292 lines
32 KiB
XML
292 lines
32 KiB
XML
<document id="5BA1C78C88E558A4F36AEC21EBD3A2B2" ID-DOI="10.5281/zenodo.187806" ID-GBIF-Dataset="9a495f32-3eb8-4b35-bd46-adca3b5db2d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="187806" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461074216081" checkinUser="plazi" docAuthor="Machado, Angelo B. M." docDate="2009" docId="1312774BFFC8FFFEFF04F979FC8CFCE5" docLanguage="en" docName="zt02108p036.pdf" docOrigin="Zootaxa 2108" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tuberculobasis guarani Machado, 2009, sp. nov." docType="treatment" docVersion="12" lastPageNumber="22" masterDocId="EF2B0F33FFD9FFEBFF93FFE7FFA7FFA6" masterDocTitle="Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae)." masterLastPageNumber="36" masterPageNumber="1" pageNumber="18" updateTime="1698585892014" updateUser="plazi">
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<mods:mods id="527ED1A1A2475B442EBA220B0D388815" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FFBEF97EF14A67C543DA3C7E80344B13">
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<mods:title id="DC0E2EB5AAA5FCF600CBD41E2C1B5BD5">Denticulobasis and Tuberculobasis, new genera close to Leptobasis, with description of ten new species (Odonata: Coenagrionidae).</mods:title>
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</mods:titleInfo>
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<mods:name id="8A6D0DC8D917B05DF460BE224890F52E" type="personal">
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<mods:role id="D330E12E80E8E1449467A13177FDDE63">
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<mods:roleTerm id="A5B162F1E2C3A8D95EE04B468412AF0B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C3443D58398C318BD98E0B1BD99EF06D">Machado, Angelo B. M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A167F3284C4AF425B26E5DE3726CCFE1">text</mods:typeOfResource>
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<mods:titleInfo id="0E550F0C74D80260A689503D32B3FEFB">
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<mods:title id="63B0C4096FD118D40E25F233F890BF9E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="8DBEA5E46E1297A0B60A1D145311FCE0">
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<mods:date id="BBB5B13AC5FA964F5EE19357451FF989">2009</mods:date>
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<mods:detail id="DA5A3C2EA5E5C1B536CED080EE4647BF" type="volume">
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<mods:number id="7F4892DF96816A7B3843172988351B94">2108</mods:number>
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</mods:detail>
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<mods:extent id="4FCFF19AF5FD05EFE3071A667406F423" unit="page">
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<mods:start id="BE72E995F4423DCA7334FB8CE92E9121">1</mods:start>
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<mods:end id="7594376AA5BBCFE8A94DB396792152E1">36</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="68DB14AB3758EB12CD98E5AB241B1326">journal article</mods:classification>
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<mods:identifier id="524F182E2BD06A5BD85D868F2C893D58" type="DOI">10.5281/zenodo.187806</mods:identifier>
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<mods:identifier id="5347CAF9CF5C4CBE8FCDBAEF300D6CE4" type="GBIF-Dataset">9a495f32-3eb8-4b35-bd46-adca3b5db2d2</mods:identifier>
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<mods:identifier id="8CA431ECBC5F0AEDFB8FCCE54E2020E7" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C4F6AE9260B8F984C5581F164DB22466" type="Zenodo-Dep">187806</mods:identifier>
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<treatment id="1312774BFFC8FFFEFF04F979FC8CFCE5" ID-DOI="http://doi.org/10.5281/zenodo.4391373" ID-GBIF-Taxon="119617259" ID-Zenodo-Dep="4391373" LSID="urn:lsid:plazi:treatment:1312774BFFC8FFFEFF04F979FC8CFCE5" httpUri="http://treatment.plazi.org/id/1312774BFFC8FFFEFF04F979FC8CFCE5" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18">
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||
<subSubSection id="D3A195D6FFC8FFFAFF04F979FE30F97D" pageId="17" pageNumber="18" type="nomenclature">
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||
<paragraph id="9B04C65DFFC8FFFAFF04F979FDBCF91E" blockId="17.[151,539,1694,1755]" box="[151,539,1694,1720]" pageId="17" pageNumber="18">
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||
<heading id="C04C7131FFC8FFFAFF04F979FDBCF91E" bold="true" box="[151,539,1694,1720]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
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||
<emphasis id="A9CF1A4FFFC8FFFAFF04F979FDBCF91E" bold="true" box="[151,539,1694,1720]" pageId="17" pageNumber="18">
