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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.827.31365" ID-GBIF-Dataset="85f74389-c078-44de-b0bd-8b7e4c3b9404" ID-PMC="PMC6472301" ID-Pensoft-Pub="1313-2970-827-31" ID-PubMed="31114425" ID-ZBK="10B663895E424E5287D8F49E2405D651" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-827-31" ModsDocOrigin="ZooKeys 827" ModsDocTitle="Mollusc species from the Pontocaspian region an expert opinion list" checkinTime="1551875342560" checkinUser="pensoft" docAuthor="Wesselingh, Frank, Poorten, Jan Johan ter, Kijashko, Pavel, Albrecht, Christian, Anistratenko, Olga Yu, Frolov, Pavel, Gándara, Alberto Martinez, Gittenberger, Arjan, Gogaladze, Aleksandre, Mikhail Karpinsky, Popa, Luis, Sands, Arthur F, Vandendorpe, Justine &amp; Wilke, Thomas" docDate="2019" docId="137F49F77908B1976FF0013D2FC9A739" docLanguage="en" docName="ZooKeys 827: 31-124" docOrigin="ZooKeys 827" docSource="http://dx.doi.org/10.3897/zookeys.827.31365" docTitle="Clathrocaspia knipowitschii" docType="treatment" docVersion="5" lastPageNumber="68" masterDocId="FFA2FA62FFEC4705FFBBFB79FFAAFFFE" masterDocTitle="Mollusc species from the Pontocaspian region - an expert opinion list" masterLastPageNumber="124" masterPageNumber="31" pageNumber="67" updateTime="1668166983143" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Mollusc species from the Pontocaspian region - an expert opinion list</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wesselingh, Frank</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poorten, Jan Johan ter</mods:namePart>
<mods:affiliation>Field Museum of Natural History, Chicago, United States of America</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Kijashko, Pavel</mods:namePart>
<mods:affiliation>Moscow State University, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Albrecht, Christian</mods:namePart>
<mods:affiliation>Justus Liebig University, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Anistratenko, Olga Yu</mods:namePart>
<mods:affiliation>Schmalhausen Instite of Zoology, National Academy of Sciences of Ukraine, Kiev, Ukraine &amp; Institute of Geological Sciences, National Academy of Sciences of Ukraine, KievUkraine</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Frolov, Pavel</mods:namePart>
<mods:affiliation>Saint-Petersburg State University, Saint Petersburg, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gándara, Alberto Martinez</mods:namePart>
<mods:affiliation>Grigore Artipa National Museum of Natural History, Bucharest, Romania</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gittenberger, Arjan</mods:namePart>
<mods:affiliation>Gittenberger Marine Research, Inventory &amp; Strategy, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gogaladze, Aleksandre</mods:namePart>
<mods:affiliation>Naturalis Biodiversity Center, Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mikhail Karpinsky</mods:namePart>
<mods:affiliation>Russian Federal Research Institute of Fisheries and Oceanography, Moscow, Russia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Popa, Luis</mods:namePart>
<mods:affiliation>Grigore Antirpa National Museum of Natural History, Bucharest, Romania</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sands, Arthur F</mods:namePart>
<mods:affiliation>Justus Liebig University, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vandendorpe, Justine</mods:namePart>
<mods:affiliation>Justus Liebig University, Giessen, Germany</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wilke, Thomas</mods:namePart>
<mods:affiliation>Justus Liebig University Giessen Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>827</mods:number>
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<mods:start>31</mods:start>
<mods:end>124</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.827.31365</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.827.31365</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-827-31</mods:identifier>
<mods:identifier type="ZBK">10B663895E424E5287D8F49E2405D651</mods:identifier>
<mods:identifier type="ZooBank">10B663895E424E5287D8F49E2405D651</mods:identifier>
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<treatment ID-GBIF-Taxon="183410485" LSID="urn:lsid:plazi:treatment:137F49F77908B1976FF0013D2FC9A739" httpUri="http://treatment.plazi.org/id/137F49F77908B1976FF0013D2FC9A739" lastPageNumber="68" pageNumber="67">
<subSubSection type="nomenclature">
<paragraph pageId="36" pageNumber="67">
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clathrocaspia knipowitschii" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="species" species="knipowitschii">Clathrocaspia knipowitschii</taxonomicName>
(Makarov, 1938)
</paragraph>
</subSubSection>
<subSubSection type="reference_group">
<paragraph pageId="36" pageNumber="67">
*1938
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia gmelini" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="species" species="gmelini">Caspia gmelini</taxonomicName>
[sic] var. Knipowitschii Makarov: 1058.
