species is known only from medium altitude area (ca. 
<quantity id="8C70BBC8355ADF3CB212EAD784F7E722" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="9.0" metricValueMin="6.0" unit="m" value="750.0" valueMax="900.0" valueMin="600.0">600-900 m</quantity>
) in north-central 
<taxonomicName id="8F3431D6C0E0966CC5C10AE0DC18F540" genus="Guadalcanal" lsidName="Guadalcanal" pageId="0" pageNumber="81" rank="genus">Guadalcanal</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="2B61C0917F6292E378639FD98DC23111" pageId="0" pageNumber="81" type="habitat">
<paragraph id="CF5C8A2126B7BF38416272386C724D96" pageId="0" pageNumber="81">Habitat.</paragraph>
<paragraph id="CD81EBAE9547CDC3FB9C0F82E22DF245" pageId="0" pageNumber="81">Largely unknown; according to label data, one specimen was collected in a water-filled hole in tree trunk.</paragraph>
</subSubSection>
</treatment>
</document>