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||
<taxonomicName id="5CBBBDDEFFC8FFFAFF04F979FE10F91E" ID-CoL="7D9XJ" box="[151,439,1694,1720]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="guarani" status="sp. nov.">
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||
<emphasis id="A9CF1A4FFFC8FFFAFF04F979FE10F91E" bold="true" box="[151,439,1694,1720]" italics="true" pageId="17" pageNumber="18">Tuberculobasis guarani</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="B2FCA734FFC8FFFAFE2DF979FDBCF91E" box="[446,539,1694,1720]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="9B04C65DFFC8FFFAFF04F926FE30F97D" blockId="17.[151,539,1694,1755]" box="[151,407,1729,1755]" pageId="17" pageNumber="18">
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||
<figureCitation id="0380DAD8FFC8FFFAFF04F926FEE5F97D" box="[151,322,1729,1755]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="17" pageNumber="18">Figures 29–30</figureCitation>
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,
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<figureCitation id="0380DAD8FFC8FFFAFEDDF926FE30F97D" box="[334,407,1729,1755]" captionStart="FIGURES 57 – 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 – 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 – 59); T. costalimai, paratypes (60 – 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="17" pageNumber="18">64–65</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A195D6FFC8FFFAFF04F8E9FF46F8E9" pageId="17" pageNumber="18" type="etymology">
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<paragraph id="9B04C65DFFC8FFFAFF04F8E9FF46F8E9" blockId="17.[151,1436,1806,1987]" pageId="17" pageNumber="18">
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<emphasis id="A9CF1A4FFFC8FFFAFF04F8E9FE8FF88E" bold="true" box="[151,296,1806,1832]" pageId="17" pageNumber="18">Etymology:</emphasis>
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The name refers to the
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<taxonomicName id="5CBBBDDEFFC8FFFAFDD7F8E9FD04F88E" box="[580,675,1806,1832]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="17" pageNumber="32" phylum="Arthropoda" rank="species" species="guarani">Guarani</taxonomicName>
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indian people who still inhabit various areas of the State of São Paulo.
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A195D6FFC8FFFAFF55F8BCFD18F83A" pageId="17" pageNumber="18" type="materials_examined">
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||
<paragraph id="9B04C65DFFC8FFFAFF55F8BCFD18F83A" blockId="17.[151,1436,1806,1987]" pageId="17" pageNumber="18">
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<emphasis id="A9CF1A4FFFC8FFFAFF55F8BCFD88F8D3" bold="true" box="[198,559,1883,1909]" pageId="17" pageNumber="18">
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<typeStatus id="440078FFFFC8FFFAFF55F8BCFEA4F8D3" box="[198,259,1883,1909]" pageId="17" pageNumber="18">Type</typeStatus>
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specimens:
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<typeStatus id="440078FFFFC8FFFAFE35F8BCFDBFF8D3" box="[422,536,1883,1909]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
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(
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</emphasis>
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3
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<emphasis id="A9CF1A4FFFC8FFFAFDCDF8BCFDC8F8D3" bold="true" box="[606,623,1883,1909]" pageId="17" pageNumber="18">).</emphasis>
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<collectingCountry id="E3AC86CDFFC8FFFAFDEFF8BBFD4EF8D0" box="[636,745,1884,1910]" name="Brazil" pageId="17" pageNumber="18">BRAZIL</collectingCountry>
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, São Paulo State, Ibitinga (
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<geoCoordinate id="FE8FA09AFFC8FFFAFBD2F8BBFB05F8D0" box="[1089,1186,1884,1910]" direction="south" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="-21.75">21º45´S</geoCoordinate>
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,
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<geoCoordinate id="FE8FA09AFFC8FFFAFB20F8BBFA87F8D0" box="[1203,1312,1884,1910]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-48.816666">48º49´W</geoCoordinate>
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), Fazenda Itaguaré,
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<date id="EF05E09DFFC8FFFAFE96F865FE35F83A" box="[261,402,1922,1948]" pageId="17" pageNumber="18" value="1965-06-17">17-VI-1965</date>