</paragraph>
<paragraph pageId="36" pageNumber="67">?1966 P. [yrgula] (Caspia) gmelini [sic] aluschtensis Golikov and Starobogatov: 354, fig. 1(8).</paragraph>
<paragraph pageId="36" pageNumber="67">1966 P. [yrgula] (Caspia) makarovi Golikov and Starobogatov: 353-354, fig. 1(5).</paragraph>
<paragraph pageId="36" pageNumber="67">
?1987
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia gmelinii" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="subspecies" species="gmelinii" subspecies="stanislavi">Caspia gmelinii stanislavi</taxonomicName>
Alexenko and Starobogatov: 33, fig. 1.
</paragraph>
<paragraph pageId="36" pageNumber="67">
1992
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia (Clathrocaspia) knipowitchi" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="species" species="knipowitchi" subGenus="Clathrocaspia">Caspia (Clathrocaspia) knipowitchi</taxonomicName>
Makarov, 1938. - Anistratenko and Prisyazhniuk: 19, fig. 2b.
</paragraph>
<paragraph pageId="36" pageNumber="67">
2006
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia knipowitchi" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="species" species="knipowitchi">Caspia knipowitchi</taxonomicName>
[sic] Makarov, 1938. - Kantor and Sysoev: 87-88, pl. 41, fig. J.
</paragraph>
<paragraph pageId="36" pageNumber="67">
2006
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia makarovi" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="species" species="makarovi">Caspia makarovi</taxonomicName>
(Golikov et Starobogatov, 1966). - Kantor and Sysoev: 88, pl. 41, fig. L.
</paragraph>
<paragraph pageId="36" pageNumber="67">
2013
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia knipowitchii" order="Littorinimorpha" pageId="36" pageNumber="67" phylum="Mollusca" rank="species" species="knipowitchii">Caspia knipowitchii</taxonomicName>
[sic] Makarov, 1938. - Anistratenko: 53-55, figs 1
<normalizedToken originalValue="AI">A-I</normalizedToken>
, 3
<normalizedToken originalValue="AD">A-D</normalizedToken>
, 5
<normalizedToken originalValue="AD">A-D</normalizedToken>
.
</paragraph>
<paragraph pageId="37" pageNumber="68">
<pageBreakToken pageId="37" pageNumber="68" start="start">2013</pageBreakToken>
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia makarovi" order="Littorinimorpha" pageId="37" pageNumber="68" phylum="Mollusca" rank="species" species="makarovi">Caspia makarovi</taxonomicName>
(Golikov &amp; Starobogatov, 1966). - Anistratenko: 56-59, figs 2
<normalizedToken originalValue="AE">A-E</normalizedToken>
, 3E.
</paragraph>
<paragraph pageId="37" pageNumber="68">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia knipowitchi" order="Littorinimorpha" pageId="37" pageNumber="68" phylum="Mollusca" rank="species" species="knipowitchi">Caspia knipowitchi</taxonomicName>
[sic] Makarov, 1938. - Vinarski and Kantor: 224.
</paragraph>
<paragraph pageId="37" pageNumber="68">
2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia makarovi" order="Littorinimorpha" pageId="37" pageNumber="68" phylum="Mollusca" rank="species" species="makarovi">Caspia makarovi</taxonomicName>
(Golikov &amp; Starobogatov, 1966). - Vinarski and Kantor: 225.
