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<document id="261194C074D9055EC68DFA113B34CA98" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFCEFFFCFF04FD3BFE0CF8F5" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tuberculobasis macuxi Machado, 2009, sp. nov." docType="treatment" docVersion="12" lastPageNumber="24" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="24" updateTime="1698585892014" updateUser="plazi">
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<mods:title id="72DD80BC4F9CAB4C1A0CDD86978E7205">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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<mods:name id="0CEC2DC6D16DCC65D2036EE01490F0C6" type="personal">
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<mods:namePart id="F54F003DD06395D12444588751E5DB98">Machado, Angelo B. M.</mods:namePart>
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<mods:date id="544C21FB10EF59257BC4D5F3DB9690B4">2009</mods:date>
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<mods:number id="4E835F3BB73B3C16C8D2DBC694982668">2108</mods:number>
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<treatment id="1312774BFFCEFFFCFF04FD3BFE0CF8F5" ID-DOI="http://doi.org/10.5281/zenodo.4391379" ID-GBIF-Taxon="119617255" ID-Zenodo-Dep="4391379" LSID="urn:lsid:plazi:treatment:1312774BFFCEFFFCFF04FD3BFE0CF8F5" httpUri="http://treatment.plazi.org/id/1312774BFFCEFFFCFF04FD3BFE0CF8F5" lastPageNumber="24" pageId="23" pageNumber="24">
<subSubSection id="D3A195D6FFCEFFFCFF04FD3BFE30FCBE" pageId="23" pageNumber="24" type="nomenclature">
<paragraph id="9B04C65DFFCEFFFCFF04FD3BFDB2FD53" blockId="23.[151,533,731,792]" box="[151,533,731,758]" pageId="23" pageNumber="24">
<heading id="C04C7131FFCEFFFCFF04FD3BFDB2FD53" bold="true" box="[151,533,731,758]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
<emphasis id="A9CF1A4FFFCEFFFCFF04FD3BFDB2FD53" bold="true" box="[151,533,731,758]" pageId="23" pageNumber="24">
<taxonomicName id="5CBBBDDEFFCEFFFCFF04FD3BFE17FD50" ID-CoL="7D9XK" box="[151,432,732,758]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="macuxi" status="sp. nov.">
<emphasis id="A9CF1A4FFFCEFFFCFF04FD3BFE17FD50" bold="true" box="[151,432,732,758]" italics="true" pageId="23" pageNumber="24">Tuberculobasis macuxi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B2FCA734FFCEFFFCFE24FD3CFDB2FD53" box="[439,533,731,757]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="9B04C65DFFCEFFFCFF04FD19FE30FCBE" blockId="23.[151,533,731,792]" box="[151,407,766,792]" pageId="23" pageNumber="24">
<figureCitation id="0380DAD8FFCEFFFCFF04FD19FEE4FCBE" box="[151,323,766,792]" captionStart="FIGURES 35 46" captionStartId="19.[151,271,1939,1963]" captionTargetBox="[375,1177,183,1913]" captionTargetId="figure@19.[363,1191,178,1917]" captionTargetPageId="19" captionText="FIGURES 35 46. Tuberculobasis, male thorax in dorsal and lateral view: T. geijskesi, holotype (35, 36); T. karitiana, holotype (37, 38); T. mammilaris, Venezuela (39, 40); T. tirio, holotype (41, 42); T. yanomami, paratype (43, 44); T. macuxi, holotype (45, 46)." httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="23" pageNumber="24">Figures 4546</figureCitation>
,