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, K. Lenko leg. (
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<collectionCode id="FDAA5E98FFC8FFFAFDDCF865FD14F83A" box="[591,691,1922,1948]" pageId="17" pageNumber="18">ABMM</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="D3A195D6FFC8FFFEFF55F84FFE0CFD24" lastPageId="21" lastPageNumber="22" pageId="17" pageNumber="18" type="description">
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<paragraph id="9B04C65DFFC8FFFAFF55F84FFDB7F865" blockId="17.[151,1436,1806,1987]" box="[198,528,1960,1987]" pageId="17" pageNumber="18">
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<emphasis id="A9CF1A4FFFC8FFFAFF55F84FFEF0F864" bold="true" box="[198,343,1960,1986]" pageId="17" pageNumber="18">Description</emphasis>
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.Male
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<typeStatus id="440078FFFFC8FFFAFE31F84EFDABF865" box="[418,524,1961,1987]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<caption id="CFC496D5FFCBFFF9FF04F852FA98F856" httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="18" pageNumber="19" targetBox="[327,1275,181,1941]" targetPageId="18">
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<paragraph id="9B04C65DFFCBFFF9FF04F852FA98F856" blockId="18.[151,1436,1973,2032]" pageId="18" pageNumber="19">
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<emphasis id="A9CF1A4FFFCBFFF9FF04F852FEC7F86B" bold="true" box="[151,352,1973,1997]" pageId="18" pageNumber="19">FIGURES 25–34.</emphasis>
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<emphasis id="A9CF1A4FFFCBFFF9FEF4F851FDB6F86B" box="[359,529,1974,1997]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="5CBBBDDEFFCBFFF9FEF4F851FDAAF86B" box="[359,525,1974,1997]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">Tuberculobasis</taxonomicName>
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,
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</emphasis>
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male thorax in dorsal and lateral view in
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<emphasis id="A9CF1A4FFFCBFFF9FC4BF851FB90F86B" box="[984,1079,1974,1997]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="5CBBBDDEFFCBFFF9FC4BF851FB94F86B" box="[984,1075,1974,1997]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="arara">T. arara</taxonomicName>
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,
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</emphasis>
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holotype (25, 26);
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<emphasis id="A9CF1A4FFFCBFFF9FA99F851FA3BF86B" box="[1290,1436,1974,1997]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="5CBBBDDEFFCBFFF9FA99F851FA3FF86B" box="[1290,1432,1974,1997]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
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,
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</emphasis>
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holotype (27, 28);
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<emphasis id="A9CF1A4FFFCBFFF9FECEF83EFE74F856" box="[349,467,2009,2032]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="5CBBBDDEFFCBFFF9FECEF83EFE69F856" box="[349,462,2009,2032]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="guarani">T. guarani</taxonomicName>
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,
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</emphasis>
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holotype (29, 30);
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<emphasis id="A9CF1A4FFFCBFFF9FD0CF83EFCA8F856" box="[671,783,2009,2032]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="5CBBBDDEFFCBFFF9FD0CF83EFCACF856" box="[671,779,2009,2032]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="inversa">T. inversa</taxonomicName>
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,
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</emphasis>
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holotype (31, 32);
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<emphasis id="A9CF1A4FFFCBFFF9FC4FF83EFBDEF856" box="[988,1145,2009,2032]" italics="true" pageId="18" pageNumber="19">
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<taxonomicName id="5CBBBDDEFFCBFFF9FC4FF83EFBD2F856" box="[988,1141,2009,2032]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="williamsoni">T. williamsoni</taxonomicName>
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,
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</emphasis>
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holotype (33, 34).
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</paragraph>
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</caption>
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<caption id="CFC496D5FFCAFFF8FF04F874FE17F849" httpUri="https://zenodo.org/record/187814/files/figure.png" pageId="19" pageNumber="20" targetBox="[375,1177,183,1913]" targetPageId="19">
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<paragraph id="9B04C65DFFCAFFF8FF04F874FE17F849" blockId="19.[151,1436,1939,2032]" pageId="19" pageNumber="20">
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<emphasis id="A9CF1A4FFFCAFFF8FF04F874FEC7F80D" bold="true" box="[151,352,1939,1963]" pageId="19" pageNumber="20">FIGURES 35–46.</emphasis>
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<taxonomicName id="5CBBBDDEFFCAFFF8FEF4F874FDACF80C" box="[359,523,1939,1962]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
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<emphasis id="A9CF1A4FFFCAFFF8FEF4F874FDACF80C" box="[359,523,1939,1962]" italics="true" pageId="19" pageNumber="20">Tuberculobasis</emphasis>
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||
</taxonomicName>
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, male thorax in dorsal and lateral view:
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<emphasis id="A9CF1A4FFFCAFFF8FC56F874FBE3F80C" box="[965,1092,1939,1962]" italics="true" pageId="19" pageNumber="20">
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<taxonomicName id="5CBBBDDEFFCAFFF8FC56F874FBE7F80C" box="[965,1088,1939,1962]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="geijskesi">T. geijskesi</taxonomicName>
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,
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</emphasis>
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holotype (35, 36);
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<emphasis id="A9CF1A4FFFCAFFF8FA84F874FA3BF80C" box="[1303,1436,1939,1962]" italics="true" pageId="19" pageNumber="20">
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||
<taxonomicName id="5CBBBDDEFFCAFFF8FA84F874FA3FF80C" box="[1303,1432,1939,1962]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="karitiana">T. karitiana</taxonomicName>
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,
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||
</emphasis>
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||
holotype (37, 38);
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||
<emphasis id="A9CF1A4FFFCAFFF8FEF5F851FDABF86B" box="[358,524,1974,1997]" italics="true" pageId="19" pageNumber="20">
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||
<taxonomicName id="5CBBBDDEFFCAFFF8FEF5F851FDAFF86B" box="[358,520,1974,1997]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="mammilaris">T. mammilaris</taxonomicName>
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||
,
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||
</emphasis>
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Venezuela (39, 40);
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||
<emphasis id="A9CF1A4FFFCAFFF8FD67F851FCE3F86B" box="[756,836,1974,1997]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="5CBBBDDEFFCAFFF8FD67F851FC99F86B" box="[756,830,1974,1997]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="tirio">T. tirio</taxonomicName>
|
||
,
|
||
</emphasis>
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||
holotype (41, 42);
|
||
<emphasis id="A9CF1A4FFFCAFFF8FB8EF851FB14F86B" box="[1053,1203,1974,1997]" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="5CBBBDDEFFCAFFF8FB8EF851FB08F86B" box="[1053,1199,1974,1997]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="yanomami">T. yanomami</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype (43, 44);
|
||
<emphasis id="A9CF1A4FFFCAFFF8FA1BF851FF4CF856" italics="true" pageId="19" pageNumber="20">
|
||
<taxonomicName id="5CBBBDDEFFCAFFF8FA1BF851FF40F856" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="macuxi">T. macuxi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype (45, 46).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CFC496D5FFCDFFFFFF04F93CFD95F8FD" httpUri="https://zenodo.org/record/187815/files/figure.png" pageId="20" pageNumber="21" targetBox="[188,1374,193,1754]" targetPageId="20">
|
||
<paragraph id="9B04C65DFFCDFFFFFF04F93CFD95F8FD" blockId="20.[151,1437,1755,1883]" pageId="20" pageNumber="21">
|
||
<emphasis id="A9CF1A4FFFCDFFFFFF04F93CFEC5F955" bold="true" box="[151,354,1755,1779]" pageId="20" pageNumber="21">FIGURES 47–54.</emphasis>
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<taxonomicName id="5CBBBDDEFFCDFFFFFEF9F93BFDB7F955" box="[362,528,1756,1779]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
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||
<emphasis id="A9CF1A4FFFCDFFFFFEF9F93BFDB7F955" box="[362,528,1756,1779]" italics="true" pageId="20" pageNumber="21">Tuberculobasis</emphasis>
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</taxonomicName>
|
||
female thorax, in dorsal view:
|
||
<emphasis id="A9CF1A4FFFCDFFFFFCF9F93BFC5CF955" box="[874,1019,1756,1779]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFCF9F93BFC50F955" box="[874,1015,1756,1779]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="cardinalis">T. cardinalis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype (47);
|
||
<emphasis id="A9CF1A4FFFCDFFFFFB34F93BFA9BF955" box="[1191,1340,1756,1779]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFB34F93BFA9FF955" box="[1191,1336,1756,1779]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
allotype (48);
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFF42F918FE9DF8B0" box="[209,314,1791,1814]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="inversa">
|
||
<emphasis id="A9CF1A4FFFCDFFFFFF42F918FE9DF8B0" box="[209,314,1791,1814]" italics="true" pageId="20" pageNumber="21">T. inversa</emphasis>
|
||
</taxonomicName>
|
||
, Peru, Loreto (49);
|
||
<emphasis id="A9CF1A4FFFCDFFFFFD9EF918FD0AF8B0" box="[525,685,1791,1814]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFD9EF918FD0EF8B0" box="[525,681,1791,1814]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="mammilaris">T. mammilaris</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
Venezuela (50);
|
||
<emphasis id="A9CF1A4FFFCDFFFFFCF1F918FC08F8B0" box="[866,943,1791,1814]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFCF1F918FC0FF8B0" box="[866,936,1791,1814]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="tirio">T. tirio</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
allotype (51);
|
||
<emphasis id="A9CF1A4FFFCDFFFFFBDDF918FB4CF8B0" box="[1102,1259,1791,1814]" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFBDDF918FB40F8B0" box="[1102,1255,1791,1814]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="williamsoni">T. williamsoni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
allotype (52);
|
||
<emphasis id="A9CF1A4FFFCDFFFFFA1BF918FEB5F89E" italics="true" pageId="20" pageNumber="21">
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFA1BF918FEA9F89E" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="yanomami">T. yanomami</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
paratype (53), and in dorso-lateral view:
|
||
<taxonomicName id="5CBBBDDEFFCDFFFFFD7FF8C6FC30F89E" box="[748,919,1825,1848]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A9CF1A4FFFCDFFFFFD7FF8C6FC30F89E" box="[748,919,1825,1848]" italics="true" pageId="20" pageNumber="21">Tuberculobasis</emphasis>
|
||
</taxonomicName>
|
||
sp. 1, Manaus, Amazonas (54) [47 by N. von Ellenrieder; 50, 53 by J. De Marmels].
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9B04C65DFFCDFFFFFF55F86AFE33F868" blockId="20.[151,1436,1933,2036]" pageId="20" pageNumber="21">
|
||
<emphasis id="A9CF1A4FFFCDFFFFFF55F86AFEAEF801" bold="true" box="[198,265,1933,1959]" pageId="20" pageNumber="21">Head</emphasis>
|
||
. Labium whitish yellow; labrum dark green; antefrons and top of head dark brown with rounded pale postocular spots.
|
||
</paragraph>
|
||
<paragraph id="9B04C65DFFCDFFFFFF55F83DFBB8F852" blockId="20.[151,1436,1933,2036]" box="[198,1055,2010,2036]" pageId="20" pageNumber="21">
|
||
<emphasis id="A9CF1A4FFFCDFFFFFF55F83DFE8DF852" bold="true" box="[198,298,2010,2036]" pageId="20" pageNumber="21">Thorax.</emphasis>
|
||
Prothorax yellowish brown except for dark brown anterior lobe.
|
||
</paragraph>
|
||
<paragraph id="9B04C65DFFCCFFFEFF55FF7FFDBAFE80" blockId="21.[151,1437,152,835]" pageId="21" pageNumber="22">
|
||
Thorax, pterothorax: mesopleuron brown; metapleuron yellowish. Poorly defined grey stripes at posterior part of mesepisternum and metepisternum. Legs yellowish, wings hyaline, venation and pterostigma brown, Px in FW 11, in HW 10. R3 arising near Px
|
||
<quantity id="5C436BB8FFCCFFFEFD36FF02FD74FF59" box="[677,723,229,255]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="21" pageNumber="22" unit="in" value="5.0">5 in</quantity>
|
||
FW, near Px
|
||
<quantity id="5C436BB8FFCCFFFEFCE6FF02FC04FF59" box="[885,931,229,255]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="21" pageNumber="22" unit="in" value="4.0">4 in</quantity>
|
||
HW. Petiolation distal to Ac by a distance equal to Ac length in both wings.
|
||
</paragraph>
|
||
<paragraph id="9B04C65DFFCCFFFEFF55FED5FB50FED5" blockId="21.[151,1437,152,835]" pageId="21" pageNumber="22">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFF55FED5FEE7FEEA" bold="true" box="[198,320,306,332]" pageId="21" pageNumber="22">Abdomen</emphasis>
|
||
. S1–2 dorsally brownish yellow ventrally yellowish with a grey dorso-lateral stripe. S3–6 dorsally brown, laterally yellow, with a dark brown ring at distal 1/7. S70–10 yellowish orange.
|
||
</paragraph>
|
||
<paragraph id="9B04C65DFFCCFFFEFF55FE98FC80FD92" blockId="21.[151,1437,152,835]" pageId="21" pageNumber="22">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFF55FE98FE72FE3F" bold="true" box="[198,469,383,409]" pageId="21" pageNumber="22">Structural characters</emphasis>
|
||
.
|
||
<collectingCountry id="E3AC86CDFFCCFFFEFE74FE67FD85FE3C" box="[487,546,384,410]" name="India" pageId="21" pageNumber="22">Hind</collectingCountry>
|
||
prothoracic lobe with very small lateral lobes (
|
||
<figureCitation id="0380DAD8FFCCFFFEFBF1FE67FB56FE3C" box="[1122,1265,384,410]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="21" pageNumber="22">Figs 29, 30</figureCitation>
|
||
), median lobe projecting caudally, semicircular, and two-lipped; both lips continuous (
|
||
<figureCitation id="0380DAD8FFCCFFFEFC4CFE41FB88FE66" box="[991,1071,422,448]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="21" pageNumber="22">Fig.29</figureCitation>
|
||
). Mesepisternal tubercles (
|
||
<figureCitation id="0380DAD8FFCCFFFEFAFBFE41FF45FE41" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="21" pageNumber="22">Figs 29, 30</figureCitation>
|
||
) very high (2.83 mm), conical, with the apex directed laterally and bases adjacent to mid-dorsal carina (
|
||
<figureCitation id="0380DAD8FFCCFFFEFF33FE13FEA6FDA8" box="[160,257,500,526]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="21" pageNumber="22">Fig. 29</figureCitation>
|
||
). Cercus 1/3 shorter than paraproct (
|
||
<figureCitation id="0380DAD8FFCCFFFEFD40FE13FC96FDA8" box="[723,817,500,526]" captionStart="FIGURES 57 – 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 – 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 – 59); T. costalimai, paratypes (60 – 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="21" pageNumber="22">Fig. 64</figureCitation>
|
||
) in posterior view (
|
||
<figureCitation id="0380DAD8FFCCFFFEFBBEFE13FB2CFDA8" box="[1069,1163,500,526]" captionStart="FIGURES 57 – 65" captionStartId="24.[151,269,1907,1931]" captionTargetBox="[289,1277,524,1859]" captionTargetId="figure@24.[264,1323,493,1917]" captionTargetPageId="24" captionText="FIGURES 57 – 65. Tuberculobasis, male appendages in dorsal, lateral, and posterior view: T. arara, holotype (57 – 59); T. costalimai, paratypes (60 – 62); T. guarani, holotype (64, 65), T. inversa holotype in dorsal (63), dorso-lateral (66), and lateral (68) views. T. inversa, Tefé, Amazonas State, in postero-lateral (68) view [66, 67 by R. W. Garrison]." httpUri="https://zenodo.org/record/187817/files/figure.png" pageId="21" pageNumber="22">Fig. 65</figureCitation>
|
||
) with ventral process subtriangular. Penis as usual for the genus (as in
|
||
<figureCitation id="0380DAD8FFCCFFFEFD41FDFDFCB1FD92" box="[722,790,538,564]" captionStart="FIGURE 1" captionStartId="4.[151,255,1006,1030]" captionTargetBox="[562,1032,498,981]" captionTargetId="figure@4.[541,1045,465,982]" captionTargetPageId="4" captionText="FIGURE 1. Denticulobasis garrisoni, paratype: penis with distal segment raised [by R. W. Garrison]." httpUri="https://zenodo.org/record/187807/files/figure.png" pageId="21" pageNumber="22">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="9B04C65DFFCCFFFEFF55FDA7FD45FDFD" blockId="21.[151,1437,152,835]" box="[198,738,576,603]" pageId="21" pageNumber="22">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFF55FDA7FE0CFDFC" bold="true" box="[198,427,576,602]" pageId="21" pageNumber="22">Dimensions (mm).</emphasis>
|
||
Abdomen 32.5; HW 19.8.