</paragraph>
<paragraph pageId="37" pageNumber="68">
?2016
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Caspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Caspia stanislavi" order="Littorinimorpha" pageId="37" pageNumber="68" phylum="Mollusca" rank="species" species="stanislavi">Caspia stanislavi</taxonomicName>
Alexenko &amp; Starobogatov, 1987. - Vinarski and Kantor: 225.
</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph pageId="37" pageNumber="68">Status. Accepted Pontocaspian species.</paragraph>
</subSubSection>
<subSubSection type="distribution">
<paragraph pageId="37" pageNumber="68">Type locality. Ukraine, in the Dniester River (exact locality not specified).</paragraph>
<paragraph pageId="37" pageNumber="68">
Distribution. Azov Sea and northern Black Sea Basin. Known from the Holocene of Danube Delta, Ukraine (
<bibRefCitation author="Anistratenko, VV" journalOrPublisher="Vestnik Zoologii" pageId="72" pageNumber="103" pagination="15 - 21" title="Novyye dannyye o mollyuskakh golotsenovykh otlozheniy Chernogo morya na Ukraine." volume="5" year="1992">Anistratenko and Prisyazhniuk 1992</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection type="discussion">
<paragraph pageId="37" pageNumber="68">
Taxonomic notes.
<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Clathrocaspia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clathrocaspia knipowitschii" order="Littorinimorpha" pageId="37" pageNumber="68" phylum="Mollusca" rank="species" species="knipowitschii">Clathrocaspia knipowitschii</taxonomicName>
,
<taxonomicName lsidName="C. makarovi" pageId="37" pageNumber="68" rank="species" species="makarovi">C. makarovi</taxonomicName>
,
<taxonomicName lsidName="C. gmelini" pageId="37" pageNumber="68" rank="subspecies" species="gmelini" subspecies="aluschtensis">C. gmelini aluschtensis</taxonomicName>
, and
<taxonomicName lsidName="C. stanislavi" pageId="37" pageNumber="68" rank="species" species="stanislavi">C. stanislavi</taxonomicName>
were all described from the northern margin of the Black Sea. After detailed morphological comparison of
<taxonomicName lsidName="C. knipowitschii" pageId="37" pageNumber="68" rank="species" species="knipowitschii">C. knipowitschii</taxonomicName>
and
<taxonomicName lsidName="C. makarovi" pageId="37" pageNumber="68" rank="species" species="makarovi">C. makarovi</taxonomicName>
syn. n. and preliminary genetic analyses (TW, unpublished data), we conclude that both taxa should be considered synonyms. Very likely, also
<taxonomicName lsidName="C. gmelini" pageId="37" pageNumber="68" rank="subspecies" species="gmelini" subspecies="aluschtensis">C. gmelini aluschtensis</taxonomicName>
and
<taxonomicName lsidName="C. stanislavi" pageId="37" pageNumber="68" rank="species" species="stanislavi">C. stanislavi</taxonomicName>
are synonyms of
<taxonomicName lsidName="C. knipowitschii" pageId="37" pageNumber="68" rank="species" species="knipowitschii">C. knipowitschii</taxonomicName>
, but a final decision on that matter requires investigation of the type material.
</paragraph>
</subSubSection>
<subSubSection type="conservation">
<paragraph pageId="37" pageNumber="68">
Conservation status. Least Concern (same for
<taxonomicName lsidName="C. makarovi" pageId="37" pageNumber="68" rank="species" species="makarovi">C. makarovi</taxonomicName>
;
<bibRefCitation author="Son, M" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="82" pageNumber="113" url="https://doi.org/10.2305/IUCN.UK.2011-1.RLTS.T156116A4900657.en" year="2011 b">Son 2011b</bibRefCitation>
,
<bibRefCitation author="Son, M" journalOrPublisher="Zoologicheskiy Zhurnal" pageId="82" pageNumber="113" url="https://doi.org/10.2305/IUCN.UK.2011-1.RLTS.T155680A4822960.en" year="2011 c">c</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>