<figureCitation id="0380DAD8FFCEFFFCFEDDFD19FE30FCBE" box="[334,407,766,792]" captionStart="FIGURES 69 77" captionStartId="25.[151,272,1670,1694]" captionTargetBox="[151,1426,187,1662]" captionTargetId="figure@25.[151,1436,138,1748]" captionTargetPageId="25" captionText="FIGURES 69 77. Tuberculobasis male, anal appendages in dorsal, lateral, and posterior views: T. macuxi, holotype (69 71); T. williamsoni, holotype (72 74); T. geijskesi, holotype (75 77)." httpUri="https://zenodo.org/record/187818/files/figure.png" pageId="23" pageNumber="24">6971</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFCEFFFCFF04FCACFE9AFC2A" pageId="23" pageNumber="24" type="etymology">
<paragraph id="9B04C65DFFCEFFFCFF04FCACFE9AFC2A" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF04FCACFE8EFCC3" bold="true" box="[151,297,843,869]" pageId="23" pageNumber="24">Etymology:</emphasis>
The name refers to the
<taxonomicName id="5CBBBDDEFFCEFFFCFDD4FCABFD06FCC0" box="[583,673,844,870]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="34" phylum="Arthropoda" rank="species" species="macuxi">Macuxi</taxonomicName>
indian people, who are settled in the region where the
<typeStatus id="440078FFFFCEFFFCFAA7FCABFA3CFCC0" box="[1332,1435,844,870]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
was collected.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFCEFFFCFF55FC7FFAA5FC7C" pageId="23" pageNumber="24" type="materials_examined">
<paragraph id="9B04C65DFFCEFFFCFF55FC7FFAA5FC7C" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55FC7FFE28FC14" bold="true" box="[198,399,920,946]" pageId="23" pageNumber="24">
<typeStatus id="440078FFFFCEFFFCFF55FC7FFEA4FC14" box="[198,259,920,946]" pageId="23" pageNumber="24">Type</typeStatus>
specimens
</emphasis>
:
<emphasis id="A9CF1A4FFFCEFFFCFE30FC7FFDB2FC14" bold="true" box="[419,533,920,946]" pageId="23" pageNumber="24">
<typeStatus id="440078FFFFCEFFFCFE30FC7FFDB2FC14" box="[419,533,920,946]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
</emphasis>
(3):
<collectingCountry id="E3AC86CDFFCEFFFCFDF3FC7EFD6AFC15" box="[608,717,921,947]" name="Brazil" pageId="23" pageNumber="24">BRAZIL</collectingCountry>
, Roraima State, Maracá Ecological Station, Maracá Island (
<geoCoordinate id="FE8FA09AFFCEFFFCFF33FC27FF50FC7C" box="[160,247,960,986]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="925" value="3.4166667">3º25´N</geoCoordinate>
,
<geoCoordinate id="FE8FA09AFFCEFFFCFE90FC27FED7FC7C" box="[259,368,960,986]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="925" value="-61.6">61º36´W</geoCoordinate>
),
<date id="EF05E09DFFCEFFFCFE17FC27FDA0FC7C" box="[388,519,960,986]" pageId="23" pageNumber="24" value="1988-02-24">24-II-1988</date>
. P.A.R. Machado leg (within forest, away from water) (
<collectionCode id="FDAA5E98FFCEFFFCFB01FC27FB51FC7C" box="[1170,1270,960,986]" pageId="23" pageNumber="24">ABMM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFCEFFFCFF55FC01FD4EF91E" pageId="23" pageNumber="24" type="description">
<paragraph id="9B04C65DFFCEFFFCFF55FC01FDBFFBA6" blockId="23.[151,1437,843,1875]" box="[198,536,998,1024]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55FC01FEF9FBA6" bold="true" box="[198,350,998,1024]" pageId="23" pageNumber="24">Description.</emphasis>
Male
<typeStatus id="440078FFFFCEFFFCFE3AFC01FDB4FBA6" box="[425,531,998,1024]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="9B04C65DFFCEFFFCFF55FBEAFCC9FBD2" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55FBEAFEAEFB81" bold="true" box="[198,265,1037,1063]" pageId="23" pageNumber="24">Head</emphasis>
. Labium yellowish white. Labrum and anteclypeus yellow; genae and base of mandibles grey; postclypeus, antefrons, and top of head brown; postocular lobe yellowish ochre. A subtriangular blue postocular spot adjacent to eye. Rear of head yellowish white.
</paragraph>
<paragraph id="9B04C65DFFCEFFFCFF55FB67FE99FAFA" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55FB67FE8DFB3C" bold="true" box="[198,298,1152,1178]" pageId="23" pageNumber="24">Thorax.</emphasis>
Prothorax: Pronotum yellow ochre with bluish markings on anterior lobe. Propleuron yellowish with greyish areas. Pterothorax: mesopleuron and metepisternum yellow ochre with a narrow light blue stripe at middle part of mesepisternum and at anterior part of metepisternum. Metepimeron yellowish white. Legs yellowish white, wings hyaline, venation brown. Pterostigma yellow. Px in FW 11, in HW 10. R3 originating between Px 5 and
<quantity id="5C436BB8FFCEFFFCFEFEFAFBFE3EFA90" box="[365,409,1308,1334]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="23" pageNumber="24" unit="in" value="6.0">6 in</quantity>
both wings. Petiolation distal to Ac by a distance equal or twice as long as length of Ac in both wings.
</paragraph>
<paragraph id="9B04C65DFFCEFFFCFF55FA8FFE3AFA76" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55FA8FFEE7FA24" bold="true" box="[198,320,1384,1410]" pageId="23" pageNumber="24">Abdomen</emphasis>
. S12 dorsally yellow ochre, laterally light blue, ventrally yellowish white with a dark brown terminal line. S37 light brown dorsally, laterally yellow with a dark brown distal ring. S810, cercus, and paraproct light brown.
</paragraph>
<paragraph id="9B04C65DFFCEFFFCFF55FA3AFBE4F934" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55FA3AFE72FA51" bold="true" box="[198,469,1501,1527]" pageId="23" pageNumber="24">Structural characters</emphasis>
.
<collectingCountry id="E3AC86CDFFCEFFFCFE75FA3AFD86FA51" box="[486,545,1501,1527]" name="India" pageId="23" pageNumber="24">Hind</collectingCountry>
prothoracic lobe with well-developed lateral lobes (
<figureCitation id="0380DAD8FFCEFFFCFB06FA3AFB48FA51" box="[1173,1263,1501,1527]" captionStart="FIGURES 35 46" captionStartId="19.[151,271,1939,1963]" captionTargetBox="[375,1177,183,1913]" captionTargetId="figure@19.[363,1191,178,1917]" captionTargetPageId="19" captionText="FIGURES 35 46. Tuberculobasis, male thorax in dorsal and lateral view: T. geijskesi, holotype (35, 36); T. karitiana, holotype (37, 38); T. mammilaris, Venezuela (39, 40); T. tirio, holotype (41, 42); T. yanomami, paratype (43, 44); T. macuxi, holotype (45, 46)." httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="23" pageNumber="24">Fig. 45</figureCitation>
). Median lobe semicircular and two-lipped; ventral lip more produced than dorsal lip. Mesepisternal tubercles (
<figureCitation id="0380DAD8FFCEFFFCFA98F9E3FA34F9B8" box="[1291,1427,1540,1566]" captionStart="FIGURES 35 46" captionStartId="19.[151,271,1939,1963]" captionTargetBox="[375,1177,183,1913]" captionTargetId="figure@19.[363,1191,178,1917]" captionTargetPageId="19" captionText="FIGURES 35 46. Tuberculobasis, male thorax in dorsal and lateral view: T. geijskesi, holotype (35, 36); T. karitiana, holotype (37, 38); T. mammilaris, Venezuela (39, 40); T. tirio, holotype (41, 42); T. yanomami, paratype (43, 44); T. macuxi, holotype (45, 46)." httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="23" pageNumber="24">Figs 45, 46</figureCitation>
) subcylindrical, high (1.51 mm), directed dorsally, with bases well separated from mid-dorsal carina. Cercus about 1/2 shorter than paraproct (
<figureCitation id="0380DAD8FFCEFFFCFDB9F9B6FD19F9CD" box="[554,702,1617,1643]" captionStart="FIGURES 69 77" captionStartId="25.[151,272,1670,1694]" captionTargetBox="[151,1426,187,1662]" captionTargetId="figure@25.[151,1436,138,1748]" captionTargetPageId="25" captionText="FIGURES 69 77. Tuberculobasis male, anal appendages in dorsal, lateral, and posterior views: T. macuxi, holotype (69 71); T. williamsoni, holotype (72 74); T. geijskesi, holotype (75 77)." httpUri="https://zenodo.org/record/187818/files/figure.png" pageId="23" pageNumber="24">Figs. 6970</figureCitation>
), ventral process in lateral view tapering into a chitinized tip directed latero-posteriorly (
<figureCitation id="0380DAD8FFCEFFFCFE4BF99FFD88F934" box="[472,559,1656,1682]" captionStart="FIGURES 69 77" captionStartId="25.[151,272,1670,1694]" captionTargetBox="[151,1426,187,1662]" captionTargetId="figure@25.[151,1436,138,1748]" captionTargetPageId="25" captionText="FIGURES 69 77. Tuberculobasis male, anal appendages in dorsal, lateral, and posterior views: T. macuxi, holotype (69 71); T. williamsoni, holotype (72 74); T. geijskesi, holotype (75 77)." httpUri="https://zenodo.org/record/187818/files/figure.png" pageId="23" pageNumber="24">Fig. 71</figureCitation>
). Penis as described for genus (as in
<figureCitation id="0380DAD8FFCEFFFCFC4CF99FFB91F934" box="[991,1078,1656,1682]" captionStart="FIGURES 87 94" captionStartId="29.[151,269,1856,1880]" captionTargetBox="[282,1311,194,1855]" captionTargetId="figure@29.[273,1313,117,1921]" captionTargetPageId="29" captionText="FIGURES 87 94. Penis in lateral and ectal views: D. dunklei, holotype (87); D. garrisoni, holotype (88); T. costalimai, paratype (89); T. geijskesi, holotype (90); T. mammilaris, lectotype (91); T. tirio, paratype (92); T. williamsoni, paratype (93); Leptobasis vacillans, Panamá (94) [91 by R. W. Garrison]." httpUri="https://zenodo.org/record/187820/files/figure.png" pageId="23" pageNumber="24">Fig. 89</figureCitation>
).
</paragraph>
<paragraph id="9B04C65DFFCEFFFCFF55F979FD4EF91E" blockId="23.[151,1437,843,1875]" box="[198,745,1694,1720]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55F979FE0CF91E" bold="true" box="[198,427,1694,1720]" pageId="23" pageNumber="24">Dimensions (mm).</emphasis>
Abdomen 21.9; HW 15.35
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFCEFFFCFF55F922FB22F88A" pageId="23" pageNumber="24" type="discussion">
<paragraph id="9B04C65DFFCEFFFCFF55F922FB22F88A" blockId="23.[151,1437,843,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55F922FE9FF979" bold="true" box="[198,312,1733,1759]" pageId="23" pageNumber="24">Remarks</emphasis>
.
<taxonomicName id="5CBBBDDEFFCEFFFCFEDBF922FE1DF978" box="[328,442,1733,1758]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="macuxi">
<emphasis id="A9CF1A4FFFCEFFFCFEDBF922FE1DF978" box="[328,442,1733,1758]" italics="true" pageId="23" pageNumber="24">T. macuxi</emphasis>
</taxonomicName>
has high mesepisternal tubercles with bases well separated from mid-dorsal carina, long paraprocts, and hind prothoracic lobe with well-developed lateral lobes. This combination makes it unique within the genus belonging to none of the two species groups recognized here.
</paragraph>
</subSubSection>
<subSubSection id="D3A195D6FFCEFFFCFF55F8DFFE0CF8F5" box="[198,427,1848,1875]" pageId="23" pageNumber="24" type="description">
<paragraph id="9B04C65DFFCEFFFCFF55F8DFFE0CF8F5" blockId="23.[151,1437,843,1875]" box="[198,427,1848,1875]" pageId="23" pageNumber="24">
<emphasis id="A9CF1A4FFFCEFFFCFF55F8DFFE8EF8F4" bold="true" box="[198,297,1848,1874]" pageId="23" pageNumber="24">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>