|
||
</paragraph>
|
||
<paragraph id="9B04C65DFFCCFFFEFF55FD80FE0CFD24" blockId="21.[151,1437,152,835]" box="[198,427,615,642]" pageId="21" pageNumber="22">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFF55FD80FE86FD27" bold="true" box="[198,289,615,641]" pageId="21" pageNumber="22">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="D3A195D6FFCCFFFEFF55FD69FC8CFCE5" pageId="21" pageNumber="22" type="discussion">
|
||
<paragraph id="9B04C65DFFCCFFFEFF55FD69FC8CFCE5" blockId="21.[151,1437,152,835]" pageId="21" pageNumber="22">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFF55FD69FE9FFD0E" bold="true" box="[198,312,654,680]" pageId="21" pageNumber="22">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="5CBBBDDEFFCCFFFEFED9FD68FDC5FD0E" box="[330,610,655,680]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="guarani">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFED9FD68FDC5FD0E" box="[330,610,655,680]" italics="true" pageId="21" pageNumber="22">Tuberculobasis guarani</emphasis>
|
||
</taxonomicName>
|
||
has the highest mesepisternal tubercle within the genus (
|
||
<figureCitation id="0380DAD8FFCCFFFEFA8BFD69FAD4FD0E" box="[1304,1395,654,680]" captionStart="FIGURES 25 – 34" captionStartId="18.[151,270,1973,1997]" captionTargetBox="[327,1275,181,1941]" captionTargetId="figure@18.[312,1277,161,1944]" captionTargetPageId="18" captionText="FIGURES 25 – 34. Tuberculobasis, male thorax in dorsal and lateral view in T. arara, holotype (25, 26); T. costalimai, holotype (27, 28); T. guarani, holotype (29, 30); T. inversa, holotype (31, 32); T. williamsoni, holotype (33, 34)." httpUri="https://zenodo.org/record/187813/files/figure.png" pageId="21" pageNumber="22">Fig. 30</figureCitation>
|
||
). It belongs in the
|
||
<taxonomicName id="5CBBBDDEFFCCFFFEFED5FD52FE66FD68" box="[326,449,693,718]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="costalimai">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFED5FD52FE66FD68" box="[326,449,693,718]" italics="true" pageId="21" pageNumber="22">costalimai</emphasis>
|
||
</taxonomicName>
|
||
species group and it is close to
|
||
<emphasis id="A9CF1A4FFFCCFFFEFCADFD52FC00FD68" box="[830,935,693,718]" italics="true" pageId="21" pageNumber="22">
|
||
<taxonomicName id="5CBBBDDEFFCCFFFEFCADFD52FC04FD68" box="[830,931,693,718]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="arara">T. arara</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
It shares with this species the presence of subtriangular short ventral process of cercus but can be separated from it by characters given in the key. Together with
|
||
<emphasis id="A9CF1A4FFFCCFFFEFED2FCE5FDF8FCBD" box="[321,607,770,795]" italics="true" pageId="21" pageNumber="22">
|
||
<taxonomicName id="5CBBBDDEFFCCFFFEFED2FCE5FE7DFCBD" box="[321,474,770,795]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="costalimai">T. costalimai</taxonomicName>
|
||
,
|
||
<taxonomicName id="5CBBBDDEFFCCFFFEFE75FCE5FDF8FCBD" box="[486,607,770,795]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="guarani">T. guarani</taxonomicName>
|
||
</emphasis>
|
||
is the only
|
||
<taxonomicName id="5CBBBDDEFFCCFFFEFD74FCE5FC3EFCBD" box="[743,921,770,795]" class="Insecta" family="Coenagrionidae" genus="Tuberculobasis" kingdom="Animalia" order="Odonata" pageId="21" pageNumber="22" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A9CF1A4FFFCCFFFEFD74FCE5FC3EFCBD" box="[743,921,770,795]" italics="true" pageId="21" pageNumber="22">Tuberculobasis</emphasis>
|
||
</taxonomicName>
|
||
so far reported from SE
|
||
<collectingCountry id="E3AC86CDFFCCFFFEFB28FCE5FAA2FCBA" box="[1211,1285,770,796]" name="Brazil" pageId="21" pageNumber="22">Brazil</collectingCountry>
|
||
, and the first species of the genus recorded for the State of São Paulo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |