303 lines
34 KiB
XML
303 lines
34 KiB
XML
<document id="9820E75720DBA3DD0B2470980B654B72" ID-CLB-Dataset="26689" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.29172" ID-GBIF-Dataset="edfd877f-a449-46b9-beab-96b10bad752b" ID-Pensoft-Pub="2535-0730-2-137" ID-ZooBank="D49CE72306C24E38B5B0C7C311C8A89E" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-2-137" ModsDocOrigin="Evolutionary Systematics 2 (2)" ModsDocTitle="New species of tube web spiders of the genus Ariadna from South Australia (Araneae, Segestriidae)" checkinTime="1558621764846" checkinUser="pensoft" docAuthor="Marsh, Jessica R., Baehr, Barbara C., Glatz, Richard V. & Framenau, Volker W." docDate="2018" docId="11EA75D9BAC85A8E479E5571602E2C0E" docLanguage="en" docName="EvolutSyst 2(2): 137-149" docOrigin="Evolutionary Systematics 2 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.2.29172" docTitle="Ariadna clavata Marsh, Baehr, Glatz & Framenau, 2018, sp. n." docType="treatment" docUuid="6C44FD1E-17E3-4F3A-A289-778289106615" docUuidSource="ZooBank" docVersion="6" lastPageNumber="137" masterDocId="EC0CFFC4C843F74AFFD2FFB8D811FFAF" masterDocTitle="New species of tube web spiders of the genus Ariadna from South Australia (Araneae, Segestriidae)" masterLastPageNumber="149" masterPageNumber="137" pageNumber="137" updateTime="1701382727419" updateUser="ExternalLinkService">
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<mods:titleInfo id="84893F17C56B53F2A31C9136FFE771FF">
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<mods:title id="B5FAFED9848B78328DADC1E97FB99A0D">New species of tube web spiders of the genus Ariadna from South Australia (Araneae, Segestriidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A29865713822F2C08EF111061374C4C5" type="personal">
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<mods:namePart id="992D2D42FC276368877AA573529C22E7">Marsh, Jessica R.</mods:namePart>
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<mods:namePart id="DAFF3DE77A3BFE5945611E3AB43E4B9E">Baehr, Barbara C.</mods:namePart>
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<mods:namePart id="2A86CBF720091D7F79980217FC8FA004">Glatz, Richard V.</mods:namePart>
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<mods:name id="DB596DECEA30D46631A8057741486580" type="personal">
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<mods:namePart id="0B264C40B6CCA711DA4C74AB49F9C0A0">Framenau, Volker W.</mods:namePart>
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<mods:titleInfo id="42B65F45A915328F916D77FEBDCB281E">
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<mods:title id="9743DEFF7EB382E51D70CBC3FE80362A">Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part id="F854B7DE8AE5CBF942E0634E3115546C">
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<mods:date id="5D6F0E94C4EEA506EEE9D6238FE4200B">2018</mods:date>
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<mods:number id="83F7CEA5A6EC7F444596F0C675E68A04">2</mods:number>
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<mods:detail id="9C832ECED52E1F6448E227F6143AB9AD" type="issue">
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<mods:number id="7C90BF668BF5E748DD475386DCE186D2">2</mods:number>
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<mods:start id="E5BC3BA0EE9EAAC4BFBBBF1B74CA23B1">137</mods:start>
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<mods:url id="A705F94E3F9B8FC14B910BAF5BED1A65">http://dx.doi.org/10.3897/evolsyst.2.29172</mods:url>
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<mods:classification id="3628752CEEF0A326D21CDF99D0CD6B57">journal article</mods:classification>
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<mods:identifier id="F512F89A996092DDB1F4E3A72BC0536B" type="DOI">http://dx.doi.org/10.3897/evolsyst.2.29172</mods:identifier>
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<mods:identifier id="04DE5FE5CDF3A098EDD8981838AF38B9" type="Pensoft-Pub">2535-0730-2-137</mods:identifier>
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<mods:identifier id="DD7B3286AD929C0A97D81D14CDC70622" type="ZooBank">D49CE72306C24E38B5B0C7C311C8A89E</mods:identifier>
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<treatment id="11EA75D9BAC85A8E479E5571602E2C0E" ID-GBIF-Taxon="157251941" LSID="urn:lsid:zoobank.org:act:6C44FD1E-17E3-4F3A-A289-778289106615" httpUri="http://treatment.plazi.org/id/11EA75D9BAC85A8E479E5571602E2C0E" lastPageNumber="137" pageId="0" pageNumber="137">
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<subSubSection id="74BD5334F0806F1203AD0278394B8A6D" pageId="0" pageNumber="137" type="nomenclature">
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<paragraph id="1FE3B3524F5251F9589F56F4B217E1E8" pageId="0" pageNumber="137">
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<taxonomicName id="711CD799C7B9244264C9F685565962BF" ID-CoL="GLKW" LSID="http://zoobank.org/6C44FD1E-17E3-4F3A-A289-778289106615" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariadna clavata" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="clavata">Ariadna clavata</taxonomicName>
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<taxonomicNameLabel id="CFF2A1039912EBC12DEAE62B40AB5E7B" pageId="0" pageNumber="137">sp. n.</taxonomicNameLabel>
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Figs 1
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<normalizedToken id="5E3659C9EF9AA0A52BBC7555C9575D9F" originalValue="A–I">A-I</normalizedToken>
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, 2
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<normalizedToken id="0F4F7307DD0674E58D07DAA6C3A20344" originalValue="A–H">A-H</normalizedToken>
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, 3
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||
<normalizedToken id="BC746133837E79E287B13E2DA06FFB4A" originalValue="A–C">A-C</normalizedToken>
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||
, 8
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="335864B9C1E22130D36022795FB6C6B9" pageId="0" pageNumber="137" type="type material">
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<paragraph id="FFE20CAED642FCFD3083DD2A7A362FF4" pageId="0" pageNumber="137">Type material.</paragraph>
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<paragraph id="A432F48F1523BBFD0AA9BB713ABC80D6" pageId="0" pageNumber="137">
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AUSTRALIA: South Australia: Holotype: ♂, Tangara Drive, American River, Kangaroo Island, -
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<geoCoordinate id="E31223006232E9B2FB1AA134E28814DE" direction="south" orientation="latitude" precision="1" value="-35.78733">35.787329S</geoCoordinate>
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,
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<geoCoordinate id="700FB4D4335BFD21D3DD7231DF0C90CD" direction="east" orientation="longitude" precision="1" value="137.76721">137.767213E</geoCoordinate>
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, 3 June 2017, under bark at 1.5 m height,
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<taxonomicName id="CC3129F012C2A2AC75508C619DA0A845" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus diversifolia" order="Myrtales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="diversifolia">Eucalyptus diversifolia</taxonomicName>
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(1 m diameter), J. Marsh (SAM NN29861). Paratypes: 1♀, same data as holotype, except 8 July 2017, observed mating with holotype male in captivity, J. Marsh (SAM NN29862); 1♂, Pelican Lagoon walking trail, American River, Kangaroo Island, -
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<geoCoordinate id="0C76F666C31753203025FB9717C6915B" direction="south" orientation="latitude" precision="1" value="-35.796497">35.796496S</geoCoordinate>
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,
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<geoCoordinate id="BBA5B0A3FCC4AC25ECF97F49BC3577F1" direction="east" orientation="longitude" precision="1" value="137.75096">137.750963E</geoCoordinate>
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, 2 Nov 2017, under bark
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<taxonomicName id="52A04882CB9369D501F50AC4EA94F205" lsidName="E. diversifolia" pageId="0" pageNumber="137" rank="species" species="diversifolia">E. diversifolia</taxonomicName>
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, J. Marsh (CeNak ZMH-A0003051); 1♀, Tangara Drive, American River, Kangaroo Island, -
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<geoCoordinate id="A55BF525C13D945B73C675223221DFC0" direction="south" orientation="latitude" precision="1" value="-35.782887">35.782887S</geoCoordinate>
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,
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<geoCoordinate id="50EBB038396AD987249025884CBEDACF" direction="east" orientation="longitude" precision="1" value="137.77156">137.771559E</geoCoordinate>
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, 16 Aug 2017, in tube web at 1 m height, in crevice of bark of
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<taxonomicName id="23CE83982F708E82D58578ECF4178EB9" class="Magnoliopsida" family="Casuarinaceae" genus="Allocasuarina" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Allocasuarina muelleriana" order="Fagales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="muelleriana">Allocasuarina muelleriana</taxonomicName>
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, J. Marsh (CeNak ZMH-A0003052).
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</paragraph>
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</subSubSection>
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<subSubSection id="3299167F46BCB10E56EA9D32A8CE6A47" pageId="0" pageNumber="137" type="materials_examined">
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<paragraph id="C45323786899567EBA4CC23157FDE87C" pageId="0" pageNumber="137">Other material examined.</paragraph>
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<paragraph id="4F0B14F7F7AFCA95B844D8787A034595" pageId="0" pageNumber="137">
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AUSTRALIA: South Australia: Kangaroo Island: 1♂, Cannery walking trail, American River, -
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<geoCoordinate id="63B013412B438A87F5DC52D4E0D0D721" direction="south" orientation="latitude" precision="1" value="-35.77371">35.773707S</geoCoordinate>
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,
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<geoCoordinate id="17AE9469D6450D601F4E4E8FDAB858F3" direction="east" orientation="longitude" precision="1" value="137.78111">137.781111E</geoCoordinate>
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, 16 Aug 2017 (SAM NN29879); 1♀, Pelican Lagoon Walking Trail, American River, -
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<geoCoordinate id="DF6680A91C6218E3E3F517853B7AF703" direction="south" orientation="latitude" precision="1" value="-35.79258">35.792580S</geoCoordinate>
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,
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<geoCoordinate id="EBE35D24FC5C9EE2D1525DA6A1B845F7" direction="east" orientation="longitude" precision="1" value="137.7579">137.757909E</geoCoordinate>
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, 1 Sept 2017 (SAM NN29876); 2♂, same data, except 10 Oct 2017 (SAM NN29881; J. Marsh reference collection, Se092); 4♂, 1♀ same data, except -
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<geoCoordinate id="89F794D3A29CF401081088C2439D9FD3" direction="south" orientation="latitude" precision="1" value="-35.796497">35.796496S</geoCoordinate>
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,
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<geoCoordinate id="8BAE2F8C9574C20F63721F157EC1F47D" direction="east" orientation="longitude" precision="1" value="137.75096">137.750963E</geoCoordinate>
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, 2 Nov 2017 (SAM NN29897, NN29899; J. Marsh reference collection, Se058); 1♂, Tangara Drive, American River, -
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<geoCoordinate id="32F48F55C1E19200946F28A762FB1063" direction="south" orientation="latitude" precision="1" value="-35.78718">35.787181S</geoCoordinate>
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,
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<geoCoordinate id="526E71116DB5FA72858B6C82DD2E0625" direction="east" orientation="longitude" precision="1" value="137.767">137.766994E</geoCoordinate>
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, 2 June 2017 (SAM NN29865); 1♂, same data, except 3 Jun 2017 (SAM NN29866); 1♂, same data, except 12 Jun 2017 (SAM NN29870); 1♂, 3♀, Antechamber Bay, Chapman River, -
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<geoCoordinate id="998AAD7E543CBFC79C5BA4B86703A862" direction="south" orientation="latitude" precision="1" value="-35.790737">35.790736S</geoCoordinate>
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,
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<geoCoordinate id="40C37788725772DAADC70646609C408B" direction="east" orientation="longitude" precision="1" value="138.06555">138.065559E</geoCoordinate>
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, 1 Sept 2017 (SAM NN29892); 2♂, Baudin Conservation Park, Dudley Peninsula, -
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<geoCoordinate id="8B1314F246FE5B3EB9F3B9BD7939EFD7" direction="south" orientation="latitude" precision="1" value="-35.734848">35.734848S</geoCoordinate>
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,
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<geoCoordinate id="D19F4E4B8D4BBA17C4E921CCCBD16740" direction="east" orientation="longitude" precision="1" value="137.95921">137.959212E</geoCoordinate>
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, 15 Jul 2017 (SAM NN29908); 1♀, Dudley Conservation Park, Dudley West, -
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<geoCoordinate id="0E981343B4E3B28860F10C9894A2E39C" direction="south" orientation="latitude" precision="1" value="-35.79532">35.795318S</geoCoordinate>
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,
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<geoCoordinate id="1C49104B9DE664AFBF982CE7A4AD9AC5" direction="east" orientation="longitude" precision="1" value="137.85">137.850007E</geoCoordinate>
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, 22 Jun 2017 (SAM NN29911); 1♀, same data, except -
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<geoCoordinate id="1D61328B5D42083D5E777823F83006D6" direction="south" orientation="latitude" precision="1" value="-35.795452">35.795452S</geoCoordinate>
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,
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<geoCoordinate id="C36B57300E89139BA4A4462E150DBC81" direction="east" orientation="longitude" precision="1" value="137.86018">137.860189E</geoCoordinate>
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(SAM NN29872); 1♂, same data, except -
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<geoCoordinate id="687DA306F831BCA605DB500A1627B970" direction="south" orientation="latitude" precision="1" value="-35.795944">35.795946S</geoCoordinate>
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,
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<geoCoordinate id="311B3F70874D60008057EA5788B757C0" direction="east" orientation="longitude" precision="1" value="137.8608">137.860794E</geoCoordinate>
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(SAM NN29868); 1♀, Pelican Lagoon Conservation Park, Dudley West, -
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<geoCoordinate id="AB4B79ACE7B69BF38A6A18A1808B5393" direction="south" orientation="latitude" precision="1" value="-35.801685">35.801685S</geoCoordinate>
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,
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<geoCoordinate id="9FE4816D3DAC2BEAC4B9568C88468FAB" direction="east" orientation="longitude" precision="1" value="137.77422">137.774214E</geoCoordinate>
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, 13 Oct 2017 (SAM NN29886); 3♂, 3♀, Simpson Road, Dudley West, -
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<geoCoordinate id="0B0743CCC5EAD17A063A33C352898625" direction="south" orientation="latitude" precision="1" value="-35.823524">35.823523S</geoCoordinate>
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,
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<geoCoordinate id="70D31F89612CD80596DD1FCC51149240" direction="east" orientation="longitude" precision="1" value="137.83104">137.831032E</geoCoordinate>
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, (SAM NN29910); 2♀, Three Chain Road, MacGillivray, -
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<geoCoordinate id="034F6FA35029FFEBADEF02A796DF803E" direction="south" orientation="latitude" precision="1" value="-35.910675">35.910674S</geoCoordinate>
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,
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<geoCoordinate id="4E7D29432506DE56892C7F41AA201050" direction="east" orientation="longitude" precision="1" value="137.55049">137.550490E</geoCoordinate>
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, 26 Jul 2017 (SAM NN29909, NN29874); 4♀, same data, except 7 Aug 2017 (SAM NN29888, NN29889, NN29890); 4♀, roadside, near Parndana Conservation Park, -
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<geoCoordinate id="5FB109FA2F37FE91B187FCBF2B4A59A3" direction="south" orientation="latitude" precision="1" value="-35.757187">35.757187S</geoCoordinate>
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,
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<geoCoordinate id="9D0C1354E6D551FA42AAD39975FD3A75" direction="east" orientation="longitude" precision="1" value="137.30731">137.307314E</geoCoordinate>
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, 17 Aug 2017 (SAM NN29891); 1♂, 2♀ roadside, near Parndana Conservation Park, -
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<geoCoordinate id="C23650BE1C6CBB864A233B8EA0EC4243" direction="south" orientation="latitude" precision="1" value="-35.75762">35.757621S</geoCoordinate>
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<geoCoordinate id="B74824C3F23AC3D89348D76481B7C30E" direction="east" orientation="longitude" precision="1" value="137.30733">137.307326E</geoCoordinate>
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, 10 Aug 2017 (SAM NN29883); 1♀, same data, except -
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<geoCoordinate id="36E89A98E3E1C2EF74D2FB8499B8F6AE" direction="south" orientation="latitude" precision="1" value="-35.756706">35.756707S</geoCoordinate>
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,
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<geoCoordinate id="789F7903618D0C6275260C6810C3653B" direction="east" orientation="longitude" precision="1" value="137.30652">137.306523E</geoCoordinate>
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(NN29895); 1♂, same data, except -35.757047S, 137.307237E (SAM NN29893); 1♂, same data, except -
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<geoCoordinate id="6945CE8FA797A64B640157618C2D1227" direction="south" orientation="latitude" precision="1" value="-35.75713">35.757131S</geoCoordinate>
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,
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<geoCoordinate id="38E49D7AEDCEE6442AE0EC7B8916DB5E" direction="east" orientation="longitude" precision="1" value="137.3072">137.307199E</geoCoordinate>
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(SAM NN29880). Mount Lofty Ranges: 1♂, 1♀, Devils Gully Native Forest Reserve, Kersbrook, -
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<geoCoordinate id="36EEAC20C33877CCFD5D08713C898FC8" direction="south" orientation="latitude" precision="1" value="-34.755554">34.7555556S</geoCoordinate>
|
||
,
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<geoCoordinate id="82520397C4E22BB3C7942B648A795615" direction="east" orientation="longitude" precision="1" value="138.8189">138.818889E</geoCoordinate>
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||
, Oct/ Nov 2000 (SAM, not accessioned); 1♂, Sixth Creek Native Forest Reserve, Forest Range, -
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<geoCoordinate id="B8D295E7072A1052031419AE034CD24C" direction="south" orientation="latitude" precision="1" value="-34.900276">34.9002778S</geoCoordinate>
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,
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<geoCoordinate id="36F3EF8B0084866D04CD0CF0823DE8D5" direction="east" orientation="longitude" precision="1" value="138.79417">138.794167E</geoCoordinate>
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, Oct/ Nov 2000 (SAM, not accessioned). Fleurieu Peninsula: 3♂, 2♀, Spring Mount, -
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<geoCoordinate id="79228F4BF4B38DA3DEAF81EE66B29794" direction="south" orientation="latitude" precision="1" value="-35.43222">35.432222S</geoCoordinate>
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,
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<geoCoordinate id="53122165E57E557529FE804B0D86921D" direction="east" orientation="longitude" precision="1" value="138.5239">138.523889E</geoCoordinate>
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Dec/ Jan 2006 (SAM, not accessioned).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3B5068B71C92C7F3A8855C52E8A918A0" pageId="0" pageNumber="137" type="etymology">
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<paragraph id="C2316DC4CEBA88D0CE6AE0796DA368C3" pageId="0" pageNumber="137">Etymology.</paragraph>
|
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<paragraph id="F14FCF41C1A0B74ECB81CD43101046F5" pageId="0" pageNumber="137">The specific epithet is an adjective (Latin, clavata - striped) referring to the striped abdominal markings of both the male and female.</paragraph>
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</subSubSection>
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<subSubSection id="2E83B05F2AF8B21A8B95659B2F802E7F" pageId="0" pageNumber="137" type="diagnosis">
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<paragraph id="08CB696A5DC95D8517691F024BC6ABF7" pageId="0" pageNumber="137">Diagnosis.</paragraph>
|
||
<paragraph id="FDD8CAF7800AB15206B566D374AC43B8" pageId="0" pageNumber="137">
|
||
Males and females of
|
||
<taxonomicName id="EDC5DC751DB0A324B2DC7955519E3378" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
sp. n. are distinguished from
|
||
<taxonomicName id="63EFA41E897013CD6BC4D785410AC31C" lsidName="A. segmentata" pageId="0" pageNumber="137" rank="species" species="segmentata">A. segmentata</taxonomicName>
|
||
by the curvature of the posterior eye row, which is strongly procurved in
|
||
<taxonomicName id="9C82ECD9622DBC9B7F86EDF7829FA4E7" lsidName="A. segmentata" pageId="0" pageNumber="137" rank="species" species="segmentata">A. segmentata</taxonomicName>
|
||
(Hickman, 1967, fig. 63) and which is slightly recurved in
|
||
<taxonomicName id="B9EFB22D0615CF578EC6D8F2E52F7095" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
(Fig. 2C). Males of
|
||
<taxonomicName id="4BFB1D9515DBAFFB3BBE669CDC83AC71" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="4D29ED97D1573C1749FB9C1317285492" lsidName="A. segmentata" pageId="0" pageNumber="137" rank="species" species="segmentata">A. segmentata</taxonomicName>
|
||
by the long, curved and distally hooked embolus (Fig. 1G, I), while the embolus of
|
||
<taxonomicName id="4E458B19EF7CEFA51AB4FB3B260EFA16" lsidName="A. segmentata" pageId="0" pageNumber="137" rank="species" species="segmentata">A. segmentata</taxonomicName>
|
||
is shorter and stouter (
|
||
<bibRefCitation id="96FF6BDE28A076542CB108414D06E941" pageId="0" pageNumber="137">Hickman 1967</bibRefCitation>
|
||
, fig. 65). The pedipalp bulb of
|
||
<taxonomicName id="938E34E26D47CCD454F902EB09EAADF6" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="5A9C149A9DCF528748AC951D6A6C9EB4" lsidName="A. muscosa" pageId="0" pageNumber="137" rank="species" species="muscosa">A. muscosa</taxonomicName>
|
||
by being roughly pyramidal in shape, while that of
|
||
<taxonomicName id="04EBD3DA72D36B4511BD52C73E59E93E" lsidName="A. muscosa" pageId="0" pageNumber="137" rank="species" species="muscosa">A. muscosa</taxonomicName>
|
||
is spheroidal (Fig. 1G, I;
|
||
<bibRefCitation id="CEB3999BFD473C1E4AF972B3B81BC65C" author="Hickman, VV" journalOrPublisher="Papers and Proceedings of the Royal Society of Tasmania" pageId="0" pageNumber="137" pagination="96 - 118" title="Studies in Tasmanian spiders. Part III." volume="1928" year="1929">Hickman 1929</bibRefCitation>
|
||
, fig. 3a). Males and females of
|
||
<taxonomicName id="7A8E50B479803BDADD407AD2AE0AE347" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="DCF8FBFE51DFFF8378276739C941D625" lsidName="A. muscosa" pageId="0" pageNumber="137" rank="species" species="muscosa">A. muscosa</taxonomicName>
|
||
by the number of teeth on the tarsal claws of the first legs;
|
||
<taxonomicName id="7A111CB0730FFD225BAFAFEFBF9C3D9D" lsidName="A. muscosa" pageId="0" pageNumber="137" rank="species" species="muscosa">A. muscosa</taxonomicName>
|
||
has 7 or 8 teeth, with the inferior claw bare,
|
||
<taxonomicName id="8B4C22348ECC4C3EC2145B0B68851D07" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
has 10 or 11 teeth, with a small tooth on the inferior claw (Fig. 2G). Females of
|
||
<taxonomicName id="19165C9F2BA41AE81410B07D11E7BBE3" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
differ from
|
||
<taxonomicName id="13C9AB4E5512349F141B76A85977DC99" lsidName="A. burchelli" pageId="0" pageNumber="137" rank="species" species="burchelli">A. burchelli</taxonomicName>
|
||
by the number of teeth on the chelicerae;
|
||
<taxonomicName id="3AAF79B466F806A906438FAFB7DD2B2E" lsidName="A. burchelli" pageId="0" pageNumber="137" rank="species" species="burchelli">A. burchelli</taxonomicName>
|
||
has one tooth either side of the fang furrow,
|
||
<taxonomicName id="981FA95F2B65B0D97B907675D4A852EE" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
has three teeth on the promargin, one tooth on the retromargin (Fig. 2B, C).
|
||
<taxonomicName id="99AA482CC1B2F1D2A7FE5A56FA196A02" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariadna clavata" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="clavata">Ariadna clavata</taxonomicName>
|
||
differs from all remaining Australian
|
||
<taxonomicName id="0B5C31B905F4FDB46A14D88E3FC1A209" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariadna" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="genus">Ariadna</taxonomicName>
|
||
by the presence of transverse abdominal markings (Fig. 1A).
|
||
</paragraph>
|
||
<caption id="98EB898ED7CD5D4C2B16DE2D4F18C870" pageId="0" pageNumber="137">
|
||
<paragraph id="7D12FAFE6FC18E496B01925642F96C98" pageId="0" pageNumber="137">
|
||
Figure 1.
|
||
<taxonomicName id="BCFFC03CED313E136710DD29E86FD80B" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariadna clavata" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="clavata">Ariadna clavata</taxonomicName>
|
||
sp. n., male holotype, from American River, Kangaroo Island (SA; SAM NN29861): A habitus, dorsal view B habitus, ventral view C chelicerae and eyes, dorsal view D left leg I femur, dorsal view E left leg IV, metatarsus preening comb, retrolateral view F left leg I, tarsus and tarsal claws, prolateral view G palp, prolateral view H palp, dorsal view I palp, retrolateral view. Scale bars: 1.0 mm (A, B, D); 0.1 mm (others).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="09EDD2471C6341E881B107F8012209E9" pageId="0" pageNumber="137" type="description">
|
||
<paragraph id="064E78EE703B1AE2115323410B575204" pageId="0" pageNumber="137">Description.</paragraph>
|
||
<paragraph id="31ABE4860FC36BFCA85C170DBFC27E31" pageId="0" pageNumber="137">
|
||
Male holotype (SAM NN29861). Carapace length 3.5 mm, dark reddish brown, becoming darker anteriorly, covered with sparse dark setae; sternum brownish yellow, darker at the edges and between coxae; endites dark brown, chelicerae dark brown to black; legs yellowish brown; abdomen length 3.9 mm, dorsally pale yellow with 8 dark transverse bands, connected by triangular extensions at the midpoint (Fig. 1A), ventrally uniform yellow/grey (Fig. 1B). Carapace oval, narrowing anteriorly and apically square; raised anteriorly, highest just behind posterior eye row; lateral margins slightly undulating (Fig. 1A). Fovea an indented pit, with faint dark radial striae extending outwards from fovea. Labium narrowing anteriorly,
|
||
<normalizedToken id="69D654205E34AC3D51BEF1A71B23A0B6" originalValue="¾">3/4</normalizedToken>
|
||
the length of maxillae. Chelicerae semi-porrect, with strong lateral ridges dorsally and with basal transverse ridges (Fig. 1C), retromargin with single tooth, promargin with three teeth. Sternum oval with precoxal triangles and with small, rounder intercoxal extensions (Fig. 1B). Posterior eye row slightly recurved (Fig. 1C). Femur I bowed in dorsal view (Fig. 1D). Leg measurements (
|
||
<normalizedToken id="0F4DB20C033B9A5D8D82054C14DC7A30" originalValue="I–IV">I-IV</normalizedToken>
|
||
): femora 2.74, 2.65, 2.06, 2.28; patellae 1.13, 1.16, 0.99, 1.06; tibiae 2.34, 2.53, 1.50, 1.76; tarsi 2.14, 2.29, 1.64, 1.72; metatarsi 0.97, 0.85, 0.79, 0.80; total 9.32, 9.48, 6.98, 7.62. Spines: Leg I: femur d1-1-1-1, dp2ap (Fig. 1D); tibia dp1-1, p1-1, vp1-1, vp1ap, v1, vr1-1/0-1/0, vr1ap, dv1-1/0; metatarsus vp1-1, vp1ap, v1, vr1-1, vr1ap. Leg II: femur d1-1-1-1-1/0, dp1ap; tibia dp1-1-1, p1, vp1ap, vr1-1-1/0, vr1ap; metatarsus vp1-1, vp1ap, v1, v1ap, vr1-1, vr1ap. Leg IV: femur d1-1-1-1/0, dp1ap, tibia dp1-1/0, vp1, vp1ap, vr1-1, vr1ap, dr1-1; metatarsus dp1, v1, vr1-1, vr2ap, r1-1, retrolateral distal preening comb with five spines (Fig. 1E). Legs I and II tarsal claws with 10 or 11 teeth, one small tooth on inferior claw (Fig. 1F). Legs III and IV tarsal claw with six teeth and inferior claw bare. Tarsi with distal ventral setae, projecting beneath claws (Fig. 1F). Pedipalp femur expanded apically, on the ventral retrolateral edge, so it appears slightly concave ventrally. Pedipalp tibia short, about 1.5 times as long as the cymbium, and swollen in the dorso-ventral plane (Fig. 1G, I). Bulb roughly pyramidal with embolus arising from the ventral point of the pyramid. Embolus long, thin, hooked apically (Fig. 1
|
||
<normalizedToken id="23E4875C10F127575B7C953EF17C46A2" originalValue="G–I">G-I</normalizedToken>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="BED2B5306152C5AC4F37D74C09730FF4" pageId="0" pageNumber="137">
|
||
Female paratype (SAM NN29862). Carapace length 3.8 mm, brown, sternum light brown, with darker patches between coxae; endites yellowish brown, chelicerae brown; legs light brown. Abdomen length 4.6 mm, dorsally pale yellow with 8 dark transverse bands, connected by triangular extensions at the midpoint (Fig. 2A). Abdomen ventrally yellowish grey with faint longitudinal darker line centrally (Fig. 2B). Carapace dorsal surface sparsely covered with black setae, with stronger and longer black setae projecting forward from the clypeus (Fig. 2C). Carapace elongated and oval, narrowing anteriorly and apically squared; carapace domed so when viewed laterally highest point is mid-way between fovea and posterior eyes; lateral margin of carapace slightly undulating, fovea a shallow indented pit (Fig. 2A). Sternum oval with precoxal triangles and small, rounder inter-coxal extensions (Fig. 2B). Labium narrowing anteriorly, about
|
||
<normalizedToken id="963CEC07CC5B299B4D5BC26D558C9568" originalValue="¾">3/4</normalizedToken>
|
||
length of maxillae. Posterior eye row slightly recurved, chelicerae hypognathous, covered in strong, long black setae and with basal transverse ridges (Fig. 2C), retromargin with single tooth, promargin with three teeth. Femur I bowed in dorsal view (Fig. 2A, E). Leg measurements (
|
||
<normalizedToken id="E27086BCAF430EF8EE540C43D98BCF2B" originalValue="I–IV">I-IV</normalizedToken>
|
||
): femora 2.83, 2.59, 1.96, 2.23; patellae 1.21, 1.26, 1.03, 1.25; tibiae 2.08, 2.16, 1.41, 1.8; metatarsi 1.66, 1.76, 1.31, 1.52; tarsi 0.84, 0.89, 0.75, 0.76; total 8.62, 8.67, 6.46, 7.55. Spines: Leg I: femur dp2ap (Fig. 2E); tibia vp1-1-1, vp1ap, v1, vr1-1, vr1ap, v1/0; metatarsus vp1-1-1, vp1ap, v1, v1ap, vr1-1, vr1ap (Fig. 2
|
||
<normalizedToken id="901C7C9CDE565EFB7DA7D264CA2320DC" originalValue="D–F">D-F</normalizedToken>
|
||
). Leg II: femur dp1ap; tibia p1, vp1ap, v1-1, v1ap; metatarsus vp1-1-1, vp1ap, v1, v1ap, vr1-1-1/0, vr1ap. Leg IV: femur d1-1, dp1ap, tibia p1, vp1ap, v1, vr1-1, vr1ap; metatarsus v1, vr1-1, vr2ap (1 spine broken), distal retrolateral preening comb with five spines (Fig. 2H). Tarsi with distal ventral hairs, projecting beneath claws, tarsal claws I and II with 11/12 teeth, inferior claw with one small tooth (Fig. 2G). Legs III and IV tarsal claw with six teeth, inferior claw bare. Pedipalps with multiple strong prolateral spines on the tarsus and tibia, and with a single toothless claw. Epigastrium a slightly raised, lightly sclerotized external plate. Internal genitalia: Anterior receptaculum bilobed, dorsal lobe rounded and about 1.5 times as long as ventral lobe. Fig. 3
|
||
<normalizedToken id="7E9BFC3E9CB0B9CC89CE0840A05161C8" originalValue="A–C">A-C</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
<caption id="BEB29A3A66768F50188B9D6DDEEDE12D" pageId="0" pageNumber="137">
|
||
<paragraph id="107F50FCA6296A9E71DB3D46DA79F434" pageId="0" pageNumber="137">
|
||
Figure 2.
|
||
<taxonomicName id="E80F46FF5C829BAA1CDEB3C2D270C70F" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariadna clavata" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="clavata">Ariadna clavata</taxonomicName>
|
||
sp. n., female paratype, from American River, Kangaroo Island (SA; SAM NN29862): A habitus, dorsal view B habitus, ventral view C chelicerae and eyes, dorsal view D left leg I spines prolateral view (mirrored) E femur, dorsal view F tibia and metatarsus, spination ventral view G tarsus and tarsal claws, prolateral view, arrow indicates inferior claw tooth H left leg IV, metatarsus preening comb, retrolateral view (mirrored). Scale bars: 1.0 mm (A, B, D); 0.1 mm (others).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="B4173D98B3157A96A39DC49379FCA71C" pageId="0" pageNumber="137">
|
||
<paragraph id="E33C262B1D83D86C9C3359B267C6DF0F" pageId="0" pageNumber="137">
|
||
Figure 3.
|
||
<taxonomicName id="97411F76406CC39A2BA1B71A8B696945" class="Arachnida" family="Segestriidae" genus="Ariadna" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ariadna clavata" order="Araneae" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="clavata">Ariadna clavata</taxonomicName>
|
||
sp. n., female,
|
||
<normalizedToken id="F46EE11401E3A0B0016937811683E1BC" originalValue="D’Estrees">D'Estrees</normalizedToken>
|
||
Bay, Kangaroo Island (SA; SAM NN29889): A internal genitalia, ventral view B same, showing structure of AR ventral view C same, showing structure of AR, lateral view. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="9A1D614C3C88FD51C147C52F9245E16D" pageId="0" pageNumber="137" type="variation">
|
||
<paragraph id="AC3B5D602F883C12ADD590294E874519" pageId="0" pageNumber="137">Variation</paragraph>
|
||
<paragraph id="2BB18D532AF0762A00D99777A5A3A856" pageId="0" pageNumber="137">
|
||
(see Suppl. material 1: Table S1): Males (n=21), carapace length: range 2.46-3.64 mm, mean 3.14 mm, standard deviation 0.38 mm. Females (n=23), carapace length: range 2.57-4.34 mm, mean 3.63 mm, standard deviation 0.47 mm. Whilst leg spination showed a large amount of variation, both between and within specimens, some spines showed a higher level of consistency and this was true for both males and females e.g. (1) number of prolateral macrosetae at the apex of the femur I, and (2) the paired macrosetae at the apices of tibiae and metatarsi I and II. The remainder of the spination varied between specimens. The structure of the
|
||
<normalizedToken id="A03385E89FF06D7B580DF155B12EE8AF" originalValue="‘preening’">'preening'</normalizedToken>
|
||
combs on the metatarsus IV was constant in all specimens examined (Figs 1E, 2H).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DD977CB962DF6E30F7F7E67C2524F1BA" pageId="0" pageNumber="137" type="distribution">
|
||
<paragraph id="87B9C0F682238A8C57CE7CC0926B95F3" pageId="0" pageNumber="137">Distribution.</paragraph>
|
||
<paragraph id="2E9655A7E7C7D80BED761B72E95ED1FF" pageId="0" pageNumber="137">South-eastern South Australia, including the Fleurieu Peninsula, the Mount Lofty Ranges and Kangaroo Island (Fig. 8).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0F903267B01F06FA9EA2A2ACA1091E74" pageId="0" pageNumber="137" type="life history and habitat preferences">
|
||
<paragraph id="CC5616E54E432FDD1D8D9B9A9B3548E0" pageId="0" pageNumber="137">Life history and habitat preferences.</paragraph>
|
||
<paragraph id="7C7ACA51BAFB6CB49583CC6963EFE039" pageId="0" pageNumber="137">
|
||
The majority of
|
||
<taxonomicName id="9A444B177E2493D9E531F318A607E164" lsidName="A. clavata" pageId="0" pageNumber="137" rank="species" species="clavata">A. clavata</taxonomicName>
|
||
specimens were collected from within tube webs located beneath rugose bark of older
|
||
<taxonomicName id="2F66D2511D0A0408936700E58B036EE4" class="Magnoliopsida" family="Myrtaceae" genus="Eucalyptus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Eucalyptus" order="Myrtales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="genus">Eucalyptus</taxonomicName>
|
||
trees. Rather than tree species, the structure of the bark (with specimens found in older, cracked or layered bark), appeared to be of importance. The species appears to be somewhat opportunistic in terms of the location of its retreats, being a variety of holes and crevices, such as borer holes in dead wood (both as standing trees and in logs lying on the ground), in tubular holes in limestone and in burrows in loose soil beneath rocks. The main habitat type was mallee woodland. Specimens were also collected from isolated trees on roadside verges and from a stand of
|
||
<taxonomicName id="40DE421D95CC664D53E7C2E488787949" class="Pinopsida" family="Pinaceae" genus="Pinus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Pinus radiata" order="Pinales" pageId="0" pageNumber="137" phylum="Tracheophyta" rank="species" species="radiata">Pinus radiata</taxonomicName>
|
||
near
|
||
<normalizedToken id="3733BF433508ECD645B039DC68EDA46E" originalValue="D’Estrees">D'Estrees</normalizedToken>
|
||
Bay (Kangaroo Island). Mature males were collected manually throughout autumn, winter and spring (March through to October). Mature females were present all year